Cardiodactylus rizali Robillard, n. sp.

(Figs. 4, 6I, 10F, 51)

Type material. Male holotype: Philippines. North Zamboanga Province, Mindanao, Dapitan, Baker, 7523 (MNHN-EO-ENSIF1192).

Type locality. Philippines, Mindanao Island, Dapitan.

Etymology. Named after Jose Rizal, the national hero of the Philippines who was exiled in the city of Dapitan.

Distribution. Philippines, north of Mindanao Island.

Diagnosis. Species of average to large size, coloration brownish with yellow brown patterns, characterized by head coloration and shape of pseudepiphallic sclerite in male genitalia with hook-like anterior expansions and slightly asymmetrical lateral expansions of dorsal ridges close to C. muria. Male genitalia close to that of C. empagatao Otte, 2007a (shape of pseudepiphallic dorsal ridges and rami little sclerotized) but the species differ strikingly in terms of coloration.

Description. Average size for the species group. General coloration orange brown with yellow and dark areas (Fig. 51A–B). Head dorsum with 4 dark brown bands, 2 wide lateral ones with an anterior notch separated from black coloration of fastigium, and 2 thin median parallel and punctuated bands; short triangular bands posterior to eyes slightly connected to lateral bands. Scapes yellow brown with a dark brown ring. Antennae orange brown. Face yellow brown, with a transverse dark brown band below antennae; front part of fastigium with faint parallel dark stripes. Mouthparts yellow brown. Maxillary palpi brown, apex darker. Lateral part of head dark brown dorsally, lateral margin gray brown. Pronotum: Dorsal disk 4 corners yellow; median area yellow brown with dark brown patterns; posterior area dark brown. Lateral lobes dark brown, ventral margin yellow. Legs I–II orange brown, with faint dark spots on femora and rings on tibiae. FIII orange brown, knees and tibiae dark brown. Hind wing tail dark brown, transverse veins yellow. Cerci orange brown. Apex of abdomen dark brown, sternites yellow brown.

Male: FW coloration mostly dark brown (Fig. 10F); area anterior to stridulatory file black; orange brown sclerotization near base of chords. Venation mostly yellowish, except part of 2A, posterior half of diagonal and apical field, orange brown. Small oval area posterior to mirror with translucent cells and yellow veins. Lateral field brown. M/R area dark red brown. Sc vein orange brown, apex yellow and bifurcations whitish, including their basis. FW venation (Fig. 10F): 1A slightly bisinuated. CuP faint but present until the file level. Harp with 1 wshaped harp vein and a faint anterior vein. Mirror area: mirror (d1) longer than wide, oval, well defined, separated in two parts by a strong transverse vein, the posterior part smaller and triangular; d2 crossed by several accessory veins; e1 not crossed by accessory veins. Apical field with 4 cell alignments posterior to mirror. Lateral field with 8 projections of Sc and 3 more ventral veins.

Male genitalia (Fig. 51C–E): Pseudepiphallus forming a narrow gutter, very sclerotized and narrowed preapically. Dorsal ridges parallel, slightly carinated innerly, with antero-lateral expansions folded externally, slightly asymmetrical as in C. muria. Anterior edge of pseudepiphallic sclerite with hook-like convergent expansions, membrane between them with thin setae. Posterior region rounded, with a small rounded translucent area at apex. Rami strong, with convergent apical stems. Ectophallic arc strong, with a short triangular posterior expansion. Bases of ectophallic apodemes with posterior lobes sclerotized at apex, without anterior sclerotization. Ectophallic fold laterally sclerotized, apex trilobate, median lobe conical and sponge-like. Endophallic sclerite with short lateral arms, with a triangular posterior expansion. Endophallic apodeme with lateral lamellas and a mediodorsal crest. Membrane of endophallic cavity finely plicate.

Female: Unknown.

Juvenile: Unknown.

Measurements. See Table 13.

Habitat and life history traits. Unknown.

Behavior. Unknown.