Published May 21, 2026 | Version v1
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Figure 7 from: Jablonski D, Tillack F, Mahlow-Tillack K, Petzold A, Wilzo M, Das A, Idrees M, Baniya CB, Masroor R, Hofmann S (2026) Integrative taxonomy reveals previously undescribed diversity within the Gloydius himalayanus complex (Squamata, Viperidae, Crotalinae) from the Himalaya and Hindu Kush. ZooKeys 1280: 83-153. https://doi.org/10.3897/zookeys.1280.182768

  • 1. Department of Zoology, Comenius University in Bratislava, Bratislava, Slovakia
  • 2. Museum für Naturkunde, Leibniz-Institut für Evolutions- und Biodiversitätsforschung, Berlin, Germany
  • 3. Museum für Naturkunde, Leibniz-Institut für Evolutions- und Biodiversitätsforschung, Berlin, Germany|Institute for Biochemistry and Biology, University of Potsdam, Potsdam, Germany
  • 4. Institute for Biochemistry and Biology, University of Potsdam, Potsdam, Germany
  • 5. Wildlife Institute of India, Chandrabani, India
  • 6. Department of Zoology, University of Peshawar, Peshawar, Pakistan
  • 7. Central Department of Botany, Tribhuvan University, Kathmandu, Nepal
  • 8. Zoological Sciences Division, Pakistan Museum of Natural History, Islamabad, Pakistan
  • 9. Museum Koenig, Leibniz Institute for the Analysis of Biodiversity Change, Bonn, Germany

Description

Figure 7 Principal Component Analysis (PCA) of osteological differentiation among species of the Gloydius himalayanus complex. A. Scatterplot of the first two principal components (PC1 and PC2) based on osteological ratio traits. Points represent individual specimens, colour-coded by species (Figs 1, 2), and convex hulls (shaded polygons) indicate the overall spread of each taxon in morphospace. Dashed arrows are included only as visual guides to displaced labels and have no analytical meaning. Axis labels indicate the percentage of variance explained by each principal component; B. Geographic distribution of sampled populations included in the osteological analysis across Pakistan, India, and Nepal. The inset shows the regional context within Central and South Asia. The map was produced in QGIS 3.28.

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Journal article: 10.3897/zookeys.1280.182768 (DOI)
Journal article: https://zenodo.org/record/20352597 (URL)