Published March 23, 2026 | Version v1

Phytoseiidae Berlese 1916

  • 1. Laboratory of Ecological Information, Graduate School of Agriculture, Kyoto University, Kyoto 606 - 8502, Japan & Department of Plant Protection, Faculty of Agriculture, Assiut University, Assiut 71526, Egypt
  • 2. NARO headquarters, Tsukuba, Ibaraki 305 - 8517, Japan
  • 3. Institute of Plant Protection, NARO, Morioka, Iwate 020 - 0123, Japan
  • 4. Laboratory of Ecological Information, Graduate School of Agriculture, Kyoto University, Kyoto 606 - 8502, Japan & Emeritus Professor, Kyoto University, Kyoto 606 - 8502, Japan & Laboratory of Ecological Information, Graduate School of Agriculture, Kyoto University, Kyoto 606 - 8502, Japan

Description

Key to subfamilies, tribes and genera of the Phytoseiidae in Japan (females)

1. Setae z3 and s6 absent (with four prolateral setae)............ Amblyseiinae Muma, 1961 ....................... 8

- Either or both setae z3 and s6 present (with more than four prolaterals)........................................ 2

2. Setae Z1, S2, S4, S5 absent............ Phytoseiinae Berlese, 1913 ...................... Phytoseius Ribaga, 1904

- At least one of setae Z1, S2, S4 or S5 present............ Typhlodrominae Scheuten, 1857 ...................... 3

3. Seta s6 absent............ Chanteiini Chant & McMurtry, 1994 ............... Chanteius contiguus (Chant, 1959)

- Seta s6 present.................................................................................... 4

4. Seta z6 present............ Paraseiulini Wainstein, 1976 ................................................. 7

- Seta z6 absent..................................................................................... 5

5. Setae S4 and JV4 present............ Typhlodromini Wainstein, 1962 .............. Typhlodromus Scheuten, 1857

- Setae S4 and JV4 absent............ Metaseiulini Chant & McMurtry, 1994 .................................. 6

6. Seta R1 absent; seta S2 present.................................. Galendromus (G.) occidentalis (Nesbitt, 1951)

- Seta R1 present; seta S2 absent..................................... Typhlodromina conspicua (Garman, 1948)

7. Seta JV2 absent....................................................... Paraseiulus soleiger (Ribaga, 1902)

- Seta JV2 present............................................ Kuzinellus yokogawae (Ehara & Hamaoka, 1980)

8. Peritremal shield not fused anteriorly with dorsal shield............ Indoseiulini Ehara & Amano, 1998 ...................................................................................... Gynaeseius Wainstein, 1962

- Peritremal shield fused anteriorly with dorsal shield....................................................... 9

9. Sternal shield with median posterior projection; pre-anal setae JV2 and ZV2 migrated forward............ Euseiini Chant & McMurtry, 2005a ............................................................................... 20

- Sternal shield without posterior projection; without forward migration of pre-anal setae JV2 and ZV2 .............. 10

10. Seta S4 absent.................................................................................... 14

- Seta S4 present................................................................................... 11

11. Ratio seta s4: Z1> 3:1; setae s4, Z4 and usually Z5 distinctly longer than other dorsal setae; seta J2 present/absent............ Amblyseiini Muma, 1961 ...................................................................... 17

- Ratio seta s4: Z1 <3:1; setae s4, Z4 and usually Z5 not greatly longer than other dorsal setae; seta J2 present......... 12

12. GeII and GeIII rarely with macroseta; fixed digit of chelicera usually with fewer than six teeth..... Neoseiulini Chant & McMurtry, 2003a .............................................................. Neoseiulus Hughes, 1948

- GeII and GeIII usually with macrosetae; fixed digit of chelicera usually with more than six teeth............ Typhlodromipsini Chant & McMurtry, 2005b .......................................................................... 13

13. Dorsal shield with waist at level of seta R1; seta R1 not inserted on dorsal shield; posterior margin of sternal shield straight or concave and never with posterior projection................................... Typhlodromips De Leon, 1965

- Dorsal shield without waist at level of seta R1; seta R1 inserted on edge of the dorsal shield; posterior margin of sternal shield sometimes with a median projection, lightly sclerotised or indistinct Scapulaseius Karg & Oomen-Kalsbeek, 1987

14. Ventrianal shield usually long, narrow, with 3 pairs of pre-anal setae............ Kampimodromini Kolodochka, 1998 ............................................................................................... 15

- Ventrianal shield reduced, neither long nor narrow, with 0-1 pair of pre-anal setae............ Phytoseiulini Chant & McMurtry, 2006 ................................................ Phytoseiulus persimilis Athias-Henriot, 1957

