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Published February 19, 2026 | Version v1
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A new kentriodontid dolphin (Cetacea, Odontoceti, Delphinida) from the Middle Miocene of the southeastern Pacific

Authors/Creators

  • 1. D.O. Terre et Histoire de la Vie, Institut royal des Sciences naturelles de Belgique, 29 rue Vautier, B-1000 Brussels (Belgium) olambert@naturalsciences.be (corresponding author)
  • 2. Université de Strasbourg, CNRS, IPHC UMR 7178, F-67000 Strasbourg (France) and Centre d'Études biologiques de Chizé, CNRS, UMR 7372, F-79360 Villiers en Bois (France) maelle.oberlin.pro@gmail.com
  • 3. CR2P (CNRS, MNHN, Sorbonne Université), Département Origines et Évolution, Muséum national d'Histoire naturelle, case postale 38, 57 rue Cuvier, F-75231 Paris cedex 05 (France) muizon@mnhn.fr
  • 4. Dipartimento di Scienze della Terra, Università di Pisa, via S. Maria 53, I-56126 Pisa (Italy) alberto.collareta@unipi.it
  • 5. Departamento de Paleontología de Vertebrados, Museo de Historia Natural-Universidad Nacional Mayor de San Marcos, Avenida Arenales 1256, P-15072 Lima (Peru) mariourbina01@hotmail.com
  • 6. Dipartimento di Scienze della Terra, Università di Pisa, via S. Maria 53, I-56126 Pisa (Italy) giovanni.bianucci@unipi.it

Description

Lambert, Olivier, Oberlin, Maëlle, Muizon, Christian de, Collareta, Alberto, Urbina, Mario, Bianucci, Giovanni (2026): A new kentriodontid dolphin (Cetacea, Odontoceti, Delphinida) from the Middle Miocene of the southeastern Pacific. Geodiversitas 48 (3): 25-50, DOI: 10.5252/geodiversitas2026v48a3

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URN
urn:lsid:zoobank.org:pub:urn:lsid:zoobank.org:pub:6DFFE442-7DE6-42CF-B184-5B94B354FB36
LSID
urn:lsid:plazi.org:pub:E673DD781C2EFD46D35BFFB12466FF91
URL
http://publication.plazi.org/id/E673DD781C2EFD46D35BFFB12466FF91

Cites
Publication: 10.1671/0272-4634(2001)021[0573:ANGOKC]2.0.CO;2 (DOI)
Publication: 10.4435/BSPI.2022.04 (DOI)
Publication: 10.1016/j.palaeo.2018.08.013 (DOI)
Publication: 10.1016/j.cub.2022.02.041 (DOI)
Publication: 10.1127/nos/2019/0525 (DOI)
Publication: 10.1016/j.jsames.2019.102399 (DOI)
Publication: 10.1016/j.jsames.2022.103909 (DOI)
Publication: 10.1080/17445647.2020.1850535 (DOI)
Publication: 10.3390/jmse9111188 (DOI)
Publication: 10.1127/njgpa/2023/1128 (DOI)
Publication: 10.1080/02724634.1996.10011290 (DOI)
Publication: 10.13130/2039-4942/8373 (DOI)
Publication: 10.1186/1471-2148-11-112 (DOI)
Publication: 10.5479/si.1943-6688.100 (DOI)
Publication: 10.7717/peerj.10945 (DOI)
Publication: 10.1371/journal.pone.0280218 (DOI)
Publication: 10.1111/j.1440-1738.1994.tb00126.x (DOI)
Publication: 10.1002/ar.24585 (DOI)
Publication: 10.1111/j.1440-1738.1994.tb00127.x (DOI)
Publication: 10.1002/mmng.200410004 (DOI)
Publication: 10.1671/0272-4634(2005)025[0929:ROSMKC]2.0.CO;2 (DOI)
Publication: 10.1080/02724634.2014.857347 (DOI)
Publication: 10.1080/02724634.2019.1566739 (DOI)
Publication: 10.2517/PR220027 (DOI)
