Published January 14, 2026
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TABLE 1 in Five new earthworm species of the Pheretima sangirensis-group (Clitellata: Megascolecidae) from Mount Banahaw, Quezon Province, Philippines
Authors/Creators
- 1. Department of Plant Medicine, College of Agriculture & Life Sciences, Jeonbuk National University, Jeonju 54896, Korea
- 2. Department of Regenerative Organic Agriculture, Maharishi International University Fairfield, Iowa, USA 52556
Description
TABLE 1. Comparison of species of the Philippine sangirensis group and P. sangirensis of Indonesia. Setal counts are given as a pair of numbers separated by a comma, the first of the pair being the count in segment vii, and the second being the number in xx. Setal gaps are represented as D for dorsal gap, either present (+D) or absent (–D), and likewise V for ventral. Reproductive pore spacings are given as the estimated fraction of body circumference. Typhlosole presence accompanied by the segment number of its origin if not in XXVII. Septal presence and absence are indicated by + or –, the first of each pair being for septum 8/9. For bursal pads, those found on the floor of the bursa are represented by a number followed by F (e.g. 4F), and those attached to the roof of the bursa are represented by a number followed by R (e.g. 4R). Missing data are shown by a question mark (?). Data from Aspe & James (2014; 2016; 2017), Hong & James (2008; 2011; 2021), – James (2004b), and Michaelsen (1896). *individuals with spermathecae; some are athecal.
| quincunxia | diesmosi | rugosa | monoporata | vicinipora | baungon-ensis | pauci-setosa | alba | rubida | virgata | sangirensis | apoensis | baracatan-ensis | camiguin-ensis | floresi | sibucal-ensis | solisi | timpoong-ensis | malindang-ensis | misamis-ensis | wati | nunezae | tigris | longi-prostata | maculo-dorsalis | immanis | lago | boniao | mariae | lagunaensis | vergrandis | concepcionensis | riparian-ensis | bontoc-ensis | banahaw-ensis sp. nov. | Pheretima sp 2 aurora | sancta sp. nov. | taceae sp. nov. | apolinarioi sp. nov. | madii sp. nov. | |
|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|
| setae vi, xxv | 18, 25 | 28, 38 | 24, 32 | 30, 34 | 52, 40 | 38, 50 | 22, 28 | 38, 36 | 34, 36 | 76, 80 | ? | 38–48, 50–58 | 28, 38 | 32, 42 | 37, 45 | 70, 90 | 52, 40 | 29–43, 50–72 | 44–48, 50–58 | 42–51, 43–48 | 60–70, 52–60 | 46, 51 | 38, 48–60 | 36–40, 38–48 | 74, 64–74 | 62–68, 64–68 | 49, 54 | 60–78, 62–68 | 32, 32 | 46, 52 | 37–42; 50–54 | 40–50; 56–68 | 42, 52 | 40, 62 | 46, 64 | ? | 28, 38 | 31, 36 | 39, 45 | 52, 56 |
| setal gaps D, V | Irreg. | +D, –V | +D, –V | +D, –V | +D, –V | +D, –V | +D, –V | +D, –V | +D, –V | –D, –V | +D,?V | +D,–V | +D, –V | +D, –V | +D, –V | +D, –V | +D, –V | +D, –V | –D, +V | +D, –V | +D, –V | +D, –V | +D, –V | +D, –V | +D, –V | –D, –V | +D, –V | +D, –V | +D, –V | +D, –V | +D, +V | +D, +V | +D, +V | +D, +V | +D, +V | -D, –V | –D, +V | +D, –V | +D, +V | +D, +V |
| male pore setae | 0–2 | 0 | 0 | 0 | 0 | 8 | 2 | 0 | 4 | 4–5 | 6 | 4–5 | 4–5 | 1–4 | 5 | 9 | 3–5 | 2–5 | 0 | 6–7 | 0 | 9 | 0 | 0–5 | 2–4 | 5 | 0–2 | 0 | 1 | 0 | – | 2 | 5–7 | 4–6 | 4–7 | 5 | 6–7 | 1 | 0 | 0 |