15. Without distinctive notch or incision in lateral margin of dorsal shield at level of seta s4.......... Okiseius Ehara, 1967

- With distinctive notch or incision in lateral margin of dorsal shield at level of seta s4 ........................... 16

16. Setae S2 present............................................................. Amblyseiulella Muma, 1961

- Setae S2 absent.................................................. Paraphytoseius Swirski & Schechter, 1961

17. Sternal shield narrow; ventrianal shield usually longer than wide; setae z2 and z4 usually short/minute; setae j5, S2 and S4 present, setae J2, S5 and Z1 present/absent............................................................. 19

- Sternal shield broad; ventrianal shield usually broader; legs II–IV with/without macrosetae; setae z2 and/or z4 often longer; setae j5, J2, S2, S4 and/or Z1 present/absent............................................................ 18

18. Seta J2 absent, seta j5 present.................................................. Proprioseiopsis Muma, 1961

- Seta J2 usually present or, if absent, seta j5 also absent............. Phytoscutus japonicus Ehara & Kishimoto, 2007a

19. Ratio seta s4: S2 <2.7:1................................................. Transeius Chant & McMurtry, 2004

- Ratio seta s4: S2> 3:1........................................................... Amblyseius Berlese, 1914

20. Chelicerae normal in size and shape, with teeth evenly distributed along fixed digit; peritremes usually extending to level of seta j1............................................ Amblydromalus limonicus (Garman & McGregor, 1956)

- Chelicerae reduced in size, with a group of teeth clustered at distal end of fixed digit; peritremes usually not extending beyond setae j3................................................................................... 21

21. Ventrianal shield divided into separate ventral and anal shields................. Iphiseius degenerans (Berlese, 1889)

- Ventrianal shield entire........................................................... Euseius Wainstein, 1962

Notes

Published as part of Negm, Mohamed W., Toyoshima, Shingo, Kishimoto, Hidenari & Amano, Hiroshi, 2026, Phytoseiid mites of Japan (Acari: Mesostigmata), pp. 51-85 in Zootaxa 5782 (1) on pages 74-75, DOI: 10.11646/zootaxa.5782.1.3, http://zenodo.org/record/19181174

Files

Files (7.9 kB)

Name Size Download all
md5:a3d057b9708c45ecc2c92de8f1773037
7.9 kB Download

System files (41.7 kB)

Name Size Download all
md5:6d93707d935f5dd28721cb56ac0a8d8e
41.7 kB Download

Linked records

Additional details

Biodiversity

Scientific name authorship
Berlese
Kingdom
Animalia
Phylum
Arthropoda
Order
Mesostigmata
Family
Phytoseiidae
Taxon rank
family
Taxonomic concept label
Phytoseiidae Berlese, 1916 sec. Negm, Toyoshima, Kishimoto & Amano, 2026

