Published February 3, 2026 | Version v1
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An exceptionally well-preserved starfish fauna (Asteroidea, Echinodermata) from the Early Miocene of southeastern France

  • 1. Université Claude Bernard Lyon 1, Villeurbanne, France
  • 2. Muséum d'Histoire Naturelle de Genève, Geneva, Switzerland|Université de Genève, Geneva, Switzerland|National Museum of Natural History, Department of Paleobiology, Washington, United States of America
  • 3. Muséum National d'Histoire Naturelle, Département Origine et Évolution, Paris, France
  • 4. Réserve Naturelle Nationale Géologique du Luberon, Parc Naturel Régional du Luberon, Apt, France

Description

A new starfish fauna from the Late Burdigalian (Early Miocene) of the Apt basin (Rhodano-Provençal gulf, southeastern France) is described here. This new starfish fauna comes from accumulation levels of marine invertebrates, including exceptionally well-preserved pectinid bivalves and echinoderms. Among the echinoderms, the sea urchins are the most abundant, closely followed by the starfish. This is the first time the starfish fauna is being studied in detail. Here are described twelve additional specimens of Lacosteaster lauerorum Gale & Ward, 2024 (Solasteridae, Valvatida) with the first description of the abactinal face of the species, an undetermined species of Astropecten (Astropectinidae, Paxillosida), and two new genera and species: Cruciformaster pedicellarius gen. nov. et sp. nov. (Oreasteridae, Valvatida) and Menerbesaster bongrainae gen. nov. et sp. nov. (Echinasteridae, Spinulosida).

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References

  • Agassiz L (1836) Prodrome d'une Monographie des Radiaires ou Echinodermes. Mémoires de la Société des Sciences naturelles de Neuchâtel 1: 168–199.
  • Agassiz L (1840) Description des Échinodermes fossiles de la Suisse. Partie 2: Cidarides. Mémoires de la Société helvétique de Sciences naturelles 4: 1–107.
  • Alvarado JJ (2011) Echinoderm diversity in the Caribbean Sea. Marine Biodiversity 41: 261–285. https://doi.org/10.1007/s12526-010-0053-0
  • Baeta M, Ramon M (2013) Feeding ecology of three species of Astropecten (Asteroidea) coexisting on shallow sandy bottoms of the northwestern Mediterranean Sea. Marine Biology 160: 2781–2795. https://doi.org/10.1007/s00227-013-2270-0
  • Besson D (2005) Architecture du bassin rhodano-provençal miocène (Alpes, SE France). Relations entre déformation, physiographie et sédimentation dans un bassin molassique d'avant-pays. PhD thesis, École des Mines, Paris, France.
  • Blainville HMD De (1830) Zoophytes. Dictionnaire de Sciences Naturelles 60: 214–223.
  • Blake DB (1979) The affinities and origins of the crown-of-thorns sea star Acanthaster Gervais. Journal of Natural History 13(3): 303–314. https://doi.org/10.1080/00222937900770241
  • Blake DB (1986) Some new post-Paleozoic sea stars (Asteroidea: Echinodermata) and comments on taxon endurance. Journal of Paleontology 60(5): 1103–1119. https://doi.org/10.1017/S0022336000022630
  • Blake DB (1987) A classification and phylogeny of post-Palaeozoic sea stars (Asteroidea: Echinodermata). Journal of Natural History 21(2): 481–528. https://doi.org/10.1080/00222938700771141
  • Blake DB (1990) Adaptative zones of the class Asteroidea (Echinodermata). Bulletin of Marine Science 46(3): 701–718.
  • Blake DB (1993) A new asteroid genus from the Jurassic of England and its functional significance. Paleontology 36(1): 147–154.
  • Blake DB, Aronson RB (1998) Eocene Stelleroids (Echinodermata) at Seymour Island, Antarctic peninsula. Journal of Paleontology 72(2): 339–353. https://doi.org/10.1017/S0022336000036325
  • Blake DB, Portell RW (2011) Kionaster petersonae, n. gen. and sp. (Asteroidea), the first fossil occurrence of the Asterodiscididae, from the Miocene of Florida. Swiss Journal of Palaeontology 130(1): 25–42. https://doi.org/10.1007/s13358-010-0005-0
  • Bongrain M (2013) Les accumulations de Gigantopecten restitutensis (Fontannes, 1884) (Mollusca: Bivalvia: Pectinidae) dans le Burdigalien supérieur des carrières de Ménerbes et de Lacoste (Vaucluse, bassin d'Apt, SE France) : analyse et hypothèse explicative. Geodiversitas 35(3): 607–628. https://doi.org/10.5252/g2013n3a5
  • Borghi E, Bajo Campos I (2009) Asteroidei fossili dell'Emilia. Notiziario della Società Reggiana di Scienze Naturali 2008: 13–29.
