Antuluciella vittata (Philippi, 1859) comb. nov.

Euclidia vittata Philippi, 1859: 1113.

Scordylia vittata (Philippi). Butler 1882: 411; Bartlett-Calvert 1886: 339; Butler 1893: 461; Angulo & Casanueva 1981: 31 (Scordylla [sic]).

Hagnagora vittata (Philippi). Scoble 1999: 420; King & Parra 2011: 7; Brehm 2015: 154; Brehm et al. 2019: in phylogeny.

Heterusia mesenata Felder & Rogenhofer, 1875: pl. 130, fig. 26; Bartlett-Calvert 1886: 339. syn. nov.

Hoplosauris mesenata (Felder & Rogenhofer). Angulo & Casanueva 1981: 31.

Hagnagora mesenata (Felder & Rogenhofer). Scoble 1999: 420; Brehm 2015: 154; Brehm et al. 2019: 467.

Type material. Euclidia vittata. Holotype, female, Chile, Valdivia (apparently lost, according to MNNC). Heterusia mesenata. Lectotype (Fig 12a, designated here), female, Chile, with the following labels: “Novara CXXX Heterusia mesenata Chile”, “TYPE”, “BMNH(E) 1378770” (NHMUK), and “Genitalia slide Geom.1951-273” (examined).

Material examined. CHILE: Choapa Prov.: 1 female, Los Vilos, Quebrada de Quereo, 08-X-2020, leg. B. Salinas (MZUC-UCCC). Cordillera Prov.: 1 male and 1 female, San José de Maipo, El Manzano, 28-X-1951, leg. L.E. Peña (MZUC-UCCC); 1 male and 2 females, San José de Maipo, El Canelo, 04-XI-1949, leg. n.n. (MZUC-UCCC); 1 male, San José de Maipo, El Canelo, 20-X-1949, leg. n.n. (MZUC-UCCC); 1 female, San José de Maipo, El Canelo, IX-1951, leg. L.E. Peña (MZUC-UCCC); 2 males, San José de Maipo, El Canelo, 29-IX-1935, leg. E. Ureta (MNNC). Melipilla Prov.: 1 male and 1 female, Alhué, 18-IX-1947, leg. Barros (MZUC-UCCC); 1 female, Alhué, 17-IX-1947, leg. Barros (MZUC-UCCC). Diguillín Prov.: 1 male and 1 female, Pinto, Atacalco, 28-XI-1951, leg. L.E. Peña (MZUC-UCCC); 1 male, Las Trancas, 26-XII-2004, leg. G. Moreno (MZUC-UCCC). Concepción Prov.: 1 male, Concepción, 06-X-1955, leg. J. Artigas (MZUC-UCCC); 1 male, Concepción, 16- XII-1958, leg. J. Artigas (MZUC-UCCC); 1 female, Concepción, 25/ 30-IX-2001, leg. J. Artigas (MZUC-UCCC); 1 male, Concepción, 16-XI-1958, leg. J. Artigas (MZUC-UCCC); 1 male, Concepción, 23-X-1962, leg. traps (MZUC-UCCC); 1 female, Concepción, Fundo El Guindo, 24-IX-2008, leg. Hernández (MZUC-UCCC); 1 male, Concepción, Fundo El Guindo, 05-X-2008, leg. Hernández (MZUC-UCCC); 1 male, Universidad de Concepción, 28-XI-2012, leg. F. Venegas (MZUC-UCCC); 1 female, Concepción, Fundo El Guindo, 17-XI-2014, leg. M. Ramos & C. Rose (MZUC-UCCC); 2 males and 1 female, Concepción, Fundo El Guindo, 21-X-2022, leg. M. Ramos & C. Rose (MZUC-UCCC); 1 male, Universidad de Concepción, 15-XII-2022, leg. M. Ramos (MZUC-UCCC); 1 male, Concepción, Cerro Caracol (P. Valdivia, “lab. Raise ”), 27-X-2022, labeled “reared on Fuchsia magellanica, Luma apiculata and Nothofagus obliqua ” and “ Agni, egg 06-XI-2021, larva emerged 13-XI-2021, pupa 13-XII-2021, adult emerged 27-X-2022 ”, leg. M. Ramos (MZUC-UCCC); 1 male and 1 female, Concepción, Cerro Caracol, 06- XI-2021, leg. J. Cancino (MZUC-UCCC); 1 female, Talcahuano, Parque Tumbes, 06-I-2024, leg. M. Ramos & C. Rose (MZUC-UCCC); 1 female, Hualpén, 30-XI-1998, leg. P. Bocaz (MZUC-UCCC); 2 females, Hualpén, Estación de Biología Terrestre, 12-X-2017, leg. L. E. Parra (MZUC-UCCC). Arauco Prov.: 2 females, Cañete, Butamalal, 12-III-1966, leg. T. Cekalovic (MZUC-UCCC). Biobío Prov.: 1 male, Tucapel, 11-X-2021, leg. F. Susana (MZUC-UCCC). Cautín Prov.: 2 males, Pucón, Termas de Río Blanco, II-1951, leg. L. Peña (MZUC-UCCC); 1 male, Pucón, Camino al Volcán, 17-X-2017, leg. H. Salazar (MZUC-UCCC). Osorno Prov.: 1 male, Puyehue, Aguas Calientes, 21-XI-1999, leg. M. Beeche (MZUC-UCCC). Llanquihue Prov.: 1 female, Maullín, 20-X-1949, leg. Barros (MZUC-UCCC); 1 female, Maullín, 17-X-1941, leg. Barros (MZUC-UCCC).

