Fig. 5. Results of BEAST and LAGRANGE analyses, and comparative morphology of dolichoderine wing venation. A, Chronogram of the Dolichoderinae, Leptomyrmex relictus sp.n. bolded; B, L. burwelli: complete (unreduced) wing venation; C, Chronoxenus javanus: loss of crossvein 2rs-m with retention of other veins; D, Dorymyrmex insanus (male): loss of Rsf2-3 in addition to loss of other abscissae; E, L. relictus sp.n.: loss of Mf2-3; F, L. fragilis: loss of Mf2-3, 2r-rs (note: pterostigmal appendage not illustrated); G, †L. neotropicus: loss of Rs + M, Mf2-3, 2rs-m, Mf4+, 2r-rs; H, †Leptomyrmula maravignae: loss of Mf2+, 2rs-m, 2r-rs; I, †Usomyrma mirabilis: loss of Rs + M with retention of 1m-cu and loss of Mf2-3. In part A blue bars are 95% confidence intervals; coloured circles near nodes are inferred inherited ranges, with the top half corresponding to the upper branch and bottom half with the lower branch; coloured circles next to terminals represent biogeographical coding; asterisks indicate calibrated nodes; bracketed letters to the right of terminal names indicate wing venation reduction pattern occurrence, with letters corresponding to the wing patterns B–F. Note that 1m-cu presence is variable for most taxa displaying loss pattern E, and some Linepithema species have lost Mf2-3. Figures H and I redrawn from Emery (1891) and Dlussky et al. (2014), respectively. Palaeo., Palaeocene; Oligo., Oligocene; Pl., Pliocene; Q, Quaternary.