  • Breton G (1992) Les Goniasteridae (Asteroidea, Echinodermata) jurassiques et crétacés de France : Taphonomie, systématique, biostratigraphie, paléobiogéographie, évolution. PhD thesis, University of Caen, Caen, France.
  • Breton G, Bourseau JP, Bernier P, Barale G, Buffetaut E, Gaillard C, Gall JC, Wenz S (1994) Les astérides (Asteroidea, Echinodermata) des calcaires lithographiques kimméridgiens de Cerin (Ain, France). Geobios 16: 49–60. https://doi.org/10.1016/S0016-6995(94)80020-0
  • Carbonnel G (1969) Les ostracodes du Miocène rhodanien : Systématique, biostratigraphie écologique, paléobiologie. Documents des Laboratoires de Géologie de la Faculté des Sciences de Lyon 32(2): 229–469.
  • Cavara F (1886) Le sabbie marnose plioceniche di Mongardino e i loro fossili. Bollettino della Societa Geologica Italiana 5: 265–276.
  • Chao SM (1999) A Revision of the Family Astropectinidae (Echinodermata: Asteroidea) from Taiwan, with Description of Five New Records. Zoological studies 38(3): 257–267.
  • Clark AM (1993) An index of names of recent Asteroidea. Part 2: Valvatida. Echinoderm Studies 4: 187–366. https://doi.org/10.1201/9781003072553-4
  • Clark AM, Downey ME (1992) Starfishes of the Atlantic. London: Chapman & Hall, 794 pp.
  • Clark AM, McGown A (1962) Asteroidea. BANZ Antarctic Research Expedition 1929–1931 9(1): 1–104.
  • Clark AM, Rowe FWE (1971) Monograph of shallow-water Indo-West Pacific echinoderms. British Museum (Nat. Hist.), London, United Kingdom, 238 pp.
  • Clark HL (1909) Notes on some Australian and Indo-Pacific echinoderms. Bulletin of the Museum of Comparative Zoology, Harvard 52: 107–135.
  • Clark HL (1938) Echinoderms from Australia. Memoirs of the Museum of Comparative Zoology at Harvard College 55: 1–596.
  • Clark RN, Jewett SC (2010) A new genus and thirteen new species of sea stars (Asteroidea: Echinasteridae) from the Aleutian Island Archipelago. Zootaxa 2571: 1–36. https://doi.org/10.11646/zootaxa.2571.1.1
  • Costello MJ, Emblow CS, White R (2001) European Register of Marine Species: a check-list of the marine species in Europe and a bibliography of guides to their identification. Collections Patrimoines Naturels 50: 463.
  • Del Prato A (1896) Asteroidei terziari del Parmense e del Reggiano. Rivista Italiana di Paleontologia 2: 42–50.
  • Delle Chiaje S (1827) Memorie sulla storia e notomia degli animali senza vertebre del regno di Napoli. Volume II. Stamp. Societa Tipografica, Napoli, Italia, 232 pp.
  • Demarcq G (1962) Étude stratigraphique du Miocène rhodanien. PhD thesis, University of Paris, Paris, France.
  • Demarcq G (1984) Importance des mégafaunes marines benthiques dans l'évolution paléothermique de la Méditerranée au Néogène. Annales de Géologie des Pays helléniques 32: 87–95.
  • Döderlein L (1917) Die Asteriden der Siboga-Expedition. II. 1. Die Gattung Astropecten und ihre Stammesgeschichte. Siboga-Expeditie. Uitkomsten op zoölogisch, botanisch, ozeanographisch en geologisch gebied verzameld in Nederlandsch Oost-Indie 1899–1900 aan boord H. M. "Siboga" 46a: 1–191 [pls 1–17]. https://doi.org/10.5962/bhl.title.11319
  • Döderlein L (1936) Die Asteriden der Siboga-Expedition. III. Die Unterfamilie Oreasterinae. Siboga-Expeditie. Uitkomsten op zoölogisch, botanisch, ozeanographisch en geologisch gebied verzameld in Neder- landsch Oost-Indie 1899–1900 aan boord H. M. "Valdivia" 46c: 295–369 [pls 21–32].
