Family Level Synthesis Phylogeny of Extant Arthropoda and Related Phyla

A synthesis tree of 3,399 extant families of Panarthropods was created manually on Treegraph2 utilizing 244 phylogenetic and taxonomic sources. It is inteded for use for ecology, biogeogaphy, comparative biology, and, if possible, conservation research. A full explanation for the construction of the synthesis tree can be accessed in the Methods and References document. 

NOTE: Several families were mistakenly put into the tree twice (Dynomenidae, Erebonnasteridae, Laemobothriiae, Paraphronimidae, and Tegeocranellidae). Duplicates were removed, and the number of standing families is at 3,223.

NOTE2: Errors in the Stomatopod phylogeny were fixed with a redundant source removed. Some explanations in the Methods document were elaborated on.

NOTE3: Phylogeny of Araneae was updated, with changes to the topology of Atypoidea, Lycosoidea, Araneoidea, Dionycha, 'Marronoidea', an Austrochiloidea. The families Hexurellidae, Megahexuride, Mecicobothriidae, Dolomedidae, Ancylometidae, Synaphridae, Fonteferridae, Udubidae, Myrmeciculatoridae, Trachycosmidae, Cithaeronidae, Cicurinidae, Telemidae, Archoleptonetidae, Melloinidae, Psilodercidae, and Rhytidicolidae were added. Additionally, Pergieopidae, Theridiidae, Lamponidae, Synotaxidae, and Senoculidae, despite being included in the original source for the topology of Araneae, were omitted by mistake and are now placed in their updated positions. Other changes include the removal of Heptathelidae and Mackenziellidae (as they are now considered synonymous with Liphistiidae and Sminthurididae respectively) (WSC https://wsc.nmbe.ch/ and Collembola.org https://www.collembola.org/), the addition of Adelphacaridae to Palaeacaroidea, and changing the positions of Heteroscorpionidae and Hemiscorpionidae.

NOTE4: Phylogeny of Araneae updated with Kulkarni, Wood, et al. (2023) taken as the primary source rather than Wheeler et al. (2016).  The families Probathylepadidae (Thecostraca: Iblomorpha) and Dithinozerconidae (Arachnida, Mesostigmata) were added. 

NOTE5: The families Cyrtocoridae (Insecta: Hemiptera) and Arthruridae (Malacostraca: Tanaidacea) were added. Metaxinidae (Insecta: Coleoptera) and Epipygidae (Insecta: Hemiptera) were included in the original source trees but ommitted earlier by mistake. Phylogeny for orders of Malacostraca was updated. Phylogeny of Eriophyoidea, Philopteroidea, Amblypygi, and Coleoptera were updated. 

NOTE6: Phylogeny of Diplura updated. Proterorhagiidae, Dendrochaetidae, Dytiscacaridae and Strandtmanniidae, Chetochelacaridae, and Dobrodesmidae added. Placements of Eriorhynchidae and Pentapalpidae updated. Phylogeny of Hydrachnidia updated. Cornirostridae, a family of mollusks placed into the tree by mistake, was removed. Phylogeny of Caridea updated.

NOTE7: Placement of Xiphosura updated

Phylogenies of Astigmata, Tanaidacea, Strepsiptera, Fulgoromorpha, Opiliones, Noctuoidea, Coccoidea, Calanoida updated

New families added:

