Kelneria rovnensis Pełczyńska & Perkovsky sp. nov.

Figures 3, 4, 5, 6, 7

Material.

Holotype: MALE, preserved in an 11 × 10 × 4 mm piece of Rovno amber (SIZK K-32069) housed in the Schmalhausen Institute of Zoology, National Academy of Sciences of Ukraine, Kyiv, Ukraine (SIZK); (Figs 3 A, 3 B, 19 [IR spectrum]). Syninclusions: SIZK K-32066 – K-32068 (Chelonariidae, Formicidae, Nematocera, stellate hairs).

Type locality and age.

Late Eocene (Priabonian, c. 36–35 Ma), Rovno amber (Volhynian Uplift), Pugach quarry, Klesov, Rovno Oblast, Ukraine.

Diagnosis.

Antennae approx. 0.8 × wing length, with distinctly separated flagellomeres densely covered with relatively long microtrichia; pedicel subequal in length to scape; palpus very large, with apical palpomere distinctly elongated, longer than subsequent one and longer than first flagellomere; Sc ending in C approx. at level of rb cell tip; m – cu joining cubital vein before level of Rs base; R 2 + 3 approx. 0.4 length of R 2 + 3 + 4 + 5 fork stem; epandrium deeply and broadly notched, bearing long, slender, spinulose lateral processes extending slightly beyond apices of gonopods. Female unknown.

Description.

Body (Fig. 3 A): approx. 2.5 mm long; wing 1.9 mm long; antennae 1.5 mm long. — Head (Fig. 4 A): wider than long; eyes large, well separated, occupying most of lateral surface of head capsule; distinct cerebral sclerite present, posteriorly rounded, with dorsal surface bearing dense, thick setae; ocelli present, lateral ocelli situated on dorsal surface of cerebral sclerite rather than at its margin; position of median ocellus obscured by antennae. — Antennae (Figs 3 A, 4 A): with 2 + 14 segments; 0.8 × wing length in male, scapus annular in shape, approx. as wide as long; pedicel slightly broader but approx. equal in length to scape, globular in shape, about 1.8 × wider than first flagellomere; flagellum with 12 distinctly separated segments, covered densely with relatively long microtrichia; first flagellomere approximately 3.2 × as long as broad; subsequent flagellomeres progressively decreasing in length, except the terminal one, which is longer than preceding segment, terminating in a thin apiculus. — Mouthparts (Fig. 4 A): palpus large, 1 + 4 segmented; small palpiger visible; all four maxillary palpomeres longer than broad; second and third palpomeres subequal in length; apical palpomere distinctly elongated, 2.6 × longer than the preceding one and 1.5 × longer than the first flagellomere; labella very large, well developed, approx. as long as the apical palpomere. — Wing (Fig. 4 B, C): broad, 2.2 × longer than wide, membrane hyaline without microtrichia and any visible markings; C with microtrichia throughout length; microtrichia visible on dorsal surface of radial and medial veins; C terminates at tip of wing, after end of R 4 + 5, on approx. two third of distance between end of R 4 + 5 and M 1; Sc ending in C approx. at the level of tip of rb cell; sc-r present on approx. half of the Rb length; R 1 ending in C approx. half-length of wing, just after level where M 1 + 2 forks, approx. at the level of Cu termination; R 2 + 3 approx. 0.4 the length of R 2 + 3 + 4 + 5 fork stem; frm ending just after level at which A 1 reaches wing margin; a faint trace of Mb is present, dividing the basal cell into two; M 1 + 2 fork stem approx. 4.8 × longer than frm, ending just before level of Cu termination; M 1 approx. 3.2 × longer than M 1 + 2 fork stem; m 2 cell opening 1.3 × wider than opening of cell M 1; M 3 + 4 base weakened; m 3 + 4 cell opening 1.3 × wider than opening of m 2 cell; basal part of M 1 + 2 distinct; m – cu distinct, joining cubital vein before level of Rs base; Cu reaching wing margin; A 1 ending on wing margin; A 2 absent. — Thorax (Fig. 4 A): higher than long; scutum weakly convex, densely covered with long, thick setae; scutellum with a row of very long trichia along its margin; anepisternum narrowed dorsally, higher than long, with few setae in the dorsoanterior part; katepisternum higher than long, bare; anepimeron reaching the ventral margin of the pleura, bare; laterotergite and mediotergite bare. — Legs (Fig. 5 B, C): fore coxa the longest, with long setae covering entire anterior surface; mid coxa with a several setae anteroapically and with at least two setae anteroexternally; hind coxa the shortest, with a several setae posteroexternally; femora densely and irregularly covered with thin setae; tibiae covered with short, thin, irregularly arranged setae on whole surface, additionally on posterior surface of hind tibia visible row of thicker and more robust setae; fore tibiae with a sensory pit and a single spur, more than 1.5 × longer than the apical width of the tibia; mid and hind tibiae without sensory pits, bearing two long spurs, subequal in length, (spurs 1.8 and 2.9 × longer apical width of tibia, respectively); claws short; empodium big, longer than claws. — Abdomen (Fig. 3 A): densely covered with long setae, all eight segments visible; segment I short, segment II the longest, segment III and IV subequal in length; subsequent segments progressively decreasing in length. — Male genitalia (Figs 5 A, 5 D – G, 6, 7): epandrium deeply and broadly notched, bearing slender, spinulose lateral processes terminating just before apices of gonopods, several apical setae present; gonocoxites short, broad, fused, oriented parallel to each other; gonostyli strongly reduced, appearing as two small lobes at outer angles of gonocoxites; cerci small; not exceeding length of epandrial processes, with visible marginal setation; anal cone prominent; paired lateral parameres present on either side of aedeagus; phallosome broadened; dorsoventrally flattened, strongly sclerotized.

Etymology.

The species epithet rovnensis refers to Rovno region of Ukraine, from which the holotype originates.

Remarks.

The specimen was found in the clear piece of amber with weight 4 g and size 38 × 10 × 25 mm; distinction between gonostyli and gonocoxites is unclear; shape of the anal angle of the wing is hypothesized and indicated by a dashed line, as the anal field of the wing of the holotype is folded.