  • Downey ME (1973) Starfishes from the Caribbean and the Gulf of Mexico. Smithsonian Contribution to Zoology 126: 1–158. https://doi.org/10.5479/si.00810282.126
  • Durham JW, Roberts WA (1948) Cretaceous Asteroids from California. Journal of Paleontology 22(4): 432–439.
  • Éléaume M, Roux M, Philippe M (2020) Discometra luberonensis sp. nov. (Crinoidea, Himerometridae), a new feather star from the Late Burdigalian. European Journal of Taxonomy 729(1): 121–137. https://doi.org/10.5852/ejt.2020.729.1193
  • Fau M, Villier L (2018) Post-metamorphic ontogeny of Zoroaster fulgens Thomson, 1873 (Asteroidea, Forcipulatacea). Journal of Anatomy 233(5): 557–686. https://doi.org/10.1111/joa.12881
  • Fau M, Villier L (2020) Comparative anatomy and phylogeny of the Forcipulatacea (Echinodermata: Asteroidea): insights from ossicle morphology. Zoological Journal of the Linnean Society 189(3): 921–952. https://doi.org/10.1093/zoolinnean/zlz127
  • Fisher WK (1911) Asteroidea of the North Pacific and adjacent waters. 1. Phanerozonia and Spinulosida. Bulletin of the United States National Museum 76: [i–xiii +] 1–420. https://doi.org/10.5479/si.03629236.76.i
  • Fisher WK (1908) Necessary changes in the nomenclature of starfishes. Smithsonian Miscellaneous collections (Quaterly) 52: 87–93.
  • Fisher WK (1919) Starfishes of the Philippine seas and adjacent waters. Bulletin of the United States National Museum 3(100): 1–547.
  • Fisher WK (1940) Asteroidea. Discovery Reports 20: 69–306.
  • Fontannes F (1884) Sur une des causes de la variation dans le temps des faunes malacologiques, à propos de la filiation des Pecten restitutensis et latissimus. Bulletin de la Société Géologique de France 12(3): 357–364.
  • Forbes E (1839) On the Asteriadae of the Irish Sea. Memoirs of the Wernerian Natural History Society of Edinburgh 8: 114–129.
  • Forbes E (1856) Solaster moretonis. Memoirs of the Geological Survey of Great Britain, British Organic Remains 5: 1–3.
  • Gale AS (1987a) Phylogeny and classification of the Asteroidea (Echinodermata). Zoological Journal of the Linnean Society 89: 107–132. https://doi.org/10.1111/j.1096-3642.1987.tb00652.x
  • Gale AS (1987b) Goniasteridae (Asteroidea, Echinodermata) from the Late Cretaceous of North-West Europe. 1. Introduction. The genera Metopaster and Recurvaster. Mesozoic Research 1: 1–69.
  • Gale AS (1989) Migration and evolution in Late Cretaceous Goniasteridae (Asteroidea, Echinodermata) from north-west Europe. Proceedings of the Geologists' Association 100(3): 281–291. https://doi.org/10.1016/S0016-7878(89)80048-3
  • Gale AS (2011a) The phylogeny of post-Palaeozoic Asteroidea (Neoasteroidea, Echinodermata). Special Papers in Paleontology 85: 1–112.
  • Gale AS (2011b) Asteroidea (Echinodermata) from the Oxfordian (Late Jurassic) of Savigna, Département du Jura, France. Swiss Journal of Paleontology 130(1): 69–89. https://doi.org/10.1007/s13358-010-0008-x
  • Gale AS, Jagt JWM (2021) The fossil record of the family Benthopectinidae (Echinodermata, Asteroidea), a reappraisal. European Journal of Taxonomy 755: 149–190. https://doi.org/10.5852/ejt.2021.755.1405
  • Gale AS, Ward DJ (2024) A new sun star (Echinodermata, Asteroidea, Solasteridae) from the mid-Miocene of Lacoste, France. Proceedings of the Geologists' Association 135(6): 685–694. https://doi.org/10.1016/j.pgeola.2024.10.001
  • Germain C, Liouvillen M, De Bouchony P, Roch E, Demarcq G (1966) Carte géologique au 1/50000e, n°967, feuille de Cavaillon. Bureau des Recherches Géologiques et Minières, Orléans.
