Scione maculipennis (Schiner, 1868)

(Figs 54A–D, 55A–F)

Diclisa maculipennis Schiner, 1868: 102, pl. 2, figs.; Röder, 1886: 262 (tax.); 1892: 8 (tax.); Osten Sacken, 1886: 47 (tax.); Kertész, 1900: 14 (cat.); Ricardo, 1900a: 104 (NHMUK spp.); Hunter, 1900: 135 (cat.).

Rhinotriclista maculipennis; Kröber, 1930: 165, fig. 10 (rev.).

Scione maculipennis; Ricardo, 1902: 435 (NHMUK spp.); Aldrich, 1905: 192 (cat.); Kertész, 1908: 150 (cat.); Hine, 1925: 9 (tax.); Surcouf, 1919: 221 (Ecuador spp.); 1921: 113 (comm.); Szilády, 1926: 28 (rev.); Kröber, 1934: 234 (cat.); Fairchild, 1942: 189, figs. 1–2, 5–6 (tax.); Bequaert & Renjifo-Salcedo, 1946: 62 (Colombia spp.); Fairchild, 1947: 566 (tax.); 1961: 25 (Costa Rica spp.); 1967c: 248, fig. 7 (Schiner spp.); 1971: 26 (cat.); Moucha, 1976: 40 (cat.); Hogue & Fairchild, 1974: 14 (Costa Rica spp.); Wilkerson, 1979: 168, fig. 39 (Colombia spp.); Fairchild, 1986: 31 (Panama spp.); Fairchild & Burger, 1994: 50 (cat.); Cárdenas & Vieira, 2005: 154 (Ecuador spp.); Coscarón & Papavero, 2009: 53 (cat.); Buestán et al., 2007: 32 (tax.); Henriques, 2016: 176 (AMNH spp.).

Scione aurea Szilády, 1926: 28 (rev.); Kröber, 1930: 144 (as synonym for S. minor) (rev.); 1934: 234 (cat.); Fairchild, 1961: 25 (Costa Rica spp.); 1971: 27 (as synonym for S. maculipennis) (cat.); Fairchild & Burger, 1994: 50 (cat.); Coscarón & Papavero, 2009: 53 (cat.).

Type locality. “ South America ” (Venezuela).

Diagnosis. Body orange brown. Thorax with conspicuous whitish longitudinal stripes. Wing smoky brown with yellow patch in the middle of cells r 1 and r 2+3.

Redescription. Female. Body length: 11.2–13.4 mm (X = 12.2 mm, N = 13), wing length: 11–13.2 mm (X = 12.1 mm, N = 14).

Head. Frons nearly parallel, divergence index 0.8–0.9 (X = 0.9, N = 14), frontal index 2.0–2.4 (X = 2.2, N = 14). Vertex dark brown with long, dense black setulae, ocellar triangle little prominent, ocelli yellowish-brown. Frons and subcallus brown with yellowish pruinosity, frons with long, dense black setulae. Parafacial brown with sparse black setulae. Gena with white pruinosity and long, dense white setulae. Clypeus brown with sparse black setulae, clypeal index 0.7–0.9 (X = 0.8, N = 14). Antenna orange brown. Scape and pedicel with yellowish pruinosity and short, dense black setulae. Palpus orange brown; first segment with black and yellowish setulae; second segment with sparse black setulae and quite tapered at the tip. Proboscis dark brown, about 3x the height of the frons.

Thorax. Scutum and scutellum orange brown with dense coppery setulae and some black setulae. Scutum with conspicuous whitish longitudinal stripes. Median stripe narrow and not very evident. Sublateral stripes wider than the median and quite evident. The sublateral and lateral stripes connected by a narrow white stripe at the transverse suture. The sublateral stripes connect at 2/3 of the posterior margin of the scutum and form a rectangular whitish patch. Notopleuron with tuft of black and coppery setulae. Anepisternum, katepisternum and anepimeron with dense and long white and coppery setulae. Postalar callus with tuft of yellowish white setulae. Veins yellowish-brown with faint brown spots on crossveins. Femora orange brown with long, dense black setulae. Tibiae and tarsi yellowish-brown with short dense black setulae, hind tibiae and tarsi darkened. Wing smoky brown with yellow patch in the middle of cells r 1 and r 2+3 and no lighter areas in the middle of cells m 1 and m 2. Abdomen. Tergites orange brown with black and coppery setulae. Central posterior margin of tergites 2 to 5 with triangles of setulae and white pruinosity. Lateral margin of tergites with white setulae. Sternites with coppery and white setulae.

Distribution. Costa Rica to Venezuela, Ecuador (Morona Santiago).

Type material examined. Syntype, ♀, deposited at the NMW (examined through photographs).

