Published January 13, 2020 | Version v1
Taxonomic treatment Open

Dichotomius (Cephagonus) mundus

Description

1.2.: Dichotomius (Cephagonus) mundus (Harold 1869) (Figure 17)

Pinotus mundus Harold 1869 ı p. 136 (description)

Pinotus singularis Felche 1907 ı p. 278 (description) new synonym

Pinotus simulans Luederwaldt 1925 ı p. 68 (description) new synonym

Pinotus (Cephagonus) simulans ı Luederwaldt 1929 ı p. 111

Pinotus (Cephagonus) singularis ı Luederwaldt 1929 ı p. 112

Pinotus (Cephagonus) simulans ı Luederwaldt 1931 ı p. 358

Pinotus (Cephagonus) mundus ı Luederwaldt 1931 ı p. 358

Pinotus (Cephagonus) simulans ı Luederwaldt 1936 ı p. 214

Pinotus (Cephagonus) singularis ı Luederwaldt 1936 ı p. 215

Pinotus (Cephagonus) mundus ı Luederwaldt 1936 ı p. 215

Pinotus mundus ı Blackwelder 1944 ı p. 207

Pinotus simulans ı Blackwelder 1944 ı p. 207

Pinotus singularis ı Blackwelder 1944 ı p.207

Dichotomius (selenocopris) simulans ı Vaz-de-Mello 2000 ı p. 193

Dichotomius (selenocopris) singularis ı Vaz-de-Mello 2000 ı p. 193

Dichotomius (selenocopris) mundus ı Vaz-de-Mello 2000 ı p. 193

Diagnosis. D. mundus is distinguished from other species in the group by the following combination of characters: pronotal disc and elytra dorsal surface with microsculpture (chagrinated) (as D. carbonarius of the subgenus selenocopris); metasternum glabrous or with few setae; larger males bearing a single emargination on the anterior portion of pronotal disc and basal excavation on males paramera (dorsal view) never reaching paramera sides (Figure 17 (d-f)).

LECTOTYPE of Pinotus mundus (maleı here designated) (Figure 17 (g)): Brasilia. P. mundus Harold {white with red marginsı handwritten}/Ex. Musaeo Harold {white with black marginsı printed}/Museum Paris ex Coll. R. Oberthur {greenı printed} [MNHN];

PARALECTOTYPE of Pinotus mundus (female): munda m. {whiteı handwritten}/Ex. Musaeo Harold {white with black marginı printed}/Museum Paris ex Coll. R. Oberthur {greenı printed} [MNHN].

HOLOTYPE of Pinotus singularis (female): singularis Felsche. N. Brasilien {white with purple marginı handwritten}/Para {greenı printed}/Typus {redı printed}/Coll. C. Felsche Kauf 26ı 1918 {greenı printed}/Staatl. Musum für Tierkunde Dresden {whiteı printed}/ HOLOTYPE {red with black marginsı printed} [at SNSD].

LECTOTYPE of Pinotus simulans (maleı here designated): Rio de Janeiro. Serra de Macahé. 14.624 {whiteı partially handwritten}/COTIPO {red with black marginsı printed}/ Pinotus simulans Lüd Det. 25 {whiteı handwritten}/25959 {whiteı printed} [at MZSP].

Non type-material. BRASIL: Rio de Janeiro: Nova Friburgo. XI-1993. E Grossi [1 Ƌ at CEMT]; same municipality but i-1995. FZ Vaz-de-Mello [1 ♀ at CEMT]; same municipality but XI-1998. P & E Grossi [2 Ƌ. 2 ♀ at CEMT]; Macaé de Cima. 1500 m. III-2000. C Lopes- Andradeı Gumier & FZ Vaz-de-Mello [2 ƋƋı 3 ♀♀ at CEMT]; same data but xii-2005. E &

P Grossi [2 ƋƋ at CEMT]; São Paulo: São Luis do Paratininga. Pq. Estadual da Serra do Mar Núcleo Sta. Virgínia. xii-2004. M Uehara [1 Ƌ at CEMT].

