Deltochilum abdominale Martínez, 1947

Figs 3 B, 3 G, 4 A, 5, 6, 7 A (blue circle)

Deltochilum (D.) abdominalis Martínez, 1947: 269–274. Lámina VIII (description).

Deltochilum (Deltohyboma) abdominalis Martínez 1954: 47 (cited from Colombia, error; see D. ventripuncturatus sp. nov.).

Deltochilum (Deltohyboma) abdominale Vulcano & Pereira, 1964: 652 (catalogue).

Deltochilum (Deltohyboma) abdominalis Blanco 1987: 43 (distribution, Venezuela).

Deltochilum abdominale Medina et al. 2001: 136 (cited from Guaviare (gv?) Colombia with doubts).

Deltochilum (Deltohyboma) abdominalis González-Alvarado & Vaz-de-Mello, 2021: 62, 74 (species group).

Deltochilum abdominale Lozano De La Rosa et al. 2024: 72 (declination of the species epithet in neuter, distribution, Venezuela).

Type material examined.

Holotype ♂ Venezuela, D. [istrito] F. [ederal], 720 mts alt., Mo. del Naiguatá, AGO. 941. R. Lichy. Legit. Coll Martínez. [white label] HOLOTIPO. [red label] Deltochilum abdominalis [male symbol] sp. nov. A. Martínez-DET. 946. [white label] FICHADO. [white label] MACN-En 790. (MACN).

Additional material examined

(81 specimens): Colombia: Norte de Santander: Toledo, IP Santa María, Alto de Herrera, Vda. Diamante, Fca. La Primavera. PNN Tamá, 7°7'N, 72°13'W, 1000 m, 1 ♀, 2 ♂, 1999. ix. 22, González E., T. Exc. Hum (IAvH), Toledo, Vda. San Alberto. Bosque, 07°11'08.8"N, 072°18'39.6"W, 823 m, 2 ♂, 11. iv. 2019, Ramírez, M. & Lopera, A., T. Exc. H # SAF 1 (24 h) (1 ♂ IAvH, 1 ♂, CALT-ECC), Toledo, Vda. San Alberto. Bosque, 07°11'15.0"N, 072°18'28.3"W, 1104 m, 1 ♀, 11. iv. 2019, Ramírez, M. & Lopera, A., T. Exc. H # SAF 3 (24 h) (IAvH). Venezuela: Chichiriviche (Litoral, D. F), [10°56'1.51"N], [68°16'40.14"W], 1 ♀, 27. i. 1977, Bordón coll. (BDGC); Aragua: Cuyagua, [10°26'42"N], [67°41'44"W], 800 m, 2 ♀, 1 ♂, x. 1972, Martínez coll. (CMNC), Quebrada Coirral da Piedra, El Limón, [10°19'N], [67°39'W], 700 m, 1 ♂, 7. vi. 1973, Bordón coll. (CMNC), Racho Grande, [10°4'1.40"N], [67°32'34.02"W], 1200 m, 1 ♂, 27–30. viii. 1983, B Gill coll. (BDGC), Racho Grande, [10°4'1.40"N], [67°32'34.02"W], 1150 m, 1 ♂, 4–9. vii. 1986, B Gill coll. (BDGC); Miranda: Parque Nacional Guatopo, [10°5'37"N], [66°29'41"W], 6 ♀, 6 ♂, 3–8. iii. 1988, F. Génier coll., pitfall trap, human feces (CMNC), Parque Nacional Guatopo, 50 km SE Caracas. forest, [10°5'N], [66°29'W], 400 m, 2 ♀, 1 ♂, 5–6. iii. 1971, S. Peck coll., pitfall trap, human dung (CMNC), Santa Crucita, 33 km N Altagracia, Parque Nacional Guatopo. forest, [10°14'N], [67°1'W], 400 m, 1 ♂, 7–10. vi. 1987, S. & J. Peck coll., dung trap (CMNC); Monagas: Cueva Guácharo, Caripe. forest over coffee, [10°11'N], [63°32'30"W], 750 m, 1 ♀, 1 ♂, 20–31. vii. 1987, S. & J. Peck coll. (CMNC), Teresén, [10°11'N], [63°29'W], 800 m, 3 ♂, 12. iv. 1963, R. Hernandez coll. (CMNC); Táchira: 20 km NE San Cristobal, [7°57'34"N], [72°5'3"W], 3000 m, 1 ♂, 20. v. 1974, S. Peck coll. (CMNC), 20 km NE San Cristobal, [7°57'34"N], [72°5'3"W], 1200 m, 1 ♂, 20–22. v. 1974, S. Peck coll. (CMNC), La Trampita, Presa La Honda, [7°56'1"N], [71°43'24"W], 1100 m, 1 ♀, 17. xii. 1988, D. Havranek coll., mini-dung cup traps (CMNC), Las Trampitas, Pregonero. sub-mountain forest, [8°0'40"N], [71°45'38"W], 1240 m, 10 ♀, 11 ♂, 9–14. vii. 1989, S. & J. Peck coll., carrion traps (CMNC), Pregonero Las Trampitas, [7°59'48.89"N], [71°45'49.08"W], 1240 m, 2 ♀, 4 ♂, 9–14. vii. 1989, J Peck & S Peck coll. (BDGC), Rio Frio, [7°50'37.11"N]?, [72°6'33.48"W]?, 500 m, 3 ♀, 3 ♂, 11–18. viii. 1983, B Gill coll. (BDGC), San Cristobal, [7°45'30.99"N], [72°12'20.62"W], 1200 m, 3 ♀, 6 ♂, 10–18. viii. 1983, B Gill coll. (BDGC); Yaracuy: Bolívar. Aroa, 10°20'22.74"N, 68°50'5.71"W, 1372 m, 1 ♀, 1 ♂, 19. VII. 2009, Asmüssen M, Colmenares P., Martínez H., Cebo Heces humanas (CEMT).