Additional material examined. 1 ♀, “ 14 km. N. Puerto, Quepos, San Jose Prov. Costa Rica, 1-14-63, R. Casebeer / LACM ENT 286749/ Scione maculipennis ” (CEUFT); 1 ♀, “ Costa Rica, Puntarenas, La Amistad Nat. Park, 8º57’N, 82º50’W, 23 Feb 1991, 1500 m, Norman D. Penny Cal. Acad. Sci. Coll. ” (CAS); 3 ♀ “ Costa Rica, Puntarenas, La Amistad Nat. Park, 8º57’N, 82º50’W, 1500 m, 18 February 1991, Norman D. Penny Cal. Acad. Sci. Coll. ” (CAS); 1 ♀, Volcan Cacao, 1km W. del Cerro Cacao, Prov. Guana, Costa Rica, 1400 m., 7 FEB 1995, F. Alvarado, L N 323700 #4453/ Costa Rica INBIO CRI002, 196112/ Scione maculipennis (Schiner) det. J. Burger 2000/ Scione maculipennis (Schiner) det. Krolow, T.K., 2012” (MNCR); 1 ♀, “ Panama, Chiriqui Prov., 2km NE. of Boquete, 24-IV-1976, D.C. Rentz, M.S. Carter and C.L. Mullinex / Scione maculipennis Schin, Det. 1981, G.B. Fairchild/ Scione maculipennis (Schiner) det. Krolow, T.K., 2012” (CAS); 1 ♀, “ Panama, Provincia de Chiriqui, S. slope of Volcan de Chiriqui, (Barú) 1000 m, 28.I.1987, Edward S. Ross / Scione maculipennis Schin., Det. G.B. Fairchild 19[?]” (CAS); 1 ♀, “ Boquete, Panama, Chiriqui Prov., Marc. 26. 1923, F.M. Gaige, 293/ Cornelius B. Philip Collection, Donated to California Academy Sciences/ Scione maculipennis Schin., det. C.B. Philip / eyes unbanded/ Compared with type ♀ Scione maculipennis Schin, C.B. Philip 53, agrees (over), some of cotyp have yellower beards [on the back of the label]/ Genotype Rhinotriclista End. (= Scione Wik.)/ Scione maculipennis (Schiner), det. Krolow, T.K., 2012” (CAS); 1 ♀, “ Rio Chanjena, Boca del Toro, 2400ff./ Panama, 1 sept. 1961 / Scione maculipennis Schin. det. G.B. Fairchild 1981/ Scione maculipennis ” (INPA); 1 ♀, “ Potrerillos, Panama, Elev. 2000, VI-14-35/ I.W. Mac Swain coll./ Cornelius B. Philip Collection, Donated to California Academy Sciences/ Scione maculipennis Macq. ♀ det. C.B. Philip 41” (CAS); 1 ♀, “ Potrerillos, Panama, Elev. 2000, VI-14-35/ I.W. Mac Swain coll./ T. Aitken Collectin/ Scione maculipennis Macq. ♀ det. C.B. Philip 41” (CAS); 1 ♀, “ Panama, Chiriqui Prov. 2km NE. of Boquete, 24-IV-1976, D.C. Rentz, M.S. Carter and C.L. Mullinex ” (CAS); 1 ♀, “ Panama, Chiriqui Prov.: 45Km N. of Chiriqui, Fortuna water shed, 20 June 1992, Jean & Keve Ribarbo ” (CAS); 1 ♀, “ Panama, Chiriqui, Alto Lino, 4000, 25 june 1965, H.G. Real / Herman G. Real Collection / Scione albifasciata ♀ (Macq) Det. C.B. Philip 72” (CAS); 1 ♀, “ Rio Pomeca, 2500—2600 m, Colombia, 18-21-VII-1948 / L. Richter coll., Frank Johnson Donor / Cornelius B. Philip Collection, Donated to California Academy of Sciences” (CAS); 1 ♀, “E. Loja, Ecuador, 2800 m, XI. 1970, L.E. Peña col./ MZSP” (MZUSP).

Comments. As mentioned in the original description, the boundaries of the wing spots are nowhere sharp, giving the wing surface a variegated, cloudy appearance that is readily visible to the naked eye. Triangles of setulae and white pruinosity extend across the entire abdominal tergites in some specimens. This species resembles S. albifasciata, from which it differs by the absence of patch on the frons, almost parallel frons and the pattern of spots on the wings. S. maculipennis has yellow spots scattered across more regions of the wing, whereas in S. albifasciata the yellow spots are restricted to cells r 1 and r 2+3. Wilkerson (1979) provided a redescription for S. maculipennis, the specimens examined here differs mainly in length and general body coloration. The specimens examined by Wilkerson measured between 9 and 11 mm, the specimens examined here measure between 11.2 and 13.4 mm, furthermore, according to the author, the species has a yellowish-brown body, but when examining the photographs of the types and the specimens on loan we saw that the general coloration of the body is orange brown. This species was redescribed because we examined a relatively large series of specimens and because S. maculipennis has many similar species (see comments on S. acris, S. claripennis, S. costaricana, and S. flavescens), thus an updated redescription may aid in distinguishing these species.