  • Gray JE (1840) XXXII. A synopsis of the genera and species of the class Hypostoma (Asterias, Linnaeus). Annals of the Magazine of Natural History 6: 275–290. https://doi.org/10.1080/03745484009443296
  • Hayashi R (1940) Contributions to the Classification of the Sea-stars of Japan. Records of the Faculty of Science of Hokkaido Imperial University 7(3): 107–204.
  • Heller C (1858) Uber neue fossile Stelleriden. Sitzungsberichte der Kaiserlichen Akademie der Wissenschaften, Mathematisch-Naturwissenschaftliche Classe 28: 155–170.
  • Hess HW (1955) Die fossilen Astropectiniden (Asteroidea). Schweizerische Paläeontologische Abhandlung 71: 1–113.
  • Hess H (1966) Mikropaläontologische Untersuchungen an Ophiuren, V: Die Ophiuren aus dem Argovien (unteres Ober-Oxford) vom Guldenthal (Kt Solothurn) und von Savigna (Dépt. Jura). Eclogae Geologicae Helvetiae 59: 1025–1063.
  • Hess HW (1972) Lower Tertiary Crinoidea, Asteroidea and Ophiuroidea from Northern Europe and Greenland. Biologiske skrifter fra det Danske Videnskabernes Selskab 19: 1–83.
  • Jagt JW, Codrea V (2010) A goniasterid starfish (Echinodermata, Asteroidea) preserved in a mid-Miocene rhyolitic ignimbrite, northwest Romania. Acta Geologica Polonica 60(2): 275–282.
  • Jaselli L, Collareta A (2018) Redescription and first illustration of the holotype of Astropecten montalionis (Meneghini, 1852) [Paxillosida: Astropectinidae]. Atti della Societa Toscana di Scienze Naturali 125: 35–43. https://doi.org/10.2424/ASTSN.M.2018.21
  • Kano YT, Komatsu, M, Oguro, C (1991) Morphological changes of the cushion star, Culcita novaeguineae Müller et Troschel, during growth. In: Yanagisawa T, Yasumasu I, Oguro C, Suzuki N, Motokawa T (Eds) Biology of Echinodermata. AA Balkema, Rotterdam, The Netherland, 323–325. https://doi.org/10.1201/9781003077565-74
  • Kato M, Oji T (2013) A new species of Doraster (Echinodermata: Asteroidea) from the lower Miocene of central Japan: implications for its enigmatic paleobiogeography. Paleontological Research 17(4): 330–334. https://doi.org/10.2517/1342-8144-17.4.330
  • Koehler R (1909) Résultats des campagnes scientifiques du prince de Monaco. Fascicule XXXIV: Échinodermes provenant des campagnes du yatch PRINCESSE-ALICE (Astéries, Ophiures, Échinides et Crinoïdes). Imprimeries de Monaco, Monaco.
  • Koehler R (1920) Echinodermata: Asteroidea. Scientific Reports of the Australasian Antarctic Expedition 8: 1–308. https://doi.org/10.5962/bhl.title.85380
  • Koehler R (1921) Faune de France. Vol. 1: Échinodermes. Kraus Reprint, Nendeln, Liechtenstein. https://doi.org/10.5962/bhl.title.11514
  • Lamarck JBM (1816) Histoire naturelle des animaux sans vertèbres, Tome troisième. Imprimerie d'Abel Lanoe, Paris, France.
  • Koukouras A, Sinis AI, Bobori D, Kazantzidis S, Kitsos MS (2007) The echinoderm (Deuterostomia) fauna of the Aegean Sea, and comparison with those of the neighbouring seas. Journal of Biological Research 7: 67–92.
