Published April 21, 2026 | Version v1
Journal article Open

Range expansion of short-winged grasshopper Pezotettix giornae (Orthoptera, Acrididae) along the northern edge of its range in Central Europe

  • 1. Faculty of Forestry and Wood Sciences, Czech University of Life Sciences, Praha, Czech Republic
  • 2. Regional Museum in Mikulov, Mikulov, Czech Republic
  • 3. Beleco, Praha, Czech Republic
  • 4. Department of Biology and Ecology, University of Ostrava, Ostrava, Czech Republic
  • 5. Podyjí National Park Administration, Znojmo, Czech Republic
  • 6. Department of Physical Geography and Geoecology, University of Ostrava, Ostrava, Czech Republic
  • 7. Unaffiliated, Wien, Austria

Description

Orthoptera species distribution ranges are dynamically shifting because of climate change. This article documents the expansion of the short-winged grasshopper Pezotettix giornae in Western and Central Europe between 1990 and 2024. Published records from Central Europe (Austria, Slovakia, Hungary, and the Czech Republic) are supplemented with data from public databases and targeted field surveys, particularly in southeastern Moravia and the Burda Hills region of southern Slovakia. Whereas P. giornae was previously abundant mainly in the Mediterranean reaching northwards into central Hungary, its range has gradually shifted northward over the past 30 years, now reaching central France, Austria, Slovakia, and the Czech Republic, with single individuals transported to northern France, northern Switzerland, central Germany, and the Netherlands. The increase in abundance and the number of newly recorded localities indicate the dynamic spread of this species despite its inability to fly. This expansion is facilitated by climate warming, the species' ability to colonize ruderal and successional habitats, and passive dispersal via transportation, particularly along road and railway corridors. The case of P. giornae demonstrates that even brachypterous species with limited dispersal ability can undergo significant range shifts. This trend has important implications for ecological modeling and the prediction of future species distributions and biodiversity conservation strategies in the changing European landscape.

Files

JOR_article_173175.pdf

Files (5.2 MB)

Name Size Download all
md5:5044a7e3e53e5998b96eb5654c95b46a
5.1 MB Preview Download
md5:d27c55dc32178619e6bad620486c4e64
114.8 kB Preview Download