Astigmata: Psoralgidae, Knemidokoptidae, Apionacaridae, Heteropsoridae, Pontoppidaniidae, Cerophagidae, Rhizoglyphidae, Bulacaridae, Horstiidae, Lycogylphidae
Tanaidacea: Julmarichardiidae, Caudalongidae, Hamatipedidae, Meromonakanthidae, Paranarthrurellidae, Paratyphlotanaidae
Sphaerotheriida: Cyliosomatidae
Decapoda: Sternostylidae, Polyjacquesidae, Poupiniidae, Pirimelidae
Araneae: Systariidae
Amphipoda: Iciliidae, Spelaeorchestiidae
Blattodea: Hodotermopsidae
Dermaptera: Gonolabinidae
Orthoptera: Mastacideidae, Lathiceride, Pteroplistidae
Opiliones: Ampycidae, Metasarcidae, Prostygnidae, Gerdesiidae, Askawachidae, Icaleptidae, Kimulidae, Guasinidae, Stygnommatidae, Pyramidopidae, Tithaeidae, Beloniscidae, Cryptomastridae, Lomanellidae, Buemarinoidae, Parasironidae
Odonata: Rimanellidae, Protoneuridae, Pseudocorduliidae, Gomphomacromiidae, Austrocorduliidae, Idomacromiidae, Idionychidae, Lauromacromiidae, Macromidiidae, Neophyidae, Aeschnosomatidae
Trombidiiformes: Iolinidae, Triophtydeidae, Stigmocheylidae, Pomerantziidae, Allochaetophoridae, Tuckerellidae, Tanaupodidae, Amphotrombiidae, Kodiakellidae, Oribellidae, Anderemaeidae, Cerocepheidae
Mesostigmata: Discourellidae
Trichoptera: Plectrotarsidae
Lepidoptera: Ashinagidae, Epimarptidae, Pterolonchidae, Schistonoeidae, Anomosetidae, Prodidactidae, Acanthopteroctetidae, Scranciidae, Dryadaulidae, Ustyurtiidae, Argyresthiidae, Scythropiidae, Tonzidae, Simaethistidae, Whalleyanidae
Hemiptera: Xenococcidae, Quinococcidae, Rhizoecidae, Micrococcidae, Cryptococcidae, Halimococcidae, Polyctenidae, Medocostidae, Thyreocoridae, Phloeidae
Hymenoptera: Austrocynipidae, Paraulacidae, Proctorenyxidae, Peradeniidae, Eremiaspheciidae, Entosericidae, Geoseclionidae, Austroniidae, Ismaridae, Maamingidae
Calanoida: Discoidae, Parkiidae, Rostrocalanidae, Kyphocalanidae, Ryocalanidae
Cyclopoida: Nodocopsidae, Enteropsidae, Pseudotaeniacanthidae
Siphonostomatoida: Samarusidae, Scottomyzontidae, Rataniidae
Anaspidacea: Patagonaspididae

Families included in existing sources but previously omitted by mistake. This includes clade which were not considered families when these studies were published but have since been raised to family rank:

Thysanoptera: Thripidae
Cyclopoida: Clausiidae
Calanoida: Ryocalanidae
Embioptera: Paedembiidae, Embonychidae
Blattodea: Pasammotermitidae, Heterotermitidae
Lepidoptera: Macropiratidae, Tridentaformidae, Thyatiridae, Ethmiidae, Meessiidae
Hymenoptera: Diplolepididae

Families removed: 

Calanoida: Calocalanidae, Ridgewayiidae, Mecynoceridae (synonymization)
Harpacticoida: Cancrincolidae (synonymization)
Ostracoda: Protocytheridae (fossil)
Tanaidacea: Protoapseudoidae, Ophthalmapseudidae and Eucryptocaridae (fossil)
Araneae: Fonteferreidae (dubious)
Hemiptera: Epipygidae, Phacopteronidae, Homotomidae (synonymization)
Decapoda: Pisidae (synonymization)
Psocodea: Trimenopodidae, Lipeuridae, Trichophilopteridae (synonymization)
Acariformes: Gretacaridae, Bandakiopsidae, Nudomideopsidae, Pterochthoniidae, Pterobatidae (synonymization)
Neuroptera: Ascalaphidae (synonymization)
Dermaptera: Karschiellidae (synonymization)
Trichoptera: Ptilocolepidae, Anomalopsychidae (synonymization)
Ephemeroptera: Chromarcyidae (synonymization)
Diptera: Cremifaniidae, Rhinophoridae, Manotinae, Rhiniidae, Tanynderidae (synonymization)
Coleoptera: Drillidae, Propalticidae (synonymization)
Collembola: Microfalculidae (synonymization)