(®gures 1, 4, 6, 7)

Type material. HOLOTYPE male. Malaysia, Sabah, Danum Valley, Sapat Kalisan, locality 15, 12 February 1997, H. Zettel (NMW). PARATYPE: 1 female, same label data as holotype (NMW).

Size. Length of beetle 2.9 mm (male), 2.8 mm (female); width 1.6 mm (male), 1.5 mm (female).

Description (male). Colour. Beetle almost completely ferruginous. Elytron with dark brown, barely contrasting maculations (®gure 1).

Sculpture. Beetle dorsally and ventrally shiny. Head smooth medially, with small punctures laterally along eyes; with microreticulation and some minute punctures posterior of an imaginary line connecting hind margins of eyes. Pronotum with dense punctation, punctures larger than those on head, regularly spaced, separated by about twice their diameter; no microreticulation present; basal stria sinuate and faint, about one-third of pronotal length. Elytron with dense punctation as on pronotum, not microreticulate; no serial rows of larger punctures present. Metasternum and metacoxal plates microreticulate and with larger punctures; sternites microreticulate with few smaller punctures.

Median lobe. Median lobe of aedeagus simple, symmetrical in ventral view. Rather straight and only slightly bent distally in lateral view; in ventral view broad, more or less parallel-sided over most of its length, narrowing apically, apex pointed. Paramere two-segmented, apical segment almost as long as basal segment; tip twisted (®gures 4, 6, 7).

Female. As in male except: beetle dull because of distinct microreticulation of small cells present all over the head, pronotum and elytron.

Etymology. Named for our colleague and friend, the collector of this species, Herbert Zettel (Vienna Museum).

Distribution. East Malaysia, northern Sabah (Borneo), see ®gure 8.

Collecting notes. In shallow, small isolated puddles, at the edge of a river.