Linked records

Additional details

References

  • Adlbauer K, Sackl P (1993) Zum Vorkommen und zur Verbreitung seltener Heuschrecken und Grillen in der Steiermark. Mitteilungen der Abteilung für Zoologie am Landesmuseum Joanneum 47: 55–66.
  • Beckmann BC, Purse BV, Roy DB, Roy HE, Sutton PG, Thomas CD (2015) Two species with an unusual combination of traits dominate responses of British grasshoppers and crickets to environmental change. PLoS ONE 10: e0130488. https://doi.org/10.1371/journal.pone.0130488
  • Berg H-M, Zuna-Kratky T (1997) Heuschrecken und Fangschrecken (Insecta: Saltatoria, Mantodea). Eine Rote Liste der in Niederösterreich gefährdeten Arten. Amt der Niederösterreichischen Landesregierung, Abteilung Naturschutz, Wien, 112 pp.
  • Berg HM, Bieringer G, Zechner L (2005) Rote Liste der Heuschrecken (Orthoptera) Österreichs. In: Zulka KP (Ed.) Rote Listen gefährdeter Tiere Österreichs. Checklisten, Gefährdungsanalysen, Handlungsbedarf. Teil 1. Grüne Reihe des Lebensministeriums 14/1, Wien, 167–209.
  • Braun VB, Lederer E, Sackl P, Zechner L (1995) Verbreitung, Phänologie und Habitatansprüche der Großen Schiefkopfschrecke, Ruspolia nitidula Scopoli, 1786, in der Steiermark und im südlichen Burgenland. Abhandlungen der Zoologischen Abteilung des Landesmuseums Joanneum 49: 57–87.
  • Browde J, Pedigo L, Degooyer T, Highley L, Wintersteen W, Zeiss M (1992) Comparison of sampling techniques for grasshoppers (Orthoptera: Acrididae) in soybean. Journal of Economic Entomology 85: 2270–2274. https://doi.org/10.1093/jee/85.6.2270
  • Budrys E, Martinaitis K, Valinčienė K, Ferenca R (2019) First records of Aiolopus thalassinus (Fabricius, 1781) and Euthystira brachyptera (Ocskay, 1826) (Orthoptera: Acrididae) in Lithuania. Bulletin of the Lithuanian Entomological Society 3: 8–10.
  • Ćato S, Zagorac D (2021) Unexpected faunistic records of Rhacocleis annulata, Eyprepocnemis plorans, and Xya pfaendleri (Orthoptera) from Croatia and Slovenia. Natura Croatica 30: 501–511. https://doi.org/10.20302/nc.2021.30.33
  • David S (2009) Stepní flóra a vegetace zemních valů hradiska v obci Bíňa (okr. Nové Zámky). Bulletin Slovenskej Botanickej Spoločnosti, Bratislava 31: 77–85.
  • Didham RK, Barbero F, Collins CM, Forister ML, Hassall C, Leather SR, Stewart AJA (2020) Spotlight on insects: Trends, threats and conservation challenges. Insect Conservation and Diversity 13: 99–102. https://doi.org/10.1111/icad.12409
  • Elmqvist H, Nyström D, Bergsten J (2020) Aiolopus thalassinus (Fabricius, 1781), en ny gräshoppa för Norden och en migrerande svärm som korsade Östersjön. Entomologisk Tidskrift 141: 161–171.
  • Fartmann T, Poniatowski D, Holtmann L (2021) Habitat availability and climate warming drive changes in the distribution of grassland grasshoppers. Agriculture, Ecosystems & Environment 320: 107565. https://doi.org/10.1016/j.agee.2021.107565
  • Fedor PJ, Majzlan O (2001) Distribution and infiltration of the tree cricket Oecanthus pellucens (Scopoli, 1763) to unoriginal conditions in Slovakia. Bulletin de la Société des Naturalistes Luxembourgeois 102: 103–108.
  • Fontana P, Buzzetti F, Cogo A, Odè B (2002) Guida al riconoscimento e allo studio di cavallette, grilli, mantidi e insetti affini del Veneto. Museo Naturalistico Archeologico di Vicenza Ed., Vicenza, 592 pp.
  • GBIF.org [Global Biodiversity Information Facility] (2025) Pezotettix giornae (Rossi, 1794) occurrence data. https://www.gbif.org/species/1705356 [Accessed: 14 June 2025]
  • Gomboc S, Šegula B (2014) Pojoče kobilice Slovenije / Singing Orthoptera of Slovenia. Center za kartografijo favne in flore, Miklavž na Dravskem polju, 256 pp.
  • Gojznikar J, Poljanec N, Mlakar Medved M (2018) An interesting new record of Egyptian locust Anacridium aegyptium (Linnaeus, 1764) (Orthoptera: Acrididae) for Slovenian inland. Natura Sloveniae 20: 33–34. https://doi.org/10.14720/ns.20.2.33-34