| male pore spacing | 0.13 | 0.15 | 0.15 | 0.05 | 0.04 | 0.19 | 0.13 | 0.10 | 0.12 | 0.09 | 0.17–0.25 | 0.15 | 0.22 | 0.17 | 0.20 | 0.17 | 0.15 | 0.23 | 0.11 | 0.23 | 0.08 | 0.22 | 0.14 | 0.16 | 0.13 | 0.14 | 0.15 | 0.03 | 0.09 | 0.10 | – | 0.14 | 0.11 | 0.10 | 0.09–0.13 | 0.23 | 0.16 | 0.13 | 0.12 | 0.04 |
| sp. pore spacing | 0.16 | 0.23 | 0.20 | 0 | 0.08 | 0.38 | 0.16 | 0.13 | 0.20 | 0.11 | 0.25–0.28 | 0.16* | 0.32 | 0.17 | 0.21 | 0.21 | 0.18 | 0.20 | 0.16 | 0.30 | 0.17 | 0.28 | 0.13 | 0.16 | 0.09 | 0.12 | 0.18–0.24 | 0.14 | 0.04 | 0.03 | – | – | 0.07 | 0.09 | 0.09–0.13 | 0.14 | 0.17–0.22 | 0.12 | 0 | 0.04 |
| dorsal pores | 12/13 | 12/13 | 11/12 | 12/13 | 13/14 | 13/14 | 11/12 | 13/14 | 11/12 | 12/13 | ? | 12/13 | 12/13 | 12/13 | 12/13 | 12/13 | 12/13 | 12/13 | 12/13 | 12/13 | 12/13 | 12/13 | 12/13 | 11/12 | 12/13 | 12/13 | 12/13 | ? | 12/13 | 12/13 | 13/14 | 12/13 | 12/13 | 12/13 | 4/5 | 11/12 | 12/13 | 5/6 | 11/12 | 12/13 |
| dorsal pigment | – | brown | brown | brown | brown | brown | – | – | red-brown | brown stripes | ? | violet brown | brown | red-brown | red- brown | gray brown | brown | violet brown | dark brown | black | brown | gray-brown | red - purple | yellowbrown | red | dark violet | dark brown | violet brown | dark violet | brown | light brown | dark brown | violet brown | brown | brown, | red | brown | brown | brown | brown |
| equatorial pigment | none | + | + | + | + | + | none | none | + | none | ? | none | + | none | + | none | + | none | none | none | + | none | none | none | none | none | none | none | + | + | present | present | + | none | none | none | + | none | + | none |
| length mm | 65–86 | 128 | 70 | 60–70 | 50 | 107 | 115 | 49 | 72 | 290–360 | ? | 160–350 | 100 | 63–122 | 48–67 | >165 | 68–86 | 145–225 | 60–80 | 55–65 | 67–75 | >116 | 230–280 | 37–41 | 226–235 | 365 | 233–315 | 101–133 | 55 | 40 | 35–75 | >89 | 75–105 | 68–83 | 130 | 90 | 65 | 80 | 33 | 54–60 |
| intestinal origin | ½xvi | xvii | xvii | xvii | xvi | xv | ½xvi | xvi | xvi | xvi | ? | xv | xv | xv | xv or xvi | xiv | xv | xv or xvi | xvi | xv | xv | xv | xvi | xv | xvii | xvi | xiv | xiv | xvi | xvi | xv | xvi | xvi | xvi | xvi | xvii | xvi | xvi | xvi | xvii |
| typhlosole | – | + | + | + | + | + | – | + | + | + | ? | + | + | + | + | + | + | + | + | + | + | + | + | + | + | + | + | + | + | + | + | + | + | + | + | + | + | + 33 | + 33 | + |
| esoph. lamellae | + | + | + | + | + | + | + | + | + | + | ? | + | + | + | + | + | + | + | + | + | + | + | + | + | + | + | + | + | ? | ? | – | – | ? | ? | + | ? | + | + | ? | ? |
| intestinal vessels | ? | ? | ? | 18–20 | 26 | 32 | 34 | 24 | 26 | 32–42 | ? | 32–38 | 28 | 26–30 | 35 | 38 | 29 | 31–44 | 30 | 32 | 34–38 | 20–23 | 56–58 | 20 | 50–54 | 28–32 | 36–38 | 33–45 | 34–36 | 28–30 | 24–28 | 30–35 | ? | 28–30 | 30 | ? | ? | ? | ? | ? |
| hearts | x–xiii | x– xiii | x–xiii | x–xiii | xi–xiii | x–xiii | x–xiii | x–xiii | x–xiii | x–xiii | ? | x–xiii | x– xiii | x–xiii | x–xiii | x–xiii | xi–xiii | x–xiii | xi–xiii | x–xiii | x–xiii | xi–xiii | x–xiii | x–xiii | x–xiii | x–xiii | x–xiii | xi–xiii | x–xiii | x–xiii | x–xiii | x–xiii | x–xiii | x–xiii | x–xiii | x–xiii | x– xiii | x–xiii | x–xiii | x–xiii |
