Eotrogaspidia amans (André, 1909)

(Figs 1B, 1F, 2A, 2E, 5B, 7B, 8 C–D, 10B, 10F, 12A)

Mutilla amans André, 1909: 173, ♂ ♀, lectotype ♀ (“Tandjong Priok”, Java, Indonesia) [RMNH], designated by Mickel (1935: 236).

Timulla (Trogaspidia) amans: Mickel 1935: 220, 226, 236, ♂ ♀.

Timulla (Trogaspidia) amans amans: Pagden 1949: 228, ♂ ♀.

Eotrogaspidia amans amans: Lelej 1996c: 22; Lelej 2005: 77; Lo Cascio 2015: 548; Okayasu et al. 2018: 304, ♀.

Eotrogaspidia amans: Williams et al. 2019a: 32, ♀.

Diagnosis. MALE. Clypeus deeply concave (Fig 5B); scape ventrally with two parallel carinae; F1 weakly compressed, not wider than F2. Pronotum and propodeum dorsally with dense pale setae obscuring cuticular sculpture (Fig. 1B). Metacoxa flattened with dense erect pale microsetae (Fig. 7B). T2 laterally with large dense punctures; T7 with medial triangular impunctate line not forming tubercle posteriorly; cuspis robust, spatulate in inner view (Fig. 10B). FEMALE. T2 pale setal spots large, densely setose, well defined (Fig. 2A); distance between T2 spots 0.3–0.5 × spot diameter; pale setal bands on T3–4 complete (Fig. 2A); pygidial plate entirely with coarser micro-reticulation and with irregular sub-rugae anteriorly (Fig. 12A); lateral margins of pygidial plate convergent posteriorly.

Material examined. 5♂ 2♀. Indonesia: JAVA: 1♂, Bantam Paroaseram, 23.V.1931, Luftnick [RMNH]; 1♂, Buitenzorg, W. Roepke [RMNH]; 2♂, same place, XII.1936, E. Fransen [CSCA, RMNH]; 1♀, Karangandoel, G. Slamat, 150 m, X.1940, F.C. Drescher [RMNH]; 1♂, Malang, IV.1933, Betrem [RMNH]; 1♀, Nglirip, Djati-Bosch by Bodjonegoro, I.1936, Walsch [RMNH].

Distribution. Indonesia: Java, Kangean Islands (Pagden 1949).

Remarks. Morphologically, both sexes of this species are quite similar to E. oryzae, suggesting recent speciation. In males, E. amans and E. oryzae are recognized in the genus by having the following combination of character states: F1 as wide as F2; both pronotum and propodeum clothed with dense appressed golden setae; T7 with longitudinal triangular impunctate line; cuspis widened posteriorly. In females, both species are recognized by having the following combination of character states: T2 with large basal spots of golden setae; both T3 and T4 with complete band of golden setae; pygidial plate entirely micro-reticulate. However, E. amans differs from E. oryzae by having the male T2 laterally with large dense punctures (with small sparse punctures in E. oryzae), female T2 spots densly setose (sparsely setose in E. oryzae), female pygidial plate with coarse microreticulation (with finer microreticulation in E. oryzae) and female pygidial plate with lateral carinae convergent posteriorly (subparallel in E. oryzae). These two species are distributed in allopatry; E. amans has been recorded from Java only, while E. oryzae is widespread in Indochina.