Publication: 10.1111/zoj.12479 (DOI)
Publication: 10.1080/14772019.2017.1359689 (DOI)
Publication: 10.1002/spp2.1332 (DOI)
Publication: 10.1127/nos/2025/0864 (DOI)
Publication: 10.1098/rsos.170560 (DOI)
Publication: 10.1017/pab.2018.33 (DOI)
Publication: 10.1016/j.ympev.2009.08.018 (DOI)
Publication: 10.1186/1471-2148-11-98 (DOI)
Publication: 10.5479/si.00810282.627 (DOI)
Publication: 10.1080/02724634.2013.816720 (DOI)
Publication: 10.3390/biology13020114 (DOI)
Publication: 10.1007/s00531-021-02003-1 (DOI)
Publication: 10.18563/journal.m3.91 (DOI)
Publication: 10.1080/02724634.2017.1411357 (DOI)
Publication: 10.1098/rstb.2022.0086 (DOI)
Has part
Taxonomic treatment: 10.5281/zenodo.18700097 (DOI)
Taxonomic treatment: http://treatment.plazi.org/id/1A4AA5001C2BFD43D3B7FF4626ACFC20 (URL)
Taxonomic treatment: 10.5281/zenodo.18693546 (DOI)
Taxonomic treatment: http://treatment.plazi.org/id/1A4AA5001C2BFD43D27EFC47259DFACA (URL)
Taxonomic treatment: 10.5281/zenodo.18693552 (DOI)
Taxonomic treatment: http://treatment.plazi.org/id/1A4AA5001C2BFD51D3ADFA2725AFF97D (URL)
Figure: 10.5281/zenodo.18700564 (DOI)
Figure: 10.5281/zenodo.18700566 (DOI)
Dataset: 10.5281/zenodo.18748494 (DOI)
Figure: 10.5281/zenodo.18700568 (DOI)
Figure: 10.5281/zenodo.18700570 (DOI)
Figure: 10.5281/zenodo.18700572 (DOI)
Figure: 10.5281/zenodo.18700574 (DOI)
Figure: 10.5281/zenodo.18700576 (DOI)
Figure: 10.5281/zenodo.18700578 (DOI)
Figure: 10.5281/zenodo.18700582 (DOI)
Figure: 10.5281/zenodo.18700584 (DOI)
Figure: 10.5281/zenodo.18700586 (DOI)
Figure: 10.5281/zenodo.18700588 (DOI)
Figure: 10.5281/zenodo.18700590 (DOI)
Figure: 10.5281/zenodo.18700592 (DOI)
Dataset: 10.5281/zenodo.18748498 (DOI)
Figure: 10.5281/zenodo.18700594 (DOI)
Is source of
Dataset: https://www.gbif.org/dataset/14836d91-e4fd-4c65-926a-159ed6a4efb5 (URL)

References

  • BARNES L. G. 1978. - A review of Lophocetus and Liolithax and their relationships to the delphinoid family Kentriodontidae (Cetacea: Odontoceti). Contributions in Science, Natural History Museum of Los Angeles County 28: 1-35.
  • BARNES L. G. 1985. - The Late Miocene dolphin Pithanodelphis Abel, 1905 (Cetacea, Kentriodontidae) from California.Los Angeles County Museum Contributions in Science 367: 1-27.
  • BARNES L. G. & MITCHELL E. D. 1984. - Kentriodon obscurus (Kellogg, 1931), a fossil dolphin (Mammalia: Kentriodontidae) from the Miocene Sharktooth Hill Bonebed in California.Contributions in Science, Natural History Museum of Los Angeles County 353: 1-23.
  • BIANUCCI G. 2001. - A new genus of kentriodontid (Cetacea: Odontoceti) from the Miocene of south Italy. Journal of Vertebrate Paleontology 21 (3): 573-577.https://doi.org/10.1671/0272- 4634(2001)021[0573:ANGOKC]2.0.CO;2
  • BIANUCCI G. 2005. - Arimidelphis sorbinii a new small killer whale-like dolphin from the Pliocene of Marecchia River (Central Eastern Italy) and a phylogenetic analysis of the Orcininae (Cetacea: Odontoceti). Rivista Italiana di Paleontologia e Stratigrafia 111 (2): 329-344.
