Published March 11, 2026 | Version v1
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APPENDIX 4 in Tracing the footprints of the Atlas Brown Bear: a metric analysis of Holocene and Late Pleistocene remains from Northern Morocco

  • 1. Instituto Universitario de Xeoloxía (IUX), Universidade da Coruña, Edificio de Servicios Centrales de Investigación (ESCI), Campus de Elviña s / n, 15071 A Coruña (Spain) & National Institute of Archaeological Sciences and Heritage (INSAP), Madinat Al Irfane, Hay Riad, BP 6828 Rabat Institutes (Morocco)
  • 2. Instituto Universitario de Xeoloxía (IUX), Universidade da Coruña, Edificio de Servicios Centrales de Investigación (ESCI), Campus de Elviña s / n, 15071 A Coruña (Spain) & Archaeoscience Platform (AsP), Research Institute of the University of Bucharest (ICUB), University of Bucharest, 90 - 92 Sos. Panduri, 5 th District, 50663 Bucharest (Romania)
  • 3. Instituto Universitario de Xeoloxía (IUX), Universidade da Coruña, Edificio de Servicios Centrales de Investigación (ESCI), Campus de Elviña s / n, 15071 A Coruña (Spain)
  • 4. National Institute of Archaeological Sciences and Heritage (INSAP), Madinat Al Irfane, Hay Riad, BP 6828 Rabat Institutes (Morocco) & Max Planck Institute for Evolutionary Anthropology, Deutscher Platz 6, 04103 Leipzig (Germany)

Description

APPENDIX 4. — Main measurements (in mm) of brown bear phalanges from the sites of Keft-el-Hammar (KEH) and Hattab II (HAT II). Abbreviations: APW prox, proximal anteroposterior width; H prox, proximal height; n/n, no number; TL, total length; TW dia, transversal width of diaphysis; TW dist, distal transversal width; TW prox, proximal transversal width. The asterisk (*) indicates elements that were published previously (Ouchaou 2000).

First phalanx
SiteLevelNumberTLTW proxAPW proxTW diaTW distSource
HAT II755830.716.315.611.412.21
HAT II311533.817.814.811.113.01
HAT II324534.118.313.810.613.31
HAT II655734.718.313.310.811.81
HAT II636037.217.514.811.813.81
HAT II637238.718.314.512.013.71
HAT II325938.817.615.411.713.81
KEH1937.316.415.09.514.12
KEH1F7/2737.621.512.310.512.32
KEH1HST837.821.016.313.213.42
KEH1I9/438.017.016.417.314.02
KEH1J2946.018.515.011.514.02
KEH227/A32.018.012.011.013.02
KEH28932.017.012.011.014.02
KEH25834.017.014.011.013.02
KEH2n/n36.017.016.012.014.02
KEH2n/n36.018.015.012.014.02
KEH2n/n37.217. 812.313.42
KEH2n/n38.017.015.012.014.02
KEH2n/n41.417.213.012.012.32
KEH436.917.514.811.413.83
KTGC35.017.012.59.812.83
KTGD34.017.013.410.312.43
KTGF35.017.013.09.512.53
KTGF37.018.314.911.213.33
KTGF40.020.015.813.014.93
KTGG39.020.014.012.03
KBS3c37.820.515.512.013.53
GKHIIIb40.020.016.013.513.93
KEBsalle43.021.017.414.615.73
IOS05/2151.023.013.24
IOS05/3643.016.914.59.24
IOS05/3719.414.511.34
IOS06/I6/2143.023.918.314.916.44
IOS06/I8/238.017.514.411.413.94
IOS06/I10/846.017.615.09.614.04
IOS06/I10/1936.317.215.011.513.84
IOS06/I10/2037.519.115.011.914.04
IOS06/I10/2831.016.013.09.312.14
IOS06/I10/2933.016.013.09.312.14
IOS06/I11/1731.015.913.39.412.24
IOS06/I11/1831.515.813.59.912.14
IOS06/I11/2133.518.214.410.412.04
IOS06/I11/2431.016.013.310.112.24
IOS06/I11/2533.417.314.110.412.84
IOS06/I11/2733.318.214.310.512.24
IOS06/I11/2934.017.314.410.912.94
IOS06/J4/139.224.017.713.415.24
Second phalanx
SiteLevelNumberTLTW proxAPW proxTW diaTW distSource
HAT II643021.814.413.311.212.31
KEH2n/n27.016.015.011.014.02
KEH2n/n27.016.013.011.014.02
KEH2n/n29.017.016.214.015.02
KEH2n/n30.019.515.313.014.02
KEH2n/n30.018.017.013.816.02
KEH428327.517.015.313.014.12
KTGE28.815.913.411.114.03
KTGF26.210.513.03
KTGF27.515.713.110.214.13
KTGF22.617.313.912.314.93
GKHV19.916.012.911.113.03
IOS05/928.815.913.411.114.04
IOS05/3826.210.513.04
IOS06/I10/627.515.713.110.214.14
IOS06/I5/222.617.313.912.314.94
IOS06/I6/1719.916.012.911.113.04
IOS06/I10/2120.014.911.29.812.94
IOS06/I10/3118.914.112.110.712.14
IOS06/I11/1019.414.211.59.812.24
IOS06/I11/1618.514.112.110.512.34
IOS06/I11/2017.914.911.310.412.34
IOS06/J4/1021.518.315.614.315.84
Third phalanx
SiteLevelNumberTLH proxTW proxSource
HAT II6104144.523.814.11
HAT II667435.725.013.91
HAT II687931.026.011.31
KEHsurfacen/n11.02
KEHsurfaceHST726.013.02
KEH1130.021.011.22
KEH25246.023.014.02
KEH2n/n36.426.015.52
KEH2n/n30.021.011.02
KEH3n/n30.026.011.02
KEH3n/n41.024.014.02
KEH3n/n49.013.73
KEH4n/n56.328.015.53
KEH7n/n24.012.32
KTGD50.533.016.73
KTGE56.030.014.53
KTGF54.028.514.43
KTGF50.029.014.23
KTGF51.029.815.03
KTGF45.014.03
KBS3c50.526.014.73
KEBsalle36.026.015.03
IOS05/1345.027.514.04
IOS06/I10/950.028.215.04
IOS06/I10/1133.023.012.24
IOS06/I10/3028.022.012.44
IOS06/I10/3231.423.212.24
IOS06/I11/933.123.412.74
IOS06/I11/1530.021.013.04
IOS06/I12/132.223.014.24
IOS06/I13/534.021.514.24
IOS06/I10/1032.025.014.24
IOS06/J5/631.823.713.44

Notes

Published as part of Iken, Shaymae, García-Vázquez, Ana, González-Fortes, Gloria, Bouzouggar, Abdeljalil & Grandal-D'Anglade, Aurora, 2026, Tracing the footprints of the Atlas Brown Bear: a metric analysis of Holocene and Late Pleistocene remains from Northern Morocco, pp. 75-76 in Comptes Rendus Palevol 25 (4) on page 75, DOI: 10.5852/cr-palevol2026v25a4, http://zenodo.org/record/19005282

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