  • Lawrence JM, Agatsuma Y (2007) Ecology of Tripneustes. Developments in Aquaculture and Fisheries Science 37: 499–520. https://doi.org/10.1016/S0167-9309(07)80088-3
  • Linnaeus C (1758) Systema Naturae per regna tria naturae, secundum classes, ordines, genera, species, cum characteribus, differentiis, synonymis, locis. Laurentius Salvius, Stockholm, Sweden, 1–824. https://doi.org/10.5962/bhl.title.542
  • Mah CL (2018) New genera, species and occurrence records of Goniasteridae (Asteroidea; Echinodermata) from the Indian Ocean. Zootaxa 4539(1): 1–116. https://doi.org/10.11646/zootaxa.4539.1.1
  • Mah CL (2023) A new species of Astrosarkus from Western Australia including new Mesophotic occurrences of Indian Ocean Oreasteridae (Valvatida, Asteroidea). Memoirs of Museum Victoria 82: 143–165. https://doi.org/10.24199/j.mmv.2023.82.08
  • Mah CL (2025) World Asteroidea Database. Astropecten Gray, 1840. [Accessed through: World Register of Marine Species] https://www.marinespecies.org/aphia.php?p=taxdetails&id=123245
  • Mah CL, Blake DB (2012) Global Diversity and Phylogeny of the Asteroidea (Echinodermata). PLoS ONE 7(4): e35644. https://doi.org/10.1371/journal.pone.0035644
  • Mah CL, Fujita T (2020) New species and occurrence records of Japanese Solasteridae and Ganeriidae including a new species of Paralophaster from the North Pacific with an overview of Hyalinothrix. Zootaxa 4750(1): 67–100. https://doi.org/10.11646/zootaxa.4750.1.4
  • Mecho A, Billett DS, Ramírez-Llodra E, Aguzzi J, Tyler PA (2014) First records, rediscovery and compilation of deep-sea echinoderms in the middle and lower continental slope of the Mediterranean Sea. Scientia Marina 78(2): 281–302. https://doi.org/10.3989/scimar.03983.30C
  • Meneghini G (1852) Nuovi fossili Toscani illustrati. Estratto dagli Annali dell'Università Toscana 3.
  • Meneghini G (1886) Goniodiscus ferrazii MGH. Nuova stelleride terziaria del Vicentino. Atti della Società Toscana di Scienze Naturali 8(2): 1–8.
  • Müller J, Troschel FH (1840) Ueber die Gattungen der Asterien. Archiv für Naturgeschichte 6: 318–326.
  • Müller J, Troschel FH (1842) System der Asteriden. 1. Asteriae. 2. Ophiuridae. Vieweg, Braunschweig, Germany, [xxx+]134 pp. [12 pls.] https://doi.org/10.5962/bhl.title.11715
  • Nosowska E (1997) Asteroids from the Nawodzice Sands (Middle Miocene; Holy Cross Mountains, Central Poland). Acta Geologica Polonica 47(3–4): 225–241.
  • Otto AW (1823) Beschreibung einiger neueu Mollusken und Zoophyten. Nova Acta Physico-Medica Academiae Caesareae Leopoldino-Carolinae Naturae Curiosum 11(2): 273–314. https://doi.org/10.5962/bhl.title.118700
  • Parenzan P (1934) Revisione delle specie mediterranee del genere Astropecten. Italian Journal of Zoology 5(1): 201–223. https://doi.org/10.1080/11250003409426274
  • Pennant T (1777) British Zoology. Vol. IV. Crustacea. Mollusca. Testacea. William Eyres, London, United Kingdom, 374 pp.
  • Perrier E (1884) Mémoire sur les étoiles de mer recueillies dans la mer des Antilles et le golfe du Mexique : durant les expéditions de dragage faites sous la direction de M. Alexandre Agassiz. Archives (Muséum national d'histoire naturelle (France)) 6(2): 127–276. https://doi.org/10.5962/bhl.title.82184
  • Philippe M (1998) Les échinides miocènes du bassin du Rhône: Révision systématique. PhD thesis, University of Lyon, Villeurbanne, France, 3–241. https://doi.org/10.3406/mhnly.1998.1154
  • Philippi RA (1837) Ueber die mit Asterias auranciaca verwandten und verweschselton Asterien der Sicilianschen Kuste. Archiv für Naturgeschichte 3: 193–194.
  • Pillon R (2022) Identification key of the starfish Astropecten of the Mediterranean Sea. Les cahiers de la fondation Biotope 38: 1–33.
  • Rasmussen HW (1972) Lower Tertiary Crinoidea, Asteroidea and Ophiuroidea from Northern Europe and Greenland. Biologiske skrifter fra det Danske Videnskabernes Selskab 19: 1–83.