  • BIANUCCI G. & COLLARETA A. 2022. - An overview of the fossil record of cetaceans from the East Pisco Basin (Peru).Bollettino della Societa PaleontoIogica Italiana 61: 19-60. https://doi.org/10.4435/ BSPI.2022.04
  • BIANUCCI G., COLLARETA A., BOSIO G., LANDINI W., GARIBOLDI K., GIONCADA A., LAMBERT O., MALINVERNO E., MUIZON C. DE, VARAS-MALCA R., VILLA I. M., COLETTI G., URBINA M. & DI CELMA C. 2018. - Taphonomy and palaeoecology of the lower Miocene marine vertebrate assemblage of Ullujaya (Chilcatay Formation, East Pisco Basin, southern Peru). Palaeogeography, Palaeoclimatology, Palaeoecology 511: 256-279.https://doi.org/10.1016/j. palaeo.2018.08.013
  • BIANUCCI G., GEISLER J. H., CITRON S. & COLLARETA A. 2022. - The origins of the killer whale ecomorph. Current Biology 32 (8): 1843-1851. https://doi.org/10.1016/j.cub.2022.02.041
  • BOSIO G., MALINVERNO E., VILLA I. M., DI CELMA C., GARIBOLDI K., GIONCADA A., BARBERINI V., URBINA M. & BIANUCCI G. 2020a.-Tephrochronology and chronostratigraphy of the Miocene Chilcatay and Pisco formations (East Pisco Basin, Peru). Newsletters on Stratigraphy 53:213-247.https://doi.org/10.1127/nos/2019/0525
  • BOSIO G., MALINVERNO E., COLLARETA A., DI CELMA C., GIONCADA A., PARENTE M., BERRA F., MARX F. G., VERTINO A., URBINA M. & BIANUCCI G. 2020b. - Strontium Isotope Stratigraphy and the thermophilic fossil fauna from the middle Miocene of the East Pisco Basin (Peru). Journal of South American Earth Sciences 97: 102399. https://doi.org/10.1016/j.jsames.2019.102399
  • BOSIO G., BIANUCCI G., COLLARETA A., LANDINI W., URBINA M. & DI CELMA C. 2022. - Ultrastructure, composition, and 87Sr/86Sr dating of shark teeth from lower Miocene sediments of southwestern Peru. Journal of South American Earth Sciences 118: 103909.https:// doi.org/10.1016/j.jsames.2022.103909
  • COLBERT E. H. 1944. - A new fossil whale from the Miocene of Peru. Bulletin of the American Museum of Natural History 83: 195-216. http://hdl.handle.net/2246/389
  • COLLARETA A., DI CELMA C., BOSIO G., PIERANTONI P. P., MALINVERNO E., LAMBERT O., MARX F. G., LANDINI W., URBINA M. & BIANUCCI G. 2021a. - Distribution and paleoenvironmental framework of middle Miocene marine vertebrates along the western side of the lower Ica Valley (East Pisco Basin, Peru). Journal of Maps 17 (2): 7-17. https://doi.org/10.1080/17445647.2020.1850535
  • COLLARETA A., LAMBERT O., MARX F. G., MUIZON C. DE, VARAS-MALCA R., LANDINI W., BOSIO G., MALINVERNO E., GARIBOLDI K., GIONCADA A., URBINA M. & BIANUCCI G. 2021b. - Vertebrate palaeoecology of the Pisco Formation (Miocene, Peru): glimpses into the ancient Humboldt Current ecosystem. Journal of Marine Science and Engineering 9 (11): 1188. https://doi.org/10.3390/ jmse9111188
  • COLLARETA A., BOSIO G., VARAS-MALCA R., BIANUCCI G., MERELLA M., URBINA M. & DI CELMA C. 2023. - The extinct nautiloid Aturia in the Middle Miocene of Pacific South America: new data from the Pisco Lagerstatte of Peru. Neues Jahrbuch fur Geologie und Palaontologie, Abhandlungen 308 (1): 23-32. https://doi. org/10.1127/njgpa/2023/1128
  • COMMITTEE ON TAXONOMY 2024. - List of marine mammal species and subspecies. Society for Marine Mammalogy, https://www. marinemammalscience.org, consulted on October 18, 2024.
