Published May 6, 2021 | Version v1
Taxonomic treatment Open

Convexicaris Schram 1990

  • 1. Centre de Recherche en Paléontologie (CR 2 P, UMR 7207), Sorbonne Université-MNHN-CNRS, Muséum national d'Histoire naturelle, Département Origines et Évolution, case postale 38, 57 rue Cuvier, F- 75231 Paris cedex 05 (France) and Department of Biology, Ludwig-Maximilians-Universität München, Grosshaderner Strasse 2, 82152 Planegg-Martinsried (Germany)
  • 2. Department of Biology, Ludwig-Maximilians-Universität München, Grosshaderner Strasse 2, 82152 Planegg-Martinsried (Germany) and GeoBio-Center, Ludwig-Maximilians-Universität München, Richard-Wagner-Strasse 10, 80333 München (Germany)

Description

Genus Convexicaris Schram, 1990

TYPE SPECIES. — Convexicaris mazonensis Schram, 1990 from Carbondale Formation, Francis Creek Shale Member, Mazon Creek area, Illinois, United States.

DIAGNOSIS (repeated fromSchram 1990). — Sinuously shaped carapace displaying a rounded antero-dorsal part, a convex antero-lateral margin, and a blunt postero-ventral aspect. Elongate subchelate raptorial limbs composed of narrow articles. Body ending by long caudal rami.

Convexicaris mazonensis Schram, 1990 (Fig. 5)

TYPE MATERIAL. — Holotype:PE 32958.— Paratypes:PE 11255, 23525, 38169, 39350, 40076, 45692, 45695; SDSNH 36781; MCP 594.

TYPE HORIZON. — Carbondale Formation, Francis Creek Shale Member, c. 307 Ma, Westphalian D, Middle Pennsylvanian, Carboniferous

TYPE LOCALITY. — Mazon Creek area, Pit 11, Illinois, United States.

EXAMINED MATERIAL. — ROMIP 47987 (lateral view).

DIAGNOSIS. — Same as for genus.

DESCRIPTION

The new specimen provides important details about the trunk, especially about the appendages (Fig. 5 A-D). The shield is longer than high, 2.1× (ls = 16.7 mm; h = 7.85 mm). The posterior trunk is obviously organised into eight discrete segments, especially indicated by eight rectangular appendages emerging from under the shield. The distal end of the appendages appears bifid. The appendages are 1.69 mm to 1.89 mm long.

REMARKS

Trunk appendages have been known for several species of Thylacocephala (Arduini et al. 1980; Secretan 1985; Arduini 1992; Polz 1990, 1994, 2001; Lange et al. 2001; Haug et al. 2014; Ji et al. 2017; Braig et al. 2019; Jobbins et al. 2020). In most cases they appear to be either paddle-like or multiannulated. The morphology observed here, paddle-like, but with a bifid distal end, appears new. Yet, in fact our knowledge of the posterior trunk appendages is rather incomplete. Therefore, this may in fact be a more common feature.

Although Schram (1990) suggested that Co. mazonensis should possess eight posterior trunk segments with swimming appendages, this was not really apparent in his specimens. The new specimen now clearly shows eight distinct such appendages and hence supports the assumption by Schram (1990).

Notes

Published as part of Laville, Thomas, Haug, Joachim T. & Haug, Carolin, 2021, New species of Thylacocephala, Eodollocaris keithflinti n. gen., n. sp., from the Mazon Creek Lagerstätte, Illinois, United States (c. 307 Ma) and redescription of other Mazon Creek thylacocephalans, pp. 295-310 in Geodiversitas 43 (10) on pages 300-301, DOI: 10.5252/geodiversitas2021v43a10, http://zenodo.org/record/4746542

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Linked records

Additional details

Biodiversity

Collection code
MCP , SDSNH , TYPE, MATERIAL
Material sample ID
MCP 594 , SDSNH 36781
Scientific name authorship
Schram
Kingdom
Animalia
Phylum
Arthropoda
Genus
Convexicaris
Taxon rank
genus
Type status
holotype , paratype
Taxonomic concept label
Convexicaris Schram, 1990 sec. Laville, Haug & Haug, 2021

References

  • SCHRAM F. R. 1990. - On Mazon Creek Thylacocephala. Proceedings of the San Diego Society of Natural History 3: 1 - 16
  • ARDUINI P., PINNA G. & TERUZZI G. 1980. - A new and unusual Lower Jurassic cirriped from Osteno in Lombardy: Ostenia cypriformis ng, n. sp. Atti della Societa italiana di scienze naturali e del museo civico di storia naturale di Milano 121: 360 - 370
  • SECRETAN S. 1985. - Conchyliocarida, a class of fossil crustaceans: relationships to Malacostraca and postulated behaviour. Transactions of the Royal Society of Edinburgh: Earth Sciences 76 (2 - 3): 381 - 389. https: // doi. org / 10.1017 / S 0263593300010592
  • ARDUINI P. 1992. - Clausocaris pinnai n. sp. (Order Clausocarida nov.), thylacephalan crustacean from the Norian of the Preone Valley (Udine, N. Italy) and morphological considerations on Thylacocephala. Atti della Societa italiana di Scienze naturali e del Museo civico di Storia naturale di Milano 132 (21): 265 - 272
  • POLZ H. 1990. - Clausocaris lithographica (? Crustacea, Thylacocephala). Archaeopteryx 8: 93 - 109
  • POLZ H. 1994. - Mayrocaris bucculata gen. nov. sp. nov. (Thylacocephala, Conchyliocarida) aus den Solnhofener Plattenkalken. Archaeopteryx 12: 35 - 44
  • POLZ H. 2001. - Dollocaris michelorum, sp. nov. (Thylacocephala, Concavicarida) aus den Solnhofener Plattenkalker. Archaeopteryx 19: 45 - 55
  • LANGE S., SCHRAM F. R., STEEMAN F. A. & HOF C. H. J. 2001. - New Genus and Species from the Cretaceous of Lebanon Links the Thylacocephala To the Crustacea. Palaeontology 44 (5): 905 - 912. https: // doi. org / 10.1111 / 1475 - 4983.00207
  • HAUG C., BRIGGS D. E. G., MIKULIC D. G., KLUESSENDORF J. & HAUG J. T. 2014. - The implications of a Silurian and other thylacocephalan crustaceans for the functional morphology and systematic affinities of the group. BMC Evolutionary Biology 14 (159). https: // doi. org / 10.1186 / s 12862 - 014 - 0159 - 2
  • JI C., TINTORI A., JIANG D. & MOTANI R. 2017. - New species of Thylacocephala (Arthropoda) from the Spathian (Lower Triassic) of Chaohu, Anhui Province of China. PalZ 91 (2): 171 - 184. https: // doi. org / 10.1007 / s 12542 - 017 - 0347 - 7
  • BRAIG F., HAUG J. T., SCHADEL M. & HAUG C. 2019. - A new thylacocephalan crustacean from the Upper Jurassic lithographic limestones of southern Germany and the diversity of Thylacocephala. Palaeodiversity 12 (1): 69 - 87. https: // doi. org / 10.18476 / pale. v 12. a 6
  • JOBBINS M., HAUG C. & KLUG C. 2020. - First African thylacocephalans from the Famennian of Morocco and their role in Late Devonian food webs. Scientific Reports 10: 5129. https: // doi. org / 10.1038 / s 41598 - 020 - 61770 - 0