  • Rowe FWE, Gates J (1995) Echinodermata. Zoological Catalogue of Australia 33: 1–510. https://doi.org/10.1086/419606
  • Rowe FEW, Albertson EL (1988) A new genus and four new species in the Family Echinasteridae (Echinodermata: Asteroidea). Proceedings of The Linnean Society of New South Wales 110(1): 83–100.
  • Sacco F (1893) Sopra alcuni Asteroidei fossili. Atti della Reale Accademia delle Scienze di Torino 28: 739–745.
  • Scheibling RE (2012) Oreaster reticulatus. In Lawrence JM (Ed.) Biology and Ecology of the Asteroidea, Part II: Integrative Biology. The Johns Hopkins University Press, Baltimore, United States of America, 142–153.
  • Sladen WP (1891) A Monograph on the British Fossil Echinodermata from the Cretaceous Formations. Volume Second. The Asteroidea. Part First. Monographs of the Palaeontographical Society, 1–28. https://doi.org/10.1080/02693445.1891.12028012
  • Sladen WP (1893) A Monograph on the British Fossil Echinodermata from the Cretaceous Formations. Volume Second. The Asteroidea. Part Second. Monographs of the Palaeontographical Society, 29–66. https://doi.org/10.1080/02693445.1893.12028026
  • Spencer WK (1905) A Monograph on the British Fossil Echinodermata from the Cretaceous Formations. Volume Second. The Asteroidea. Part Third. Monographs of the Palaeontographical Society, 67–90. https://doi.org/10.1080/02693445.1905.12035516
  • Spencer WK (1907) A Monograph on the British Fossil Echinodermata from the Cretaceous Formations. Volume Second. The Asteroidea and Ophiuroidea. Part Fourth. Monographs of the Palaeontographical Society, 91–132. https://doi.org/10.1080/02693445.1907.12035532
  • Spencer WK (1908) A Monograph on the British Fossil Echinodermata from the Cretaceous Formations. Volume Second. The Asteroidea and Ophiuroidea. Part Fifth. Monographs of the Palaeontographical Society of London, 133–138. https://doi.org/10.1080/02693445.1908.12035541
  • Spencer WK, Wright CK (1966) Asterozoans. Part U: Echinodermata. Treatise on Invertebrate Paleontology 3(1): 4–107.
  • Valette A (1924) Note sur les débris de Stellérides fossiles du sud-ouest de la France. Actes de la Société linnéenne de Bordeaux 76: 167–205.
  • Verrill AE (1915) Report on the starfishes of the West Indies, Florida, and Brazil, including those obtained by the Bahama expedition from the University of Iowa in 1893. Bulletins form the Laboratories of Natural History 7(1): 1–232. https://doi.org/10.5962/bhl.title.12035
  • Verrill AE (1867) Notes on Radiata in the Museum of Yale College, with descriptions of new genera and species. No. 3. On the geographical distribution of the echinoderms of the West coast of America. Transactions of the Connecticut Academy of Arts and Sciences 1(2): 323–351.
  • Viguier C (1878) Classification des Stellérides. Comptes rendus hebdomadaires des séances de l'Académie des sciences 86: 681–683.
  • Villier L, Charbonnier S, Riou B (2009) Sea stars from Middle Jurassic Lagerstätte of La Voulte-sur-Rhône (Ardèche, France). Journal of Paleontology 83(3): 389–398. https://doi.org/10.1666/08-030.1
  • Yiu SKF, Mah CL (2024) New Ecological Observations and Occurrence for Asteroidea and Echinoidea in Hong Kong. Zootaxa 5526(1): 1–69. https://doi.org/10.11646/zootaxa.5526.1.1
  • Zulliger DE (2009) Phylogeography, Evolutionary History and Genetic Diversity of Sea Stars of the Genus Astropecten and Genetic structure within the Atlanto-Mediterranean Species A. aranciacus. PhD thesis, University of Zürich, Zurich, Switzerland.
  • Zulliger DE, Lessios HA (2010) Phylogenetic relationships in the genus Astropecten Gray (Paxillosida: Astropectinidae) on a global scale: molecular evidence for morphological convergence, species-complexes and possible cryptic speciation. Zootaxa 2504: 1–19. https://doi.org/10.11646/zootaxa.2504.1.1