  • DAWSON S. D. 1996. - A description of the skull and postcrania of Hadrodelphis calvertense Kellogg 1966, and its position within the Kentriodontidae (Cetacea; Delphinoidea). Journal of Vertebrate Paleontology 16 (1): 125-134. https://doi.org/10.1 080/02724634.1996.10011290
  • DEVRIES T. J., GROVES L. T. & URBINA M. 2006. - A new early Miocene Muracypraea Woodring, 1957 (Gastropoda: Cypraeidae) from the Pisco Basin of southern Peru. Nautilus 120 (3): 101-105. https://www.biodiversitylibrary.org/page/35185422
  • DI CELMA C., MALINVERNO E., BOSIO G., COLLARETA A., GARIBOLDI K., GIONCADA A., MOLLI G., BASSO D., VARAS-MALCA R. M., PIERANTONI P. P., VILLA I. M., LAMBERT O., LANDINI W., SARTI G., CANTALAMESSA G., URBINA M. & BIANUCCI G. 2017. - Sequence stratigraphy and paleontology of the upper Miocene Pisco Formation along the western side of the lower Ica Valley (Ica desert, Peru) Rivista Italiana di Paleontologia e Stratigrafia 123 (2): 255-273. https://doi.org/10.13130/2039- 4942/8373
  • FRASER F. C. & PURVES P. E. 1960. - Hearing in cetaceans: Evolution of the accessory air sacs and the structure of the outer and middle ear in recent cetaceans. Bulletin of the British Museum (Natural History), Zoology 7: 1-140. https://www.biodiversitylibrary.org/page/27756163
  • GEISLER J. H., MCGOWEN M. R., YANG G. & GATESY J. 2011. - A supermatrix analysis of genomic, morphological, and paleontological data for crown Cetacea. BMC Evolutionary Biology 11 (112): 1-22. https://doi.org/10.1186/1471-2148-11-112
  • GODFREY S. J. & LAMBERT O. 2023. - Miocene toothed whales (Odontoceti) from Calvert Cliffs, Atlantic Coastal Plain, USA, in GODFREY S. J. (ed.), The Geology and Vertebrate Paleontology of Calvert Cliffs, Maryland, USA. Vol. 107. Smithsonian Contributions to Paleobiology, Washington, D.C.: 49-186. https:// doi.org/10.5479/si.1943-6688.100
  • GUO Z. & KOHNO N. 2021. - A new kentriodontid (Cetacea: Odontoceti) from the early to middle Miocene of the western North Pacific and a revision of kentriodontid phylogeny. PeerJ 9: e10945. https://doi.org/10.7717/peerj.10945
  • GUO Z. & KOHNO N. 2023. - An Early Miocene kentriodontoid (Cetacea: Odontoceti) from the western North Pacific, and its implications for their phylogeny and paleobiogeography. PLoS ONE 18 (2): e0280218. https://doi.org/10.1371/journal. pone.0280218
  • HEYNING J. E. 1989. - Comparative facial anatomy of beaked whales (Ziphiidae) and a systematic revision among the families of extant Odontoceti. Contributions in Science, Natural History Museum of Los Angeles County 405: 1-64.
  • ICHISHIMA H. 1995. - A new fossil kentriodontid dolphin (Cetacea; Kentriodontidae) from the Middle Miocene Takinone Formation, Hokkaido, Japan. The Island Arc 3: 473-485. https://doi. org/10.1111/j.1440-1738.1994.tb00126.x
  • ICHISHIMA H., KAWABE S. & SAWAMURA H. 2021. - The socalled foramen singulare in cetacean periotics is actually the superior vestibular area. The Anatomical Record: 1-8. https:// doi.org/10.1002/ar.24585
  • ICHISHIMA H., BARNES L. G., FORDYCE R. E., KIMURA M. & BOHASKA D. J. 1995. - A review of kentriodontine dolphins (Cetacea; Delphinoidea; Kentriodontidae): systematics and biogeography. The Island Arc 3: 486-492. https://doi. org/10.1111/j.1440-1738.1994.tb00127.x
  • KAZAR E. 2005. - A new kentriodontid (Cetacea: Delphinoidea) from the Middle Miocene of Hungary. Mitteilungen aus dem Museum fur Naturkunde Berlin, Geowissenschaftliche Reihe 8: 53-73. https://doi.org/10.1002/mmng.200410004
  • KAZAR E. & GRIGORESCU D. 2005. - Revision of Sarmatodelphis moldavicus Kirpichnikov, 1954 (Cetacea: Delphinoidea), from the Miocene of Kishinev, Republic of Moldavia. Journal of Vertebrate Paleontology 25: 929-935.https://doi.org/10.1671/0272- 4634(2005)025[0929:ROSMKC]2.0.CO;2
  • KAZAR E. & HAMPE O. 2014. - A new species of Kentriodon (Mammalia, Odontoceti, Delphinoidea) from the middle/late Miocene of Gross Pampau (Schleswig-Holstein, North Germany). Journal of Vertebrate Paleontology 34 (5): 1216-1230.https://doi.org/10. 1080/02724634.2014.857347
  • KELLOGG R. 1927. - Kentriodon pernix, a Miocene porpoise from Maryland. Proceedings of the United States National Museum 69 (10): 1-55. http://hdl.handle.net/10088/15706
  • KIMURA T. & HASEGAWA Y. 2019. - A new species of Kentriodon (Cetacea, Odontoceti, Kentriodontidae) from the Miocene of Japan. Journal of Vertebrate Paleontology: e1566739. https://doi. org/10.1080/02724634.2019.1566739
  • KIMURA T. & HASEGAWA Y. 2024. - New fossil lipotid (Cetacea, Delphinida) from the Upper Miocene of Japan. Paleontological Research 28 (4): 1-23. https://doi.org/10.2517/PR220027
  • LAMBERT O., ESTEVENS M. & SMITH R. 2005. - A new kentriodontine dolphin from the Middle Miocene of Portugal. Acta Palaeontologica Polonica 50 (2): 239-248.