  • Grobelny S, Żurawlew P, Kutera M, Brodacki M, Kupczyk M, Gwardjan M, Radzikowski P, Itczak A, Czyżewski S (2019) Kolejne stanoviska nadrzewka południowego Meconema meridionale Costa, 1860 (Orthoptera: Tettigoniidae) w Polsce. Przegląd Przyrodniczy 30: 27–37.
  • Hatna E, Bakker M (2011) Abandonment and expansion of arable land in Europe. Ecosystems 14: 720–731. https://doi.org/10.1007/s10021-011-9441-y
  • Harz K (1975) Die Orthopteren Europas II. Series Entomologica 11. Dr. W. Junk B.V., The Hague, 1–939. https://doi.org/10.1007/978-94-010-1947-7
  • Hochkirch A, Nieto A, García Criado M, Cálix M, Braud Y, Buzzetti FM, Chobanov DP, Odé B, Presa Asensio JJ, Willemse L, Zuna-Kratky T et al. (2016) European Red List of Grasshoppers, Crickets and Bush-crickets. Publications Office of the European Union, Luxembourg, 1–95.
  • Hochkirch A, Andreä J, Bodingbauer S, Jacobi B, Klein R, Paulus C, Pittius U, Rautenberg T, Sändig S, Sattler J (2020) Heuschrecken in Deutschland 2019 – Interessante Heuschreckennachweise auf der Meldeplattform heuschrecken.observation.org aus dem Jahr 2019. Articulata 35: 93–103.
  • Hochkirch A, Franzen A, Blümel-Zimmermann E, Brozio J, Brozowski F, Füldner K, Guggemoos T, Hodges S, König S, Lehmeier S, Mähler M, Paulus C, Mehesz P, Richter W, Supperl F, Thiess L, Türk N, Winter A, Sändig S (2023) Heuschrecken in Deutschland 2022/2023 – Interessante Heuschreckennachweise auf observation.org aus den Jahren 2022 und 2023. Articulata 38: 45–68.
  • Holuša J, Kočárek P, Marhoul P (2007) Recent progress of Euchorthippus declivus in the Czech Republic. Čas. Slez. Muz. Opava (A) 56: 59–62.
  • Holuša J, Kocarek P, Vlk R, Marhoul P (2013) Annotated checklist of the grasshoppers and crickets (Orthoptera) of the Czech Republic. Zootaxa 3616: 437–460. https://doi.org/10.11646/zootaxa.3616.5.2
  • Holuša J (2012) Grasshoppers and bushcrickets regionally extinct in the Czech Republic: Consequence of the disappearance of habitats scattered on the edge of their ranges. Journal of Insect Conservation 16: 949–960. https://doi.org/10.1007/s10841-012-9534-8
  • Huntley B (1991) How plants respond to climate change: migration rates, individualism and the consequences for plant communities. Annals of Botany 67(Suppl. 1): 15–22. https://doi.org/10.1093/oxfordjournals.aob.a088205
  • IPCC (2018) Global Warming of 1.5 °C. An IPCC Special Report. Geneva: Intergovernmental Panel on Climate Change, s. 1–32.
  • iNaturalist community (2025) Observations of Pezotettix giornae. https://www.inaturalist.org [Retrieved on 19.11.2025]
  • Iorgu I, Pisica E, Pai P, Lupu G, Iuan C (2008) Checklist of Romanian Orthoptera (Insecta) and their distribution by eco-regions. Travaux du Museum National d'Histoire Naturelle "Grigore Antipa" 51: 119–135.
  • Iorgu IŞ, Pisică EI (2006) Contributions to the distribution knowledge of some Orthoptera species (Insecta: Orthoptera) from Eastern Romania. Complexul Muzeal de Ştiinţele Naturii "Ion Borcea" Bacău – Studii şi Comunicări 21: 251–256.
  • Kaláb O, Pyszko P, Kočárek P (2021) Estimation of the recent expansion rate of Ruspolia nitidula (Orthoptera) on a regional and landscape scale. Insects 12: 639. https://doi.org/10.3390/insects12070639
  • Kenyeres Z, Takács G, Bauer N (2019) Response of orthopterans to macroclimate changes: A 15-year case study in Central European humid grasslands. Journal of Orthoptera Research 28(2): 187–193. https://doi.org/10.3897/jor.28.34102
  • Kleukers R, Decleer K, Haes E, Kolshorn P, Thomas B (1996) The recent expansion of Conocephalus discolor (Thunberg) (Orthoptera: Tettigoniidae) in western Europe. Entomologist's Gazette 47: 37–50.
  • Knechtel WK, Popovici-Biznosanu A (1959) Fauna Republicii populare Romine, Insecta.
