Published May 21, 2026 | Version v1
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Figure 2 from: Jablonski D, Tillack F, Mahlow-Tillack K, Petzold A, Wilzo M, Das A, Idrees M, Baniya CB, Masroor R, Hofmann S (2026) Integrative taxonomy reveals previously undescribed diversity within the Gloydius himalayanus complex (Squamata, Viperidae, Crotalinae) from the Himalaya and Hindu Kush. ZooKeys 1280: 83-153. https://doi.org/10.3897/zookeys.1280.182768

  • 1. Department of Zoology, Comenius University in Bratislava, Bratislava, Slovakia
  • 2. Museum für Naturkunde, Leibniz-Institut für Evolutions- und Biodiversitätsforschung, Berlin, Germany
  • 3. Museum für Naturkunde, Leibniz-Institut für Evolutions- und Biodiversitätsforschung, Berlin, Germany|Institute for Biochemistry and Biology, University of Potsdam, Potsdam, Germany
  • 4. Institute for Biochemistry and Biology, University of Potsdam, Potsdam, Germany
  • 5. Wildlife Institute of India, Chandrabani, India
  • 6. Department of Zoology, University of Peshawar, Peshawar, Pakistan
  • 7. Central Department of Botany, Tribhuvan University, Kathmandu, Nepal
  • 8. Zoological Sciences Division, Pakistan Museum of Natural History, Islamabad, Pakistan
  • 9. Museum Koenig, Leibniz Institute for the Analysis of Biodiversity Change, Bonn, Germany

Description

Figure 2 DNA-based relationships in the Gloydius himalayanus complex. A. Maximum-likelihood phylogeny inferred from concatenated mitochondrial and nuclear markers (3882 bp) showing relationships within species of the G. himalayanus complex. SH-aLRT, ultrafast bootstrap (UFBoot), and posterior probability (PP) values indicate nodal support. Coloured boxes correspond to the clades recognised in this study (Fig. 1); B. Ninety-five percent statistical parsimony allele networks for three nuclear loci (C-mos, PRLR, NT3) showing relationships among phased alleles of the G. himalayanus complex and other Gloydius species. Each circle represents one unique allele; circle size is proportional to its frequency, and colours indicate species assignment. Small black circles correspond to inferred unsampled alleles, and red dashed lines indicate mutational connections between alleles of the G. himalayanus complex and those of other Gloydius.

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Journal article: 10.3897/zookeys.1280.182768 (DOI)
Journal article: https://zenodo.org/record/20352597 (URL)