  • LAMBERT O., BIANUCCI G., URBINA M. & GEISLER J. H. 2017. - A new inioid (Cetacea, Odontoceti, Delphinida) from the Miocene of Peru and the origin of modern dolphin and porpoise families. Zoological Journal of the Linnean Society 179 (4): 919-946.https:// doi.org/10.1111/zoj.12479
  • LAMBERT O., MUIZON C. DE, MALINVERNO E., DI CELMA C., URBINA M. & BIANUCCI G. 2018. - A new odontocete (toothed cetacean) from the Early Miocene of Peru expands the morphological disparity of extinct heterodont dolphins. Journal of Systematic Palaeontology 16 (12): 981-1016. https://doi.org/10. 1080/14772019.2017.1359689
  • LAMBERT O., COLLARETA A., BENITES-PALOMINO A., DI CELMA C., MUIZON C. DE, URBINA M. & BIANUCCI G. 2021. - A new small, mesorostrine inioid (Cetacea, Odontoceti, Delphinida) from four upper Miocene localities in the Pisco Basin, Peru. Papers in Palaeontology 7 (2): 1043-1064. https://doi. org/10.1002/spp2.1332
  • MALINVERNO E., BOSIO G., GASTALDELLO M. E., PELLEGRINO L., BIANUCCI G., COLLARETA A., GARIBOLDI K., URBINA M., VILLA I. M., DI CELMA C. 2025. - The early depositional history of the Pisco Formation (Middle to Upper Miocene, Peru). Newsletters on Stratigraphy 58 (1): 99-123.https://doi.org/10.1127/nos/2025/0864
  • MARX F. G., LAMBERT O. & MUIZON C. DE 2017. - A new Miocene baleen whale from Peru deciphers the dawn of cetotheriids. Royal Society Open Science 4 (9): 170560. https://doi. org/10.1098/rsos.170560
  • MARX F. G., LAMBERT O. & UHEN M. D. 2016. - Cetacean paleobiology. John Wiley & Sons, Chichester, 319 p. (Topics in Paleobiology).
  • MCCURRY M. R. & PYENSON N. D. 2019. - Hyper-longirostry and kinematic disparity in extinct toothed whales. Paleobiology 45 (1): 21-29. https://doi.org/10.1017/pab.2018.33
  • MCGOWEN M. R., SPAULDING M. & GATESY J. 2009. - Divergence date estimation and a comprehensive molecular tree of extant cetaceans.Molecular Phylogenetics and Evolution 53 (3): 891-906. https://doi.org/10.1016/j.ympev.2009.08.018
  • MCGOWEN M. R., MONTGOMERY S. H., CLARK C. & GATESY J. 2011. - Phylogeny and adaptive evolution of the braindevelopment gene microcephalin (MCPH1) in cetaceans.BMC Evolutionary Biology 11 (1): 1. https://doi.org/10.1186/1471- 2148-11-98
  • MEAD J. G. & FORDYCE R. E. 2009. - The therian skull: a lexicon with emphasis on the odontocetes. Smithsonian Contributions to Zoology 627: 1-248. https://doi.org/10.5479/si.00810282.627
  • MUIZON C. DE 1988a. - Les vertebres fossiles de la Formation Pisco (Perou). Troisieme partie: Les Odontocetes (Cetacea, Mammalia) du Miocene. Travaux de l'Institut francais d'Etudes andines 42: 1-244.