Distribution. Above 900 metres in the Atlantic Forest of Rio de Janeiro and São Paulo states (SEE RJı ESı ETC). The type-locality of Felsche s D. singularis is Pará (state of Pará) but we consider it as doubtful since all other analysed specimens occur above 900 m and only in the states of Rio de Janeiro and São Paulo (Figure 13).

Remarks. Pinotus simulans Lectotype is the most developed male of Luederwaldt s syntype series. Harold s (1869) description emphasises the dorsal surface with microsculpture (chagrinated)ı which is the main diagnostic character of this species in the group. Felsche (1907) used specimens with a malformation on the clypeal margin to describe D. singularis. This malformation results in dorsally visible maxillary palpi 2 4 (Figure 11 (a)) and is relatively constantly observable among other Scarabaeinae. Dorsal microsculptureı metasternum almost glabrous and the female sixth ventrite shape are evidence for this new synonym with Harold s species. Luederwaldt (1926) described Pinotus simulans with a large series of males and females from Nova Friburgo municipality (state of Rio de Janeiro). The author probably did not have access to Harold s type materialı but years laterı Luederwaldt (1931) comments on the strong resemblance of Pinotus simulans with P. mundus.

Notes

Published as part of Nunes, Rafael V. & Vaz-de-Mello, Fernando Z., 2019, Taxonomic revision of Dichotomius (Cephagonus) Luederwaldt 1929 and the taxonomic status of remaining Dichotomius Hope 1838 subgenera (Coleoptera: Scarabaeidae: Scarabaeinae: Dichotomiini), pp. 2231-2351 in Journal of Natural History 53 (37) on pages 2286-2290, DOI: 10.1080/00222933.2019.1692088, http://zenodo.org/record/3665519

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Linked records

Additional details

Biodiversity

Scientific name authorship
Harold
Kingdom
Animalia
Phylum
Arthropoda
Order
Coleoptera
Family
Scarabaeidae
Genus
Dichotomius
Species
mundus
Taxon rank
species
Taxonomic concept label
Dichotomius (Cephagonus) mundus (Harold, 1869) sec. Nunes & Vaz-de-Mello, 2019

References

  • Harold E. 1869. Catalogus Coleopterorum Hucusque Descriptorum Synonymicus et Systematicus. Vol. 4. Munich: E. H. LGummi.
  • Luederwaldt H. 1925. Neue Pinotus arten. Bol Do Museu Nacional. 2 (1): 67 - 69.
  • Luederwaldt H. 1929. As especies brasileiras do genero. Pinotus. Rev Museu Paulista. 16: 603 - 776.
  • Luederwaldt H. 1931. Novos subsidios para o conhecimento do genero. Pinotus. Rev Entomol. 1: 297 - 304.
  • Luederwaldt H. 1936. Notas complementares " As especies Brasileiras do genero Pinotus ". Rev Museu Paulista. 20: 207 - 216.
  • Blackwelder RE. 1944. Checklist of the Coleopterous insects of Mexicoi Central Americai the West Indies and South America. Nat Hist Mus Bull. 185: 189 - 341.
  • Vaz-de-Mello FZ. 2000. Estado atual de conhecimento dos Scarabaeidae s. str. (Coleoptera: Scarabaeoidea) do Brasil. In: Martin-Piera Fi Morrone JJi Melic Ai editors. Hacia un projecto CYTED para el inventario y Estimacion de la Diversidad Entomologica en IberoAmerica: PrIBES- 2000. Zaragoza: Monografias Tercer Milenio; p. 183 - 195.
  • Felsche C. 1907. Coprophage Scarabaeiden (Col.). Dtsch Entomol Z. 3: 273 - 296.
  • Luederwaldt H. 1926. Novas especies do genero " Pinotus ". Rev Museu Nacional. 14: 135 - 138.