  • Kočárek P (1999) First record of Pezotettix giornae (Orthoptera, Acrididae, Catantopinae) from Slovakia. Biologia (Bratislava) 54: 150.
  • Kočárek P, Holuša J, Vlk R, Marhoul P, Zuna-Kratky T (2008) Recent expansions of the bush-crickets Phaneroptera falcata and Phaneroptera nana (Orthoptera: Tettigoniidae) in the Czech Republic. Articulata 23: 67–75.
  • Kočárek P, Holuša J, Plasgura L, Skokanova H (2013) Fragmented distribution of Isophya pienensis Maran, 1954 (Insecta, Orthoptera) in mountains: A result of permanent forest opening in highly forested landscape. Periodicum Biologorum 115: 417–423.
  • Köhler G (2024) Orthopterological highlights from the Biodiversity Monitoring South Tyrol. In: 4th European Congress on Orthoptera Conservation (ECOC IV), Abstract Booklet, Fondazione Museo Civico di Rovereto: 25.
  • Köhler G, Worschech K, Krech M, Korsch H (2022) Die Südliche Eichenschrecke (Meconema meridionale A. Costa, 1860) in Thüringen (Orthoptera: Meconematidae). Mitteilungen des Thüringer Entomologenverbandes eV 29: 9–30.
  • Koschuh A, Zechner L (2006) Aktuelle und historische Nachweise ausgewählter Heuschreckenarten in der Steiermark, Österreich – Teil II: Caelifera (Saltatoria). Joannea Zoologie 8: 55–85.
  • Krištín A, Jarčuška B, Kaňuch P (2013) Notes on Orthoptera (Insecta) and their assemblages in the Romanian Carpathians. Travaux du Muséum National d'Histoire Naturelle "Grigore Antipa" LVI: 19–32. https://doi.org/10.2478/travmu-2013-0002
  • Krištín A [Ed.] (2025) Pezotettix giornae. Orthoptera of Slovakia. http://orthoptera.sk/ [accessed 2025-06-14]
  • Krištín A, Lengyel J, Kršiak B, Fabriciusová V (2007) K výskytu rovnokrídlovcov a ekológií krátkonôžky štíhlej (Ablepharus kitaibelii) na Burde. Ochrana Prírody 9: 132–139.
  • Krištín A, Jarčuška B, Kaňuch P (2020) An annotated checklist of crickets, grasshoppers and their allies (Orthoptera) in Slovakia. Zootaxa 4869: 207–241. https://doi.org/10.11646/zootaxa.4869.2.3
  • Lapin M, Melo M (1999) Climate trends in Slovakia and Central Europe: Changes in annual air temperature and precipitation over the 20th century. In: Climate Change Impacts in Central Europe, s. 15–28. Bratislava: Slovak Academy of Sciences.
  • Lederer E (2004) Autökologische Untersuchungen an Heuschrecken (Saltatoria) und Fangschrecken (Mantodea) im südlichen Burgenland. Dipl. Arb. Univ. Graz, 114 pp. [+ Anh.]
  • Liana A (1992) Owady Prostoskrzydle Orthoptera. In: Glowacinski Z (ed) Czerwona Lista Zwierzat Ginacych i Zagrozonych w Polsce. Polska Akademia Nauk, Kraków, 85–91.
  • Löffler F, Poniatowski D, Fartmann T (2019) Orthoptera community shifts in response to land-use and climate change—Lessons from a long-term study across different grassland habitats. Biological Conservation 236: 315–323. https://doi.org/10.1016/j.biocon.2019.05.058
  • Machatz B, Ratsch A, Prasse R, Ristow M (2005) Rote Liste und Gesamtartenliste der Heuschrecken und Grillen (Saltatoria: Ensifera et Caelifera) von Berlin. In: Der Landesbeauftragte für Naturschutz und Landschaftspflege / Senatsverwaltung für Stadtentwicklung (Hrsg) Rote Listen der gefährdeten Pflanzen und Tiere von Berlin. Kulturbuch-Verlag, Berlin, pp. 1–12.
  • Mařan J (1954) Rovnokřídlý hmyz státních reservací v okolí Štúrova na jižním Slovensku. Die Othopterenfauna der staatlichen Naturschutzgebiete bei Štúrovo in der Südslowakei (ČSR). Ochrana Přírody 9: 132–139.
  • Marhoul P, Holuša J, Kaláb O, Kočárek P, Kuřavová K, Musiolek D, Stanislav R, Vlk R (2019) Mapování rozšíření rovnokřídlých (Orthoptera) ve vybraných krajích České republiky v letech 2012–2015: Mapping of the Distribution of the Orthopterans (Orthoptera) in Selected Regions of the Czech Republic. Příroda, 41–54.
