Willema Grishin, new genus

http://zoobank.org/ D92BF2E1-F842-44C6-A924-023F495F9589

Type species. Heteropterus willemi Wallengren, 1857.

Definition. A sister genus to Hovala Evans, 1937 (type species Cyclopides pardalina Butler, 1879), more distant from Metisella Hemming, 1934 (type species Papilio metis Linnaeus, 1764), where it was placed previously (Fig. 8). Keys to IV.20.B in Evans (1937). Shares many features with Metisella, such as hindwing discal cell longer than half of the wing, forewing veins R 1 and R 2 very near each other or touching (not separated). Distinguished from its relatives by the pattern of hindwing underside, which is with dark web-like pattern along veins with some streaks across and orange, ochreous, yellow or white spots. Hindwing below pale-yellow with dark web-like pattern along veins with some streaks across, but without darker markings. Genitalia with narrow and long penis, uncus with tegumen bulbous in dorsal view, uncus narrows to a point (except in W. willemi where distal end is mushroom-shaped and base of uncus with side processes), gnathos better developed than in Metisella, ampulla separated from harpe, protruding beyond its caudal end (except in W. willemi where harpe upturned embracing ampulla). In DNA, a combination of the following base pairs is diagnostic: aly363.37.1:A929C, aly 1450.10.1:A576G, aly283.4.1:T1129C, aly3555.2.2:T116C, aly451.23.2:C1177A.

Etymology. The name is a feminine noun in the nominative singular, derived from the type species name.

Species included. The type species, Cyclopides formosus var. tsadicus Aurivillius, 1905, Cyclopides angolana Karsch, 1896, Metisella kumbona Evans, 1937, Cyclopides carsoni Butler, 1898, Heteropterus formosus Butler, 1894, and Cyclopides perexcellens Butler, 1896.

Parent taxon. Tribe Heteropterini Aurivillius, 1925.