Three new species of anthericolous smut fungi on Caryophyllaceae

Based on molecular phylogenetic analyses and morphological studies, three new cryptic species of
Microbotryum, M. silenes-dioicae on Silene dioica, M. shykoffianum on Dianthus sylvestris, and M. carthusianorum on Dianthus carthusianorum, are described and illustrated. For Ustilago superba on Dianthus superbus, a new combination in Microbotryum, M. superbum, is proposed.