Astenus thoracicus (Baudi di Selve, 1857) (Figs. 55–66)

Sunius thoracicus Baud idi Selve, 1857: 105

Astenus jordanicus Coiffait, 1981: 19, syn. n. (Figs. 67–74)

Type material examined: Astenus jordanicus Coiffait, 1981: 1 ♀, “ Zerkatal b. Romana ”, “ 100 m, 8.1.1958 ”, “O. Jordan. J. Klapperich ”, “ Astenus (Astenognhus) jordanicus Coiffait 1980 ” with red printed label reading “ Allotype ” (MNHNP); 1♂, same data as Allotype, “ Astenus jordanicus Coiffait ” with red printed label reading “ Paratype ” (MNHNP); 1♀, “ Jordan. J. Klapperich ”, “ Wadi Farra, Jordan ” “ 200 m, 16.12.1957 ” with red printed label reading “ Paratype ” (MNHNP).

Additional material examined: GEORGIA: 1♂, Sukhum, Caucasus, 27.X.1911, leg. Zaitsev (ZIN). ISRAEL: 2♂♂, Galilée, Ginosan, 26.V.1973, leg. Löbl (MHNG). LEBANON: 1♂, 1♀, Soum, 29.III.1975, leg. Besuchet (MHNG). RUSSIA: 2♂♂, 1♀, Krasnodar, 23.IV.1976, leg. Korotayev (ZIN). 1♂, Russia, Krasnador Reg., Bolshoi Utrish vill., 1–5.V.2009, leg. Khachikov (AZMM). TURKEY: 1♂, Aksaray, Ekecik Dağı, 38°44’07”N, 34°02’56”E, 1830 m, 23.III.2018, leg. Örgel & Yaman (AZMM). 1♂, 1♀, Aksaray, Melendiz River, 38°12’00”N, 34°19’50”E, 1400 m, 22.III.2018, leg. Örgel & Yaman. (AZMM). 1♂, Ankara, Ayaş 10 km SE, Abdülselam Dağı, 39°56’40”N, 32°22’25”E, 1414 m, 11.III.2018, leg. Örgel & Yaman (AZMM). 1♂, Ankara, Kızılcahamam, Sorgun göleti, 40°19’51”N, 32°13’08”E, 1271 m, 24.IX.2017, leg. Örgel & Yaman (AZMM). 1♀, Ankara, Beypazarı, Kıbrıscık road, 40°20’33”N, 31°56’48”E, 1630 m, 25.IX.2017, leg. Örgel & Yaman (AZMM). 1♂, Çankırı, Kurşunlu 5 km E, Taşkaracalar, 1647 m, 40°42’11”N, 33°19’43”E, 20.V.2018, leg. Örgel & Yaman (AZMM). 1♂, Eskişehir, Sündiken Dağları, 40°00’15”N, 30°57’12”E, 1140 m, 12.III.2018, leg. Örgel & Yaman (AZMM). 1♂, 2♀♀, Karaman, Sertavul Geçidi, 36°53’39”N, 33°17’18”E, 1754 m, 02.IV.2017, leg. Örgel & Yaman (AZMM). 1♀, Kayseri, Yahyalı, 38°03’01”N, 35°24’01”E, 1427 m, 05.VI.2016, leg. Yağmur, Örgel & Yaman (AZMM). 1♂, 4♀♀, Konya, Seydişehir, Geyik Dağları, 37°27’36”N, 31°43’54”E, 1477 m, 15.IV.2016, leg. Örgel & Yaman (AZMM). 1♂, 1♀, Kırşehir, Kurtbeyliyeniyapan 2 km W, 39°15’34”N, 33°58’45”E, 1590 m, 27.III.2019, leg. Örgel & Köksal (AZMM). 3♂♂, 2♀♀, Kırşehir, Akçakent, Yaylaözü 2 km NE, 39°37’33”N, 34°12’00”E, 1512 m, 01.V.2019, leg. Örgel & Köksal (AZMM). 1♂, Konya, Hadim, Beyreli Yolu, 36°53’01”N, 32°25’10”E, 1933 m, 24.V.2016, leg. Örgel & Yaman (AZMM). 1♀, Konya, Bozkır, Tanrıdağı Tepesi, 36°58’19”N, 32°08’58”E, 2015 m, 23.V.2016, leg. Örgel & Yaman (AZMM). 3♂♂, 2♀♀, Konya, Beyşehir, Geyik Dağları, 37°25’36”N, 31°43’00”E, 2000 m, 21.V.2016, leg. Örgel & Yaman (AZMM). 2♂♂, Nevşehir, Hacıbektaş, 38°56’22”N, 34°37’40”E, 1210 m, 26.III.2018, leg. Örgel & Yaman. (AZMM). 1♂, Izmir, Karaburun 5 km SW, 01.V.2013, leg. Anlaş & Yağmur (AZMM). 4♂♂, 5♀♀, Izmir, Nif Dağı, 38°23’”N, 27°21’”E, 1200–1500 m, 16.III.2014, 05.V.2014, leg. Anlaş, Yağmur & Örgel (AZMM). 1♂, Izmir, Buca, Kaynaklar, 03.III.2013, leg. Yağmur (AZMM). 5♂♂, 3♀♀, Izmir, Bozdağlar, leg. Özgen, 11.IV.2014, Yağmur & Örgel (AZMM). 1♂, Erzincan, Üzümlü, Küçük Sarıkaya creek bank, 1713 m, 39°14’20’’N, 39°50’02’’E, 18.V.2011, leg. Anlaş, Özgen & Khachikov (AZMM). 1♂, Muş, Hasköy, Elmaçulak, 15.V.2013, leg. Gündüz (AZMM). 1♀, Aydın, Imambaba Tepesi, 37°56’06”N, 27°53’56”E, 1644 m, 24.III.2014, leg. Anlaş & Örgel (AZMM). 1♂, 1♀, Aydın, Çine 10 km S, 30.IV.1975, leg. Besuchet & Löbl (MHNG). 1♂, Denizli, Çal, Ayaz Tepesi, 38°02’00”N, 29°21’25”E, 1365 m, 01.V.2013, leg. Yağmur & Örgel (AZMM). 2♂♂, 1♀, Denizli, Çal, Erikli Mevkii, 38°02’45”N, 29°22’31”E, 1382 m, l 01.V.2013, eg. Yağmur & Örgel (AZMM). 1♂, 1♀, Denizli, Karcı Dağı, 37°44’53”N, 28°58’56”E, 724 m, 12.X.2013, leg. Özgen & Örgel (AZMM). 1♂, Manisa, Turgutlu, Baktırlı 2 km W, 38°25’38”N, 27°52’44”E, 780 m, 20.III.2013, leg. Anlaş, Yağmur & Örgel (AZMM). 1♂, Manisa, Spil Dağı Dağı Milli Parkı, 1200 m, 38°33’20’’N, 27°23’17’’E, 27.IX.2009, leg. Anlaş (AZMM). 2♂♂, 2♀♀, Spil Dağı, 38°32’57”N, 27°27’06”E, 1274 m, 15.III.2014, leg. Yağmur & Örgel (AZMM). 1♂, Manisa, Selendi, Omurlar 38°02’45”N, 29°22’31”E, 1382 m, 14.III.2014, leg. Yağmur & Örgel (AZMM). 1♂, 1♀, Denizli, Karcı Dağı, 37°44’53”N, 28°58’56”E, 724 m, 12.X.2013, leg. Özgen & Örgel (AZMM). 1♂, Manisa, Turgutlu, Baktırlı 2 km W, 38°25’38”N, 27°52’44”E, 780 m, 20.III.2013, leg. Anlaş, Yağmur & Örgel (AZMM). 1♂, Manisa, Spil Köyü, 38°52’15”N, 28°49’11”E, 974 m, 27.IX.2009, leg. Yağmur & Örgel (AZMM). 5♂♂, 2♀♀, Manisa, Soma, Yağcılı, 39°20’05”N, 27°40’23”E, 306 m, 07.IV.2015, leg. Anlaş, Yağmur & Örgel (AZMM). 5♂♂, 2♀♀, Soma, Tabanlar, 39°19’47”N, 27°44’17”E, 682 m, 07.IV.2015, leg. Yağmur & Örgel (AZMM). 3♂♂, 2♀♀, Analya, Kemer, Üçoluk, 36°38’50”N, 30°26’02”E, 1078 m, 25.III.2017, leg. Kunt (AZMM). 2♂♂, Antalya, Serik, Hapıllar Köyü, 30.I.2016, leg. Örgel (AZMM). 1♂, Kütahya, Beşdeğirmen, 961 m, 39°31’18”N, 30°53’28”E, 17.V.2013, leg. Örgel (AZMM). 2♂♂, 1♀, Kütahya, Simav, Esenbağ, 39°23’53”N, 29°00’16”E, 1012 m, 14.IV.2014, leg. Özgen & Örgel (AZMM). 1♂, 1♀, Muğla, Bozburun 3 km N, 36°41’25”N, 28°05’01”E, 215 m, 05.IV.2013, leg. Yağmur & Örgel (AZMM). For additional material examined see also in Anlaş (2017).