  • Mason SC, Palmer G, Fox R, Gillings S, Hill JK, Thomas CD, Oliver TH (2015) Geographical range margins of many taxonomic groups continue to shift polewards. Biological Journal of the Linnean Society 115: 586–597. https://doi.org/10.1111/bij.12574
  • Massa B, Fontana P, Buzzeti F, Kleukers R, Odè B (2012) Fauna d'Italia: Orthoptera. 1a edition. Calderini-Edizioni, 563 pp.
  • Masters G, Brown V, Clarke I, Whittaker J (1998) Direct and indirect effects of climate change on insect herbivores: Auchenorrhyncha (Homoptera). Ecological Entomology 23: 45–52. https://doi.org/10.1046/j.1365-2311.1998.00109.x
  • McAlpine DF, Ogden JB (2012) New and noteworthy records of Orthoptera from Maritime Canada. Journal of the Acadian Entomological Society 8: 43–47.
  • Menéndez R, González-Megías A, Jay-Robert P, Marquéz-Ferrando R (2014) Climate change and elevational range shifts: Evidence from dung beetles in two European mountain ranges. Global Ecology and Biogeography 23: 646–657.
  • Monnerat C, Gurcel K, Magnouloux M, Dunant F (2020) Premières observations de Rhacocleis annulata Fieber, 1853 en Suisse et en Haute-Savoie limitrophe (Orthoptera: Tettigoniidae). Entomo Helvetica 13: 37–44.
  • Mückstein P, Vlk R (2015) Record of Aiolopus thalassinus (Orthoptera: Acrididae) in the Žďárské vrchy Protected Landscape Area (Bohemian-Moravian Highlands, Czech Republic). Acta Rerum Naturalium 18: 1–2.
  • Munteanu C, Kuemmerle T, Boltiziar M, Butsic V, Gimmi U, Lúboš H, Kaim D, Király G, Konkoly-Gyuró É, Kozak J, Lieskovský J, Mojses M, Müller D, Ostafin K, Ostapowicz K, Shandra O, Štych P, Walker S, Radeloff VC (2014) Forest and agricultural land change in the Carpathian region—a meta-analysis of long-term patterns and drivers of change. Land Use Policy 38: 685–697. https://doi.org/10.1016/j.landusepol.2014.01.012
  • Nagy A, Rácz IA (2007) Az Orthoptera fauna védelmének prioritásai az Aggteleki Nemzeti Parkban. Állattani Közlemények 92: 53–65.
  • Nagy A, Kisfali M, Szövényi G, Puskás G, Rácz IA (2010) Distribution of Catantopinae species (Orthoptera: Acrididae) in Hungary. Articulata 25: 221–237.
  • Neff F, Korner-Nievergelt F, Rey E, Albrecht M, Bollmann K, Cahenzli F, Chittaro Y, Gossner MM, Martínez-Núñez C, Meier ES, Monnerat C, Moretti M, Roth T, Herzog F, Knop E (2022) Different roles of concurring climate and regional land-use changes in past 40 years' insect trends. Nature Communications 13: 7611. https://doi.org/10.1038/s41467-022-35223-3
  • Pavlović M (2019) Egyptian locust (Anacridium aegyptium) (Acrididae: Cyrtacanthacridinae) in the Pannonian part of Croatia. Articulata 34: 95–100.
  • Pisică EI, Iorgu IS, Murariu D (2008) Preliminary Data Regarding the Orthoptera (Insecta: Orthoptera) Specific Diversity from Bucharest Metropolitan Area. Analele Ştiinţifice ale Universităţii "Al. I. Cuza" Iaşi, s. Biologie animală, Tom LIV, 65–72.
  • Poniatowski D, Fartmann T (2010) What determines the distribution of a flightless bush-cricket (Metrioptera brachyptera) in a fragmented landscape? Journal of Insect Conservation 14: 637–645. https://doi.org/10.1007/s10841-010-9293-3
  • Prach K, Rehounková K, Lencová K, Jírová A, Konvalinková P, Mudrák O, Student V, Vaneček Z, Tichý L, Petrík P, Šmilauer P, Pyšek P (2013) Vegetation succession in restoration of disturbed sites in Central Europe: The direction of succession and species richness across 19 seres. Applied Vegetation Science 17: 193–200. https://doi.org/10.1111/avsc.12064
  • Puskás G, László M, Kiss O, Szövényi G (2024) The Egyptian Bird Grasshopper just landed in Pannonia. The present situation of Anacridium aegyptium in Hungary. In European Congress on Orthoptera Conservation (ECOC IV), Rovereto (Italy), 4–6 July 2024: Program and abstracts, 51–52.
  • Rácz IA (1998) Biogeographical survey of the Orthoptera fauna in the central part of the Carpathian Basin (Hungary): Fauna types and community types. Articulata 13(1): 53–69.
