Kelneria erroris Pełczyńska, Krzemiński & Soszyńska sp. nov.

Figures 8, 9, 10

Material.

Holotype: MALE, preserved in a 15 × 10 × 3 mm piece of Baltic amber (GZG.BST.03049) housed in the Geowissenschaftliches Zentrum, Universität Göttingen, Göttingen, Germany (GMUG) (Figs 8 A, 8 B, 19 [IR spectrum]).

Diagnosis.

Antennae approx. 0.7 × wing length, with distinctly separated flagellomeres densely covered with very short microtrichia; palpus small, with apical palpomere subequal in length to the preceding one and shorter than first flagellomere; Sc ending in C distinctly before level rb cell tip; m – cu joining cubital after level of Rs base; R 2 + 3 short, approx. 0.2 length of R 2 + 3 + 4 + 5 fork stem; epandrium with deep, triangular notch, bearing large, triangular lateral processes with very short, dense apical setation, terminating approx. at the level of gonopods apices. Female unknown.

Type locality and age.

Late Eocene (Priabonian, c. 36–35 Ma), Baltic amber (Sambian Peninsula), Kaliningrad Oblast, Russia.

Description.

Body (Fig. 8 A): approx. 2.2 mm long; wing 1.9 mm long; antennae 1.3 mm long. — Head (Fig. 9 A): wider than long; eyes large, well separated, occupying most of the lateral part of head capsule, distinct cerebral sclerite present, rounded posteriorly, projected posteriorly, dorsal surface bearing several setae approximately half the length of the setae on the scutum; three ocelli present, forming triangle, median ocellus not reduced, lateral ocelli situated on the dorsal surface of the cerebral sclerite, not on its margin. — Antennae (Fig. 9 A): with 2 + 14 segments; 0.7 × wing length in male, scapus annular in shape, wider than long; pedicel bigger than scape, as long as broad, globular in shape, about 1.7 × wider than first flagellomere; flagellum with 12 distinctly separated segments, densely covered with long microtrichia; first flagellomere approximately 2.7 × as long as broad; subsequent flagellomeres progressively decreasing in length, except for the terminal one, which is longer than the preceding one, terminating in a small apiculus. — Mouthparts (Fig. 9 A): palpi small; three apical palpomeres visible; all longer than broad; apical and subapical palpomeres subequal in length; approx. 0.7 × length of first flagellomere. — Wing (Fig. 9 B, C): broad, 2.3 × longer than wide, membrane hyaline without microtrichia and any visible markings; C with microtrichia throughout length; microtrichia visible on dorsal surface of all veins except transverse ones; C terminates at tip of wing, after end of R 4 + 5, on approx. two third of distance between end of R 4 + 5 and M 1; Sc very short, ending in C distinctly before the level of tip of rb cell; R 1 ending in C before half-length of wing, approx. at level of M 1 + 2 fork, just before the level of Cu termination; R 2 + 3 approx. 0.2 the length of R 2 + 3 + 4 + 5 fork stem; frm ending before level at which A 1 reaches wing margin; a faint trace of Mb is present, dividing the basal cell into two; M 1 + 2 fork stem approx. 4.5 × longer than frm, ending just before level of Cu termination; M 1 approx. 3.1 × longer than M 1 + 2 fork stem; m 2 cell opening 1.3 × wider than opening of m 1 cell; M 3 + 4 base weakened and widely interrupted; m 3 + 4 cell opening 0.9 × opening of m 2 cell; basal part of M 1 + 2 distinct; m – cu distinct, joining cubital vein after level of Rs base; Cu reaching wing margin; A 1 termination not visible, as wing is folded; A 2 absent. — Thorax (Fig. 9 A): higher than long; scutum weakly convex, densely covered with long setae arranged in two dorsocentral and two acrostichal rows; scutellum with a row of very long trichia along its margin; anepisternum narrowed dorsally, higher than long, with at least three setae on the dorsoanterior part; katepisternum higher than long, bare; anepimeron reaching the ventral margin of the pleura, bare; laterotergite and mediotergite bare. — Legs (Figs 8 A, 10 B – D): fore coxa the longest, sparsely setulose on anterior surface; hind coxa the shortest, with a several setae posteroexternally; femora irregularly covered with thin setae; tibiae covered with short, thin, irregularly arranged setae on whole surface; fore tibiae with a sensory pit and a single spur, 1.6 × longer than the apical width of the tibia; mid and hind tibiae without sensory pits, bearing two long spurs, subequal in length, (approx. 2.4 × longer apical width of tibia); claws short; empodium big, longer than claws. — Abdomen (Fig. 8 A): covered with long, thin setae, all eight segments visible; segment I short, segment II the longest, subsequent segments progressively decreasing in length. — Male genitalia (Figs 10 A – D): epandrium with deep, triangular notch; bearing large, triangular lateral processes with very short, dense apical setation, terminating approx. at level of gonopods apices; gonocoxites short, broad, triangular, partially fused, oriented parallel to each other; gonostyli strongly reduced, appearing as two small lobes at outer angles of gonocoxites; cerci not visible; phallosome bilobed, strongly sclerotized.

Etymology.

The species epithet erroris refers to a taxonomic error whereby the holotype of this species was previously included in the type series of K. abundare.

Remarks.

Crossvein sc – r is not illustrated in the wing drawing, as its likely positioned close to the wing base and cannot be discerned in the holotype; shape of the anal angle of the wing and apical tip of vein A 1 is hypothesized and indicated by a dashed line, as anal field of the wing of the holotype is folded; specimen designated herein as the holotype originates from the type series of K. abundare as defined by Meunier (1904); this misinterpretation was subsequently retained in the generic revision by Matile (1979) and is corrected in the present study.