Distribution: A. thoracicus is a widespread species in the Eastern Mediterranean and adjacent regions. It is distributed in Georgia, Azerbaijan, Greece, Cyprus, Bulgaria, Italy, Canary Islands, Iran, Syria, Israel, Uzbekistan and Turkey. The records from Lebanon and Russia are reported for the first time.

Comments: Coiffait (1981) described A. jordanicus based on a male (holotype) from Zerkatal b. Romana (Zarka Valley near Rumana), four males and three females (paratypes) from Jordan Valley, Wadi Farra in Jordan. According to Coiffait (1981), type series of the species were deposited in the author`s collection in MNHNP and HNHM. I visited both museums, but I could not find any type specimen of A. jordanicus in HNHM. An Allotype, one male and one female (paratypes) of A. jordanicus were studied in the collection of H. Coiffait in MNHNP. Coiffait (1981) did not mention the differences of this species from other Astenus species in the original description of A. jordanicus.

A. thoracicus is probably the most common species of the genus in the East Mediterranean. Based on the original description of A. jordanicus (Figs. 67–74) and the illustrations of the aedeagus in Coiffait (1981: Figs. 1 G–H), the external morphology and aedeagal characters are found within the range of intraspecific variation of A. thoracicus (Figs. 55–66). This species is here placed in the synonymy of A. thoracicus.