1. r-m longitudinal, in line with second sector of Rs (Ohakuneinae, subfam. n.).......................................... 2

- r-m transverse or oblique............................................................................................................................. 5

2. First sector of Rs visible............................................................................................................................. 3

- First sector of Rs entirely missing.............................................................................................................. 4

3. M 1+2 complete......................................................................................... (Costa Rica) Rogambara Jaschhof

- M 1+2 incomplete at base............................................................................ (Costa Rica) Cabamofa Jaschhof

4. Rs originating before half of wing; base of M 4 complete........................................................................................................................................................ (Australia, Costa Rica, Chile, Brazil) Colonomyia Colless

- Rs originating beyond distal half of the wing; base of M 4 incomplete............................................................................................................. New Zealand, Australia, Papua New Guinea, Chile) Ohakunea Edwards

5. First sector of Rs short; r-m present............................................................................................................. 6

- First sector of Rs long; r-m absent, tb much longer than m-cu............................................................................................................................. Rangomaraminae (New Zealand) Rangomarama Jaschhof & Didham

6. R 5 running very close to C, first sector of Rs tiny, transverse, r-m scarcely visible; medial fork long, M 1 and M 2 diverging before middle of wing. Tergite 9 devoid of distal projections bearing strong spines.......................................... Heterotrichinae, subfam. n. (Europe and Japan, Baltic amber) Heterotricha Loew

- R 5 not close to C, first sector of Rs normally developed, oblique forward, r-m produced, at least slightly oblique backwards; medial fork short, M 1 and M 2 diverging beyond middle of wing. Tergite 9 bearing 1 or 2 pairs of distal projections with strong spines (Chiletrichinae, subfam. n.)............................................ 7

7. Proboscis long, maxillary palpus displaced to apex............. (South Africa) Rhynchoheterotricha Freeman

- Proboscis short, normal............................................................................................................................... 8

8. M 1+2 /M 3+4 longer than m-cu......................................................................... (Kenya) Kenyatricha Chandler

- M 1+2 /M 3+4 shorter than m-cu......................................................................................................................... 9

9. Rs originating before basal third of wing; R 1 reaching C about at middle of wing; M 1 and M 2 continuously diverging.............................................................................................(New Zealand) Insulatricha Jaschhof

- Rs originating beyond basal third of wing; R 1 reaching C at distal third of wing; M 1 not gradually diverging.............................................................................................................................................................. 10

10. m-cu reaching CuA very basally, basal cell slender..............(Chile, southern Brazil) Chiletricha Chandler

- m-cu reaching CuA more distally, basal cell wide.......................................... (Ecuador) Eratomyia n. gen.