Diagnosis.

This species can be distinguished from all other species within the plebejum species group by having the smallest interstrial punctures (Fig. 5 C). These punctures are nearly the same size as the interstrial shiny points; in all other species of the group, these shiny points are distinctly smaller than the interstrial punctures. Furthermore, the males possess a denticle on the mesofemur (Fig. 6 A, G), a character shared with D. tyba sp. nov. (Fig. 9 G), but the two species can be separated by the size of the interstrial punctures previously described.

Redescription.

Minor male. Size 10 mm. Color. Dark brown with some blue reflections dorsally. Dark blue ventrally. Head (Fig. 5 A). Interocular distance 8 × the width of one eye. Punctures on frons separated by one or less than one diameter of each puncture and subequal in size to head discal punctures. Disc punctures separated by one or less than one diameter of each puncture. Punctures from disc towards anterior area successively of almost with the same size. Genal punctures subequal in size to the discal punctures and separated by one diameter of each puncture. Pronotum (Fig. 5 B). Margin between anterior and medial-lateral angle almost straight. Medial angle of pronotum rounded. Margin between medial-lateral angle and posterior-lateral angle almost straight. Disc punctures half size to anterior-lateral ones and separated by more than one diameter of each puncture. Basal punctures are smaller than anterior-lateral ones and separated by approximately one diameter of each puncture. Hypomera. With ocellated and setigerous punctures on the anterior part (Fig. 3 B, white arrow). Most of punctures incompletely closed posteriorly and without setae (Fig. 3 B, green arrow). Anterior punctures denser and larger than posterior ones. Elytra (Fig. 5 C). Carina of the ninth interstria almost reaching middle of elytral length. Elytral apex on interstriae III, V – VII with tubercles. Striae I – VII almost inconspicuous. Width of third stria on disc ~ 1 / 55 of the distance between striae II and III. Strial punctures ~ 2 × wider than the stria. First stria as wider as second stria. Stria VIII conspicuous apically, laterally almost inconspicuous. Punctures on interstriae approximately separated by two diameters of each puncture and the same size or slightly smaller than strial punctures. Punctures of the third interstria on disc occupying ~ 1 / 18 of the distance between striae II and III. Interstriae with shiny points mixed with the punctures. Points almost the same size as the interstrial punctures. Metaventrite (Fig. 5 D). Disc with few deep excavation, occupying metaventral basal third. Anterolateral areas of metaventral process with punctures completely closed and separated by less than one diameter of each puncture. Anterocentral area of metaventral process with few and small punctures, subequal in size to the discal punctures. Punctures on the lateral lobules not completely closed (some of them) and separated by one or less than one diameter of each puncture. Abdomen (Fig. 5 E). Ventrite I expanded posteriorly, expansion reaching the fifth ventrite; on ventrite III, the expansion is as wide as the distance between mesocoxae. Basal part of the ventrite I, between the metacoxae not modified. Legs (Fig. 6 A, B). Protibia strongly broadens toward the apex. Profemora with punctures separated by less than one diameter of each puncture. Mesofemur with punctures separated by 2 × the diameter of each puncture. Mesofemur with a basal denticle on the posterior edge. Metafemur regularly broadens toward the apex. Metafemur with punctures separated by 2 × the diameter of each puncture. Metatibial spur insertion not elongated. Metatibial spur clearly distinct from the metatibia. Internal margin of metatibia with tubercles on the basal half of the metatibial length. Pygidium (Fig. 6 C). Pygidial punctures almost uniform in size and density. Punctures separated by one or less than the diameter of each puncture. Discal punctures, on the center of the pygidium, small, each puncture occupying ~ 1 / 45 of the distance between the lateral pygidial margins. Genitalia (Fig. 6 D – F). Paramera smaller than the phallobase. Apex of paramera with a large denticle in lateral view. Medial area of endophallus with two endophallites, the right ~ 4 × longer than the other.

Variation.

The female exhibits the differences described here under the sexual dimorphism section of the species group redescription. Medium-sized males have both the basal denticle of the metafemur and the metatibial tubercles (Fig. 6 J, white arrow) that are larger than those found in the minor male. The metaspur is articulated. Major males: the metatibial basal carina is irregular and strongly interrupted by setae, giving it the appearance of tubercles (Fig. 6, white arrow). This carina is significantly reduced in the mid-length of the metatibial; distal to this reduction, the carina rises again (giving the appearance of having both tubercles and carina). Other major males possess this tuberculate basal part, but the reduction is less pronounced, and the tubercles are continuous (Fig. 6 I, white arrow). The basal denticle of the metafemur is larger and more strongly curved (Fig. 6 G). The metatibia is also more strongly curved (Fig. 6 H, I). Metaspur fused.

Known distribution

(Fig. 7 A blue circle). Colombia, Norte de Santander. Venezuela, Aragua, Miranda, Monagas, Táchira, Yaracuy.

Remarks.

This species has been collected in forest, submontane forest and few specimens in forest mixed with coffee. This species is one of the members of the plebejum species group that is widely distributed, possessing a broad altitudinal range from 400 to probably 1,400 m above sea level.