Caillardia sueva Burckhardt, Serbina & Kotthoff sp. nov.

Fig. 1, Suppl. materials 1, 2

Etymology.

From the Latin adjective suevus = Swabian, referring to its provenience from Swabia.

Holotype.

Male: SMNS -P-108003-1.

Type locality.

Germany, Baden-Württemberg, Randeck Maar.

The preservation of the specimen is relatively good considering the small size of the specimen (see below). The right forewing is probably slightly deformed, while the left forewing seems to be non-deformed.

Sediment.

Dark grey limestone.

Description.

General colour dark brown. Thorax with small light dots dorsally. Protibia pale yellow; metafemur, metatibia and perhaps metatarsus dark brown. Forewing with dark brown veins; membrane light, densely covered with dark brown dots which are slightly more spaced in basal half and denser and partly confluent in apical half; with light area in cell cu 2 adjacent to vein Cu 1 b. Hindwing light.

Vertex subrectangular, about twice as wide as long. Pronotum about two thirds as long as vertex along medial longitudinal body axis. Forewing destroyed along fore margin in the middle, broad, about twice as long as wide, irregularly oval; widest in the middle; apex broadly rounded; pterostigma not visible; basal two thirds of vein C + Sc weakly curved; vein R slightly shorter than M + Cu; vein Rs almost straight; vein M about as long as M 1 + 2; vein M 1 + 2 weakly curved; cell m 1 long and broad; vein Cu 1 a weakly curved in basal two thirds, strongly curved apically; vein Cu 1 b short, almost straight; cell cu 1 relatively long and low. Male subgenital plate, in ventral view, relatively massive, as long as wide.

Measurements.

Head width 0.57 mm; forewing length 1.46 mm.

Comments.

Within Caillardia, C. sueva sp. nov. resembles the males of C. maroccana Loginova, 1972 and C. springatei Spodek & Burckhardt, 2017 in the oval, apically broadly rounded forewing densely covered with dark dots (Burckhardt 1989; Spodek et al. 2017). Based on these characters, the three species make up Loginova’s (1978) species-group 1. Caillardia sueva sp. nov. shares with C. springatei the relatively straight vein Rs. Caillardia sueva sp. nov. differs from the two species in the vein C + Sc of the forewing which is weakly curved rather than straight, vein M which is about as long as M 1 + 2 rather than distinctly longer, and the longer and lower cell cu 1.

The assignment of C. sueva sp. nov. to Caillardia is mostly for its resemblance to C. maroccana and C. springatei. The morphological characters used by Burckhardt and Queiroz (2013) in their cladistic analysis and identification key concern characters not visible in the fossil specimen at hand: frons, metacoxa, metatibia and inclination of the head relative to the longitudinal body axis.