Literature records. CHILE: Valparaíso Prov.:Valparaíso (August–September) (Butler, 1882). ARGENTINA: Neuquén Prov. (Chalup 2020).

Additional records. CHILE: Choapa Prov.: Los Vilos, Quebrada de Quereo, -31.9299, -71.5098, 05-IX-2021, observed by Diego Almendras and submitted to iNaturalist in: https://inaturalist.mma.gob.cl/ observations/186754986. Valparaíso Prov.: Zapallar, -32.5547, -71.4681, 15-VIII-2016, observed by Cristóbal Costoya and submitted to iNaturalist in: https://inaturalist.mma.gob.cl/observations/229027654; Viña del Mar, - 33.0355, -71.5488, 01-IX-2024, observed by Franco Allende and submitted to iNaturalist in: https://inaturalist. mma.gob.cl/observations/239319265; Valparaíso, Las Docas, -33.1518, -71.6990, 25-IX-2023, observed by Jorgemu and submitted to iNaturalist in: https://inaturalist.mma.gob.cl/observations/184936085. San Antonio Prov.: 2 records from El Quisco, El Totoral, -33.4132, -71.6434, 06-X-2022 / 22-IX-2023, observed by Eric Rojas and submitted to iNaturalist in: https://inaturalist.mma.gob.cl/observations/137814518 and https://inaturalist.mma.gob. cl/observations/184454961; 2 records from El Tabo, Quebrada de Córdova, -33.4421, -71.6606, 05-IX-2021 / 16- IX-2021, observed by Ernesto Guzmán and submitted to iNaturalist in: https://inaturalist.mma.gob.cl/ observations/122315217 and https://inaturalist.mma.gob.cl/observations/93763260; El Tabo, Quebrada de Córdova, -33.4426, -71.6673, 12-IX-2024, observed by Isadora and submitted to iNaturalist in: https://inaturalist.mma.gob. cl/observations/241270547; El Tabo, -33.4536, -71.6686, 27-IX-2021, observed by Isadora and submitted to iNaturalist in: https://inaturalist.mma.gob.cl/observations/245073558; El Tabo, -33.4453, -71.6486, 13-X-2024, observed by Rachel Jamett Román and submitted to iNaturalist in: https://inaturalist.mma.gob.cl/ observations/247246505; Cartagena, Quebrada El Tranque, -33.5462, -71.5934, observed by Miguel Patiño and submitted to iNaturalist in: https://inaturalist.mma.gob.cl/observations/98427048; Santo Domingo, Parque Tricao (Quebrada Chincol), -33.7312, -71.6352, 29-IX-2022, observed by Josefina Rodríguez and submitted to iNaturalist in: https://inaturalist.mma.gob.cl/observations/142456250. Santiago Prov.: Las Condes, Quebrada San Ramón, - 33.4342, -70.5034, 09-X-2020, observed by Orlando Montes and submitted to iNaturalist in: https://inaturalist. mma.gob.cl/observations/62212416; Las Condes, Quebrada San Ramón, -33.4366, -70.4943, 05-X-2020, observed by Camilo Pavez and submitted to iNaturalist in: https://inaturalist.mma.gob.cl/observations/245929926; Las Condes, Quebrada San Ramón, -33.4362, -70.5007, 27-IX-2024, observed by John Horsfall and submitted to iNaturalist in: https://inaturalist.mma.gob.cl/observations/245943064. Cardenal Caro Prov.: Litueche, Topocalma, -34.1947, -71.8650, X-2023, observed by Denisse del Campo and submitted to iNaturalist in: https://inaturalist. mma.gob.cl/observations/187616369. Talca Prov.: San Clemente, Altos de Lircay N.P., -35.5932, -71.0073, 01-XI-2022, observed by Vale Guzmán Salazar and submitted to iNaturalist in: https://inaturalist.mma.gob.cl/ observations/141487163. Punilla Prov.: San Fabián de Alico, El Caracol, -36.6572, -71.3846, 12-X-2019, observed by Alexandra Castaneda and submitted to iNaturalist in: https://inaturalist.mma.gob.cl/observations/34279281. Diguillín Prov.: Pinto, Los Lleuques (Las Turbinas), -36.8717, -71.6528, 23-X-2022, observed by Romy Weinborn and submitted to iNaturalist in: https://inaturalist.mma.gob.cl/observations/139786563. Itata Prov.: Coelemu, Estero Guarilihue, -36.5672, -72.6892, 07-X-2021, observed by Claudio Maureira and submitted to iNaturalist in: https://inaturalist.mma.gob.cl/observations/101849671. Concepción Prov.: Penco, -36.7547, -72.9542, 16-XII-2019, observed by Orlando Montes and submitted to iNaturalist in: https://inaturalist.mma.gob.cl/ observations/36776128; Penco, -36.7453, -72.982, 04-XI-2019, observed by Orlando