References

  • Muma, M. H. (1961) Subfamilies, genera and species of Phytoseiidae (Acarina: Mesostigmata). Bulletin of the Florida State Museum, 5, 267-302. https://doi.org/10.58782/flmnh.tqpo4380
  • Berlese, A. (1913) Acaroteca Italica. Tipografia di M. Ricci, Firenze, 221 pp.
  • Ribaga, C. (1904) (1902) Gamasidi planticoli. Rivista di Patologia Vegetale, 10, 175-178.
  • Scheuten, A. (1857) Einiges uber Milben. Archiv fur Naturgeschichte, 23, 104-112.
  • Chant, D. A. & McMurtry, J. A. (1994) A review of the subfamilies Phytoseiinae and Typhlodrominae (Acari: Phytoseiidae). International Journal of Acarology, 20, 223-310. https://doi.org/10.1080/01647959408684022
  • Chant, D. A. (1959) Phytoseiid mites (Acarina: Phytoseiidae). Part I. Bionomics of seven species in southern England. Part II. A taxonomic review of the family Phytoseiidae, with descriptions of 38 new species. The Canadian Entomologist, 91, 1-166.
  • Wainstein, B. A. (1976) A new tribe of the family Phytoseiidae (Parasitiformes). Zoologicheskii Zhurnal, 55, 696-700.
  • Wainstein, B. A. (1962) Revision du genre Typhlodromus Scheuten, 1857 et systematique de la famille des Phytoseiidae (Berlese, 1916) (Acarina: Parasitiformes). Acarologia, 14, 5-30.
  • Nesbitt, H. (1951) A taxonomic study of the Phytoseiinae (Family Laelaptidae) predacious upon Tetranychidae of economic importance. Zoologische Verhandelingen, 12, 1-64.
  • Garman, P. (1948) Mite species from apple trees in Connecticut. Bulletin of the Connecticut Agricultural Experimental Station, 520, 5-27.
  • Ehara, S. & Hamaoka, K. (1980) A new Typhlodromus from Japan with notes on four other species of phytoseiid mites. Acta Arachnologica, 29, 3-8. https://doi.org/10.2476/asjaa.29.3
  • Ehara, S. & Amano, H. (1998) A revision of the mite family Phytoseiidae in Japan (Acari: Gamasina), with remarks on its biology. Species Diversity, 3, 25-73. https://doi.org/10.12782/specdiv.3.25
  • Chant, D. A. & McMurtry, J. A. (2005 a) A review of the subfamily Amblyseiinae Muma (Acari; Phytoseiidae): Part VI. The tribe Euseiini n. tribe, subtribes Typhlodromalina n. subtribe, Euseiini n. subtribe, and Ricoseiina n subtribe. International Journal of Acarology, 31, 187-224. https://doi.org/10.1080/01647950508684424
  • Chant, D. A. & McMurtry, J. A. (2003 a) A review of the subfamily Amblyseiinae Muma (Acari: Phytoseiidae): Part I. Neoseiulini new tribe. International Journal of Acarology, 29, 3-46. https://doi.org/10.1080/01647950308684319
  • Hughes, A. M. (1948) The mites associated with stored food products. Ministry of Agriculture and Fisheries, H. M. Stationary Office, London, 168 pp.
  • Chant, D. A. & McMurtry, J. A. (2005 b) A review of the subfamily Amblyseiinae Muma (Acari: Phytoseiidae): Part VII. Typhlodromipsini n. tribe. International Journal of Acarology, 31, 315-340. https://doi.org/10.1080/01647950508683673
  • De Leon, D. (1965) A note on Neoseiulus Hughes 1948 and a new synonymy. Proceedings of the Entomological Society of Washington, 87, 23.
  • Karg, W. & Oomen-Kalsbeek, F. (1987) Neue Raubmilbenarten der Gattung Amblyseius Berlese (Acarina, Parasitiformes, Phytoseiidae) - Antagonisten der unechten Spinnmilbe Brevipalpus phoenicis Geijskes. Zoologische Jahrbucher Systematik, 114, 131-140.
  • Kolodochka, L. A. (1998) Two new tribes and the main results of a revision of Palearctic phytoseiid mites (Parasitiformes, Phytoseiidae) with the family system concept. Vestnik Zoologii, 32, 51-63. [in Russian]
  • Chant, D. A. & McMurtry, J. A. (2006) A review of the subfamily Amblyseiinae Muma (Acari: Phytoseiidae): Part VIII. The tribes Macroseiini Chant, Denmark and Baker, Phytoseiulini n. tribe, Afroseiulini n. tribe and Indoseiulini Ehara & Amano. International Journal of Acarology, 32, 13-25. https://doi.org/10.1080/01647950608684439
  • Athias-Henriot, C. (1957) Phytoseiidae et Aceosejidae (Acarina: Gamasina) d'Algerie. I. Genres Blattisocius Keegan, Iphiseius, Berlese, Amblyseius Berlese, Phytoseius Ribaga, Phytoseiulus Evans. Bulletin de la Societe d'Histore Natrelle de l'Afrique du Nord, 48, 319-352.
  • Ehara, S. (1967 a) Phytoseiid mites from Hokkaido (Acarina: Mesostigmata). Journal of the Faculty of Science, Hokkaido University, Series VI, Zoology, 16, 212-233.
  • Swirski, E. & Schechter, R. (1961) Some Phytoseiid mites (Acarina: Phytoseiidae) of Hong Kong, with a description of a new genus and seven new species. Israel Journal of Agricultural Research, 11, 97-117.
  • Ehara, S. & Kishimoto, H. (2007 a) Description of a new species of Phytoscutus (Acari: Phytoseiidae) from Kyushu, Japan. International Journal of Acarology, 33, 111-113. https://doi.org/10.1080/01647950708684509
  • Chant, D. A. & McMurtry, J. A. (2004) A review of the subfamily Amblyseiinae Muma (Acari: Phytoseiidae): Part III. The tribe Amblyseiini Wainstein, subtribe Amblyseiina n. subtribe. International Journal of Acarology, 30, 171-228. https://doi.org/10.1080/01647950408684388
  • Berlese, A. (1914) Acari nuovi. Manipulus IX. Redia, 10, 113-150.
  • Garman, P. & McGregor, E. A. (1956) Four new predaceous mites (Acarina: Phytoseiidae). Southern California Academy of Science Bulletin, 55, 7-13.
  • Berlese, A. (1889) Acari, Myriapoda et Scorpiones hucusque in Italia seperta. Tipografia del Seminario, Padova, 4 pls. [pls. 6 + 7-9]