20. Bagous frit group

Diagnosis. Apical margin of pygidium deeply emarginate (char. 55.2). Inner surface of dorsal process covered with a dense area of denticles (char. 100.1). Sublateral margin of penis body behind dorsal process with dorsal, sublateral, carinate, V-shaped, denticulate ridge bordering median depression behind dorsal process (char. 106.2). Lateral margin laterad and distad of orifice denticulate (char. 66.1). Penis body in cross-section T-shaped, the sides deeply compressed behind dorsal process and almost joined at midline (char. 58.1). Sublateral margin of penis body behind dorsal process denticulate (char. 105.1). Apical third of penis body asymmetrically deflected to left and asymmetrically flattened (char. 61.1).

Remarks. This group, composed of the European species B. frit and the Japanese species B. fritodes, is characterized by several synapomorphies, especially in the male genitalia. Concerning character state 57.1, this must not be confused with a slight left deflection of the strict apex present in several unrelated species. B. frit and the species belonging to the previous group formed a monophyletic clade, supported in BI and ML (respectively, 69% pp and 57% bs) and not supported in MP; relationships between these groups were also suggested by the molecular analysis, where the reciprocal position of B. frit and B. limosus was, however, reversed.

Species included. PAL: §*¹ Bagous frit (Herbst, 1795), ¹ B. fritodes O'Brien & Morimoto, 1994.