Montes and submitted to iNaturalist in: https://inaturalist.mma.gob.cl/observations/174148108; Concepción, -36.7968, -73.0195, 02-XI-2022, observed by Antonio Maureira and submitted to iNaturalist in: https://inaturalist.mma.gob.cl/ observations/142706006; Hualpén, Caleta Chome, -36.7704, -73.2068, 13-IX-2022, observed by Antonio Maureira and submitted to iNaturalist in: https://inaturalist.mma.gob.cl/observations/141032830; Coronel, Lomas Coloradas, -36.9096, -73.1270, 21-X-2023, observed by Carolina Sobarzo and submitted to iNaturalist in: https://inaturalist. mma.gob.cl/observations/190026168; Chiguayante, -36.9385, -73.0054, 23-XI-2018, observed by Claudio Maureira and submitted to iNaturalist in: https://inaturalist.mma.gob.cl/observations/26132981; Chiguayante, -36.9248, - 72.9796, 07-X-2020, observed by Claudio Maureira and submitted to iNaturalist in: https://inaturalist.mma.gob.cl/ observations/62466121; Chiguayante, -36.9359, -73.0002, 07-X-2020, observed by Claudio Maureira and submitted to iNaturalist in: https://inaturalist.mma.gob.cl/observations/62466170. Biobío Prov.: Tucapel, Polcura, -37.2862, -71.7268, 07-XI-2022, observed by Flor Susana and submitted to iNaturalist in: https://inaturalist.mma.gob.cl/ observations/142183354; Tucapel, -37.3016, -71.9526, 11-X-2021, observed by Flor Susana and submitted to iNaturalist in: https://inaturalist.mma.gob.cl/observations/97994449. Malleco Prov.: Angol, Santa Elena, 23-IX-2020,observed by Mario Ramos and submitted to iNaturalist in:https://inaturalist.mma.gob.cl/observations/61204466; Curacautín, Laguna Blanca, -38.3696, -71.6565, 06-XII-2023, observed by Romipendola and submitted to iNaturalist in: https://inaturalist.mma.gob.cl/observations/193266940. Cautín Prov.: Vilcún, -38.6319, -72.2107, 08-XII-2020, observed by Nitsuga74 and submitted to iNaturalist in: https://inaturalist.mma.gob.cl/observations/66292278; Temuco, -38.7520, -72.6629, 19-X-2021, observed by Benjamín Silva and submitted to iNaturalist in: https:// inaturalist.mma.gob.cl/observations/109338512; Temuco, Bosque Vivo CMPC Pumalal Park, -38.6472, -72.5596, 13-X-2024, observed by Nitsuga74 and submitted to iNaturalist in: https://inaturalist.mma.gob.cl/ observations/247160864; Pucón, -39.2988, -71.8888, 02-X-2022, observed by Claudia Ríos and submitted to iNaturalist in: https://inaturalist.mma.gob.cl/observations/137305842; 2 records from Villarrica, Ñancul, -39.2799, -72.3214, 13-XII-2023 / 16-IX-2024, observed by Aart Broekema and submitted to iNaturalist in: https://inaturalist. mma.gob.cl/observations/194240309 and https://inaturalist.mma.gob.cl/observations/242196014; Pucón, Pichare, -39.2538, -71.7383, 27-XII-2023, observed by Rajan Rao and submitted to iNaturalist in: https://inaturalist.mma. gob.cl/observations/195220415; Villarrica, Correntoso, -39.3396, -72.0924, 31-X-2023, observed by Michael Weymann and submitted to iNaturalist in: https://inaturalist.mma.gob.cl/observations/189700057; Villarrica, Licanray, -39.4875, -72.1475, 30-X-2021, observed by Alexis Catalán and submitted to iNaturalist in: https:// inaturalist.mma.gob.cl/observations/100029622. Valdivia Prov.: Panguipulli, Tracalpulli, -39.5836, -72.1907, 17- X-2020, observed by Yerko Lloncon and submitted to iNaturalist in: https://inaturalist.mma.gob.cl/ observations/62851963; Valdivia, Humedal Angachilla, -39.8562, -73.2318, 09-X-2022, observed by Gabriela Antonia and submitted to iNaturalist in: https://inaturalist.mma.gob.cl/observations/138320020; Valdivia, Isla Teja, -39.8070, -73.2529, 22-XI-2022, observed by Nightmare8 and submitted to iNaturalist in: https://inaturalist.mma. gob.cl/observations/143649636; Valdivia, Niebla, -39.8656, -73.3945, 29-X-2022, observed by Nodora Moyano and submitted to iNaturalist in: https://inaturalist.mma.gob.cl/observations/141423674; Los Lagos, Mocho-Choshuenco NR, -39.9385, -72.0964, 14-XI-2022, observed by René Fuentes and submitted to iNaturalist in: https://inaturalist.mma.gob.cl/observations/207980687. Osorno Prov.: Osorno, Rahue River, -40.5562, -73.1623, 28-IX-2022, observed by Ricardo Huenuanca and submitted to iNaturalist in: https://inaturalist.mma.gob.cl/ observations/136878192; Puyehue, Rupanco Lake, -40.7657, -72.62819, 03-X-2023, observed by Daniel Stange and submitted to iNaturalist in: https://inaturalist.mma.gob.cl/observations/186069493; Puyehue, Aguas Calientes, -40.7392, -72.3070, observed by Tristan Jobin and submitted to iNaturalist in: https://inaturalist.mma.gob.cl/ observations/147108624. Llanquihue Prov.: Purranque, Hueyusca, -40.9266, -73.5295, 19-XII-2023, observed by Fernanda Zamorano and submitted to iNaturalist in: https://inaturalist.mma.gob.cl/observations/194401175; Frutillar, -41.0890, -73.0566, 27-XI-2024, observed by Paulina Hechenleither and submitted to iNaturalist in: https://inaturalist.mma.gob.cl/observations/253320036; Puerto Varas, -41.3205, -72.9846, 28-XI-2024, observed by Ricardo Huenuanca and submitted to iNaturalist in: https://inaturalist.mma.gob.cl/observations/253441838; Puerto Varas, -41.3829, -73.0422, 30-IX-2024, observed by Gina Dnisse and submitted to iNaturalist in: https://inaturalist. mma.gob.cl/observations/244965263; Cochamó, La Junta, -41.4118, -72.1257, 25-XI-2022, observed by Rocío Elisa and submitted to iNaturalist in: https://inaturalist.mma.gob.cl/observations/144035937; Puerto Montt, Katalapi Park, -41.5216, -72.7535, 14-X-2018, observed by Carlos Zamora and submitted to iNaturalist in: https://inaturalist. mma.gob.cl/observations/34541922. Chiloé Prov.: Ancud, Lechagua, -41.8802, -73.8789, 30-XI-2024, observed by Marisella Vargas and submitted to iNaturalist in: https://inaturalist.mma.gob.cl/observations/253649885; Ancud, Senda Darwin Biological Station, -41.8815, -73.666, 05-XI-2020, observed by Ericlopresti and submitted to iNaturalist in: https://inaturalist.mma.gob.cl/observations/52665305; Ancud, Manao, -41.8893, -73.5189, 15-X-2020, observed by Leonardo Tempesta and submitted to iNaturalist in: https://inaturalist.mma.gob.cl/ observations/62675001; Castro, -42.4858, -73.7769, 08-IX-2024, observed by Verónica Díaz and submitted to iNaturalist in: https://inaturalist.mma.gob.cl/observations/240640971; Chonchi, Quinched, -42.5780, -73.7598, 01- XI-2020, observed by Natalia Carrasco and submitted to iNaturalist in: https://inaturalist.mma.gob.cl/ observations/64307734. Palena Prov.: Hualaihué, Vodudahue, -42.4822, -72.3866, observed by Rodrigo Barahona and submitted to iNaturalist in: https://inaturalist.mma.gob.cl/observations/109722984. Aysén Prov.: Puerto Aysén, -45.4169, -72.6754, 22-X-2023, observed by Ernesto Guzmán and submitted to iNaturalist in: https://inaturalist. mma.gob.cl/observations/188593886; Puerto Aysén, Simpson River, -45.4532, -72.4225, 23-XII-2023, observed by Ernesto Guzmán and submitted to iNaturalist in: https://inaturalist.mma.gob.cl/observations/194823718. ARGENTINA: Neuquén Prov.: Huiliches, Laguna Verde, -39.8374, -71.5238, 21-XII-2024, observed by Gra-moll and submitted to iNaturalist in: https://inaturalist.mma.gob.cl/observations/258755710; Los Lagos, Lago Espejo Chico, -40.5947, -71.7169, 24-XII-2022, observed by Gra-moll and submitted to iNaturalist in: https:// inaturalist.mma.gob.cl/observations/148697209; Los Lagos, Lago Espejo Grande, -40.6716, -71.7356, 25-XII-2021,observedbyLeonelRogetandsubmittedtoiNaturalistin:https://inaturalist.mma.gob.cl/observations/104745053; Los Lagos, Villa Angostura, -40.751, -71.6484, 05-I-2022, observed by Tilda Rosander and submitted to iNaturalist in: https://inaturalist.mma.gob.cl/observations/105394323. Río Negro Prov.: Bariloche, Lago Gutiérrez, -41.1939, -71.4206, observed by Gustavo Fernando Durán and submitted to iNaturalist in: https://inaturalist.mma.gob.cl/ observations/193921363; Bariloche, Tambo de Báez, -41.14, -71.4762, 04-I-2025, observed by Ezequiel Racker and submitted to iNaturalist in: https://inaturalist.mma.gob.cl/observations/257411412.