  • MUIZON C. DE 1988b. - Les relations phylogenetiques des Delphinida. Annales de Paleontologie 74: 159-227.
  • MURAKAMI M., SHIMADA C., HIKIDA Y., SOEDA Y. & HIRANO H. 2014. - Eodelphis kabatensis, a new name for the oldest true dolphin Stenella kabatensis Horikawa, 1977 (Cetacea, Odontoceti, Delphinidae), from the upper Miocene of Japan, and the phylogeny and paleobiogeography of Delphinoidea. Journal of Vertebrate Paleontology 34 (3): 491-511. https://doi.org/10.108 0/02724634.2013.816720
  • NOBILE F., COLLARETA A., PERENZIN V., FORNACIARI E., GIUSBERTI L. & BIANUCCI G. 2024. - Dawn of the delphinidans: New remains of Kentriodon from the Lower Miocene of Italy shed light on the early radiation of the most diverse extant cetacean clade. Biology 13 (2): 114. https://doi.org/10.3390/biology13020114
  • OCHOA D., SALAS-GISMONDI R., DEVRIES T. J., BABY P., MUIZON C. DE, ALTAMIRANO A., BARBOSA-ESPITIA A., FOSTER D. A., QUISPE K. & CARDICH J. 2021. - Late Neogene evolution of the Peruvian margin and its ecosystems: a synthesis from the Sacaco record. International Journal of Earth Sciences: 1-31. https://doi. org/10.1007/s00531-021-02003-1
  • ORLIAC M., ORLIAC C., ORLIAC M. & HAUTIN A. 2020. - A delphinid petrosal bone from a gravesite on Ahu Tahai, Easter Island: taxonomic attribution, external and internal morphology. MorphoMuseum 6 (2): 1-12.https://doi.org/10.18563/journal.m3.91
  • PEREDO C. M., UHEN M. D. & NELSON M. D. 2018. - A new kentriodontid (Cetacea: Odontoceti) from the early Miocene Astoria Formation and a revision of the stem delphinidan family Kentriodontidae. Journal of Vertebrate Paleontology: e1411357. https://doi.org/10.1080/02724634.2017.1411357
  • PERRIN W. F. 1975. - Variation of spotted and spinner porpoise (genus Stenella) in the eastern Pacific and Hawaii. Bulletin of the Scripps Institution of Oceanography University of California, San Diego La Jolla, California 21: 1-206.
  • PERRIN W. F. & MYRICK A. JR 1980. - Age determination of toothed whales and sirenians, in Proceedings International conference on determining age of odontocete cetaceans (and sirenians) La Jolla, California, September 5-19, 1978. Reports of the International Whaling Commission: 1-229.
  • RENSBERGER J. M. 1969. - A new iniid cetacean from the Miocene of California. University of California Publications in Geological Sciences 82: 1-43.
  • ROSTON R. A., BOESSENECKER R. W. & GEISLER J. H. 2023. - Evolution and development of the cetacean skull roof: a case study in novelty and homology.Philosophical Transactions of the Royal Society B 378 (1880): 20220086.https://doi.org/10.1098/rstb.2022.0086
  • SALINAS-MARQUEZ F. M., BARNES L. G., FLORES-TRUJILLO J. G. & ARANDA-MANTECA F. J. 2014. - Una especie de delfin fosil (Cetacea; Delphinoidea; Kentriodontoidae) del Mioceno medio de Baja California. Boletin de la Sociedad Geologica Mexicana 66 (1): 145-164.
  • SEAGARS D. J. 1982. -Jaw structure and functional mechanics of six delphinids (Cetacea, Odontoceti) [Master thesis: San Diego State University, 179 p.
  • SWOFFORD D. L. 2003. - PAUP*. Phylogenetic analysis using parsimony (*and other methods). Version 4. Sinauer Associates, Sunderland, Massachusetts.
  • TRUE F. W. 1912. - Description of a new fossil porpoise of the genus Delphinodon from the Miocene Formation of Maryland.Journal of the Academy of Natural Sciences of Philadelphia 15: 165-194.
  • WHITMORE F. C. JR & KALTENBACH J. A. 2008. - Neogene Cetacea of the Lee Creek Phosphate Mine, North Carolina. Virginia Museum of Natural History Special Publication 14: 181-269.
  • WURSIG B., THEWISSEN J. G. M. & KOVACS K. M. 2018. - Encyclopedia of Marine Mammals, third edition. Academic Press, London, 1157 p.