Published May 21, 2026
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Figure 1 from: Jablonski D, Tillack F, Mahlow-Tillack K, Petzold A, Wilzo M, Das A, Idrees M, Baniya CB, Masroor R, Hofmann S (2026) Integrative taxonomy reveals previously undescribed diversity within the Gloydius himalayanus complex (Squamata, Viperidae, Crotalinae) from the Himalaya and Hindu Kush. ZooKeys 1280: 83-153. https://doi.org/10.3897/zookeys.1280.182768
Authors/Creators
- 1. Department of Zoology, Comenius University in Bratislava, Bratislava, Slovakia
- 2. Museum für Naturkunde, Leibniz-Institut für Evolutions- und Biodiversitätsforschung, Berlin, Germany
- 3. Museum für Naturkunde, Leibniz-Institut für Evolutions- und Biodiversitätsforschung, Berlin, Germany|Institute for Biochemistry and Biology, University of Potsdam, Potsdam, Germany
- 4. Institute for Biochemistry and Biology, University of Potsdam, Potsdam, Germany
- 5. Wildlife Institute of India, Chandrabani, India
- 6. Department of Zoology, University of Peshawar, Peshawar, Pakistan
- 7. Central Department of Botany, Tribhuvan University, Kathmandu, Nepal
- 8. Zoological Sciences Division, Pakistan Museum of Natural History, Islamabad, Pakistan
- 9. Museum Koenig, Leibniz Institute for the Analysis of Biodiversity Change, Bonn, Germany
Description
Figure 1 A. Phylogenetic placement of the G. himalayanus complex within the genus Gloydius (for details see Fig. 2A); B. Geographic distribution (dark shading) of recognised lineages within the G. himalayanus complex, along with the sampling localities of genetically investigated specimens (see Table 1, Suppl. material 1: table SS1). The position of the locality for the lectotype BMNH 1946.1.19.64 is only for map purposes (see remarks on the lectotype in the species account). Due to incomplete locality data in Kuttalam et al. (2022) (see Fig. 2A), some sample locations are not depicted in panel (B); an inset shows the approximate extent of the major mountain ranges in the region, following Spicer et al. (2021, 2025). Question marks indicate uncertain or potential occurrences in eastern Afghanistan. The map was generated using QGIS 3.28 (https://qgis.org/) with the WGS 84 coordinate reference system. Sketch shows a syntype of Halys himalayanus, taken from Günther (1864). This individual corresponds to the specimen BMNH 1946.1.19.64, which is designated here as the lectotype for Gloydius himalayanus.
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- Journal article: 10.3897/zookeys.1280.182768 (DOI)
- Journal article: https://zenodo.org/record/20352597 (URL)