| septa 8/9/10 | +, + | +, + | +, + | +, + | +, – | +, – | +, + | –, – | +, + | +, – | ? | +, – | +, – | +, – | +, – | +, – | +, – | –, – | –, – | +, + | –, + | +, – | +, – | –, – | +, – | +, – | +, – | +, – | –, – | –, – | –,– | –,– | –, – | –, – | –, – | –, – | –, – | –, – | –, – | +, + |
| bithecate vs mono | bi | bi | bi | mono | bi | bi | bi | bi | bi | bi | bi | bi | bi | bi | bi | bi | bi | bi | bi | bi | bi | bi | bi | bi | bi | bi | bi | bi | bi | bi | mono | mono | bi | bi | bi | bi | bi | bi | mono | bi |
| thecal segm. | vii | vii | vii | vii | viii | vii | vii | vii | vii | viii | ? | viii* | viii | viii | viii | vii | viii | viii | viii | vii | viii | vii | viii | vii | viii | viii | vii | viii | vii | vii | vii | viii | vii | vii | vii | viii | vii | vii | viii | vii |
| penes | – | – | + | + | – | – | + | + | + | + | + | – | – | + | – | – | – | + | + | + | + | + | – | + | + | – | + | – | + | + | + | + | + | + | + | + | + | + | + | + |
| bursal pads | 4F | 2F | 3F | 0 | 2R | 2R | 3F | 0 | 0 | 5F, 4R | ? | 1R | 3R | 1R | 2R | 1R | 2R 1F | 2R | 1R, 2F | 1F | 1R | 2R, 2F | 5F | 0 | 2R | 2F | 2R,1F | 1R | 0 | 0 | 0 | 0 | ring of wedges | 4F wedges | 5F, 1R | 2F oval pads | 4F, 1R | ring F | 5–6F wedges | 4–5F wedges |
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Notes
Published as part of Hong, Yong & James, Samuel W., 2026, Five new earthworm species of the Pheretima sangirensis-group (Clitellata: Megascolecidae) from Mount Banahaw, Quezon Province, Philippines, pp. 363-366 in Zootaxa 5748 (3) on page 363, DOI: 10.11646/zootaxa.5748.3.2, http://zenodo.org/record/19168509
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Identifiers
References
- Aspe, N. M. & James, S. W. (2014) New species of Pheretima (Oligochaeta: Megascolecidae) from the Mt. Malindang Range, Mindanao Island, Philippines. Zootaxa, 3881 (5), 401-439. https://doi.org/10.11646/zootaxa.3881.5.1
- Aspe, N. M. & James, S. W. (2016) New species of Pheretima, Amynthas, Polypheretima and Pithemera (Clitellata: Megascolecidae) from Mindanao and associated islands, Philippines. Zoological Studies, 55, 8-49.
- Aspe, N. M. & James, S. W. (2017) Pheretimoid earthworms (Clitellata: Megascolecidae) from Mt. Apo, Mindanao Island, Philippines with description of eight new species. Raffles Bulletin of Zoology, 65, 357-372.
- Hong, Y. & James, S. W. (2008) Three new earthworms of the genus Pheretima (Oligochaeta: Megascolecidae) from Mt. Makiling, Luzon Island, Philippines. Zootaxa, 1695 (1), 45-52. https://doi.org/10.11646/zootaxa.1695.1.2
- James, S. W. (2004 b) New species of Amynthas, Pheretima, Pleionogaster (Oligochaeta: Megascolecidae) of the Mt. Kitanglad Range, Mindanao Island, Philippines. Raffles Bulletin of Zoology, Singapore, 52 (2), 289-313.
- Michaelsen, W. (1896) Oligochaten Kukenthal - Ergebnisse einer zool. Forschungsreise in den Molukken und in Borneo. Abhandlungen der senckenbergische naturforschenden Gesellschaft, 23, 192-243.