  • Ranner A, Riegler R (2004) Ein Vorkommen von Pezotettìx giornae (Rossi, 1794) (Caelifera: Catantopidae) im Nordburgenland. Beiträge zur Entomofaunistik 5: 125–128.
  • Redtenbacher J (1900) Die Dermapteren und Orthopteren (Ohrwürmer und Geradflügler) von Österreich-Ungarn und Deutschland. Carl Gerold's Sohn, Wien, 148 pp.
  • Šerić Jelaska L, Skejo J (2014) Catalogue of the entomological collections of the Division of Zoology of the Faculty of Science in Zagreb: Collection of Orthoptera (Polyneoptera, Orthopterida) of Boža Pokopac. Entomologia Croatica 18: 59–71.
  • Skejo J, Škorput J, Pavlović M, Tumbrinck J, Pakrac I, Rašan M, Ćato S, Srpak N, Deranja M, Adžić K, Kasalo N (2023) Endangered Transsylvanian wingless groundhopper (Tetrix transsylvanica) is not extinct in Croatia and requires urgent protection. Natura Croatica 32: 241–255. https://doi.org/10.20302/nc.2023.32.17
  • Skejo J, Stanković M (2014) Contribution to the knowledge of the Orthoptera of the Special Nature Reserve Zasavica (S Vojvodina, Serbia) with special emphasis on Zeuneriana amplipennis. Articulata 29: 9–20.
  • Staufer M, Forsthuber L (2021) Erste Vorkommen der allochthonen Geringelten Beißschrecke, Rhacocleis annulata Fieber, 1853, in Österreich und ihre Verbreitung im Pflanzenhandel (Orthoptera: Tettigoniidae). Beiträge zur Entomofaunistik 22: 33–42.
  • Špulerová J (2008) Land use changes in the Veselovianka river catchment in the Horná Orava region. Ekológia (Bratislava) 27: 326–337.
  • Szövényi G, Puskás G (2012) A Contribution to Knowledge Concerning the Orthoptera Fauna of the Slavonian Range (NE Croatia) with the First Record of Some Species in Croatia. Natura Croatica 21: 403–418.
  • Thorens P, Nadig A (1997) Atlas de distribution des Orthoptères de Suisse: Sauterelles, Grillons, Criquets (Orthoptera), et Mante religieuse (Mantodea). Neuchâtel: Centre Suisse de Cartographie de la Faune, Pro Natura.
  • Tscharntke T, Brandl R (2004) Plant-insect interactions in fragmented landscapes. Annual Review of Entomology 49: 405–430.
  • Verhees JJF (2024) Een populatie Rozevleugels (Calliptamus italicus) in groeve 't Rooth. Voorbode voor definitieve vestiging in Nederland? Natuurhistorisch Maandblad 113: 247–251.
  • Vlk R, Balvín O, Krištín A, Marhoul P, Hrúz V (2012) Distribution of the Southern Oak Bush-cricket Meconema meridionale (Orthoptera, Tettigoniidae) in the Czech Republic and Slovakia. Folia Oecologica 39: 155–165.
  • Wilson RJ, Fox R (2021) Insect responses to global change offer signposts for biodiversity and conservation. Ecological Entomology 46: 699–717. https://doi.org/10.1111/een.12970
  • Wolf S (2024) Erstnachweis der mediterranen Heuschreckenart Rhacocleis annulata Fieber, 1853 in der südlichen Oberrheinebene (Orthoptera: Tettigoniidae). Mitteilungen des Badischen Landesvereins für Naturkunde und Naturschutz 26: 143–146.
  • Wöss G (2017) Kleine Knarrschrecke Pezotettix giornae (Rossi, 1794). In: Zuna-Kratky T, Landmann A, Illich I, Zechner L, Essl F, Lechner K, Ortner A, Weißmair W, Wöss G (2017): Die Heuschrecken Österreichs. Denisia 39: 577–580.
  • Wöss G, Sehnal M, Berg C, Staufer M (2011) Erstnachweise der Kleinen Knarrschrecke Pezotettix giornae (Rossi, 1794) (Caelifera: Acrididae: Catantopinae) für Wien, Niederösterreich und Kärnten. Beiträge zur Entomofaunistik 12: 41–46.
  • Zechner L, Koschuh A, Berg HB, Paill W, Reinbacher H, Zuna-Kratky T (2005) Checkliste der Heuschrecken der Steiermark mit Kommentaren zu Verbreitung und Habitatansprüchen (Insecta: Orthoptera). Beiträge zur Entomofaunistik 6: 127–160.
  • Zuna-Kratky T, Landmann A, Illich I, Zechner L, Essl F, Lechner K, Ortner A, Weißmair W, Wöss G (2017) Die Heuschrecken Österreichs. Denisia 39: 880.