ANTHOPHORULA (ANTHOPHORULA) COMPLETA (COCKERELL)

Figures 27, 32

DIAGNOSIS: This species can be distinguished from others with projecting salivary lips by the shape of the mandibular apex, both teeth of which are obtusely rounded (unworn) or, when worn, fused as an oblique truncation. The antennal papilla is shorter than its basal diameter, unlike the elongate papilla of Eremapis parvula, and not sharply pointed as in Chilimalopsis parvula and Teratognatha modesta.

Head: Antennal papilla apically rounded, slightly shorter than basal diameter, much shorter than maxillary palpus. Labrum with apical edge weakly bilobed in frontal view; epipharyngeal surface bearing dense patch of long spicules on each side; mandibular corium finely spiculate.

Mandibular apex with dorsal and ventral teeth obtusely rounded; notch separating them shallow; on worn specimen mandibular apex becoming subtruncate, notch becoming even shallower; dorsal apical edge of mandible with moderately small teeth; spines of upper edge of cusp abundant, becoming elongate toward mandibular base; lower edge of cusp with row of approximately 5–7 stout spines, with largest one at leading edge of cusp, smaller spines decreasing in size extending basad and meeting upper edge. Labial palpus nearly twice as long as basal diameter. Salivary lips strongly projecting, their apex much wider than one-half distance between bases of labial palpi in frontal view.

Body: Spiracular atrium consisting of about 12 chambers. Male with transverse median scar on apex of ventral protuberance of abdominal segment 9.

MATERIAL STUDIED: Four postdefecating and two predefecating larvae: AZ: Pima Co., Desert Station E side of Tucson Mts. IV-28-1994 (J.G. Rozen).

REMARKS: Nesting biology of this species is described below.