Diagnosis. Same as for the genus.

Redescription. Adults (Fig. 1a). Head with dark brown filiform antennae in both sexes; frons and vertex covered with flattened dark brown scales; labial palpi dark brown, porrect. Thorax with patagia and tegulae bearing orange scales mottled with dark brown scales; legs grayish-white; tibial formula 0-2-4. Abdomen brownish-orange. Forewings subtriangular, bright orange speckled with brown scales toward the base, termen slightly bulged between M 1 and CuA 2; medial band blackish, oblique and slightly sinuate, extending from the costa to the anal angle; terminal band blackish, strongly broadened towards the apex; costa with brownish-gray scales on the proximal half, yellowish-white between the medial and terminal bands up to the level of R 2. Hindwings trapezoidal, bright orange speckled with brown scales toward the base, termen rounded; postmedial band blackish, slightly sinuate; terminal band blackish, subapically broadened and narrowing toward the anal angle; costa with a blackish stripe, interrupted at the postmedial band level in some specimens revealing the ground color, in others continuous. Male genitalia (Fig. 6a). Uncus hook-shaped, apex acute; gnathos absent; valvae scalpel-shaped, costa slightly sclerotized, with an apical spine and a basal brush of curved and elongated androconial setae; sacculus sclerotized; juxta mushroom-shaped, distinctly wider distally; transtilla projected as a falciform process; saccus subquadrate, strongly sclerotized, posterior margin subtriangular. Aedeagus tubular, slightly sinuate; vesica armed with a group of cornuti. Female genitalia (Fig. 6b). Corpus bursae membranous and globose in the distal half, proximal half strongly sclerotized and slightly twisted; signum lanceolate; ductus bursae short, 1/3 the length of the corpus bursae; Antrum short, broad, funnel-shaped; anterior apophyses half the length of the posterior apophyses.

Distribution. As for the genus.

Flight period. Adults have been recorded from August to March.

Biology. Immature stages, host plant, and life cycle were described by King & Parra (2011). Based on the rearing of a clutch of 40 eggs of this species, we report that the larvae, in addition to being associated with the foliage and fruits of “chilco ” (Fuchsia magellanica; Onagraceae), also feed on the foliage of “arrayán ” (Luma apiculata; Myrtaceae) and “hualle” (Nothofagus obliqua; Nothofagaceae), although feeding on this last host was observed only in the absence of the other plants and at low frequency. Larval eclosion takes place after 7 days (n= 40). Larval development extends for 30 days (n= 32), after which pupation occurs, and the adult of one reared individual emerged after 318 days (n= 1). During the day, adults visit flowers of native plants such as Cryptocarya alba, Leptocarpha rivularis, Gaultheria mucronata, and F. magellanica, in addition to flowers of exotic plants such as Myosotis sp., Prunus sp., Cardamine sp., Brassica rapa, Anthriscus caucalis, Erigeron karvinskianus, and Rubus ulmifolius.