Published December 5, 2025 | Version v1
Taxonomic treatment Open

Dicharax khasiacus

  • 1. Centre for Agricultural Research, HUN-REN Centre for Agricultural Research, 2462 Martonvásár, Brunszvik u. 2, Hungary. & Department of Soil, Water and Natural Sciences, Albert Kázmér Faculty of Agricultural and Food Sciences of Széchenyi István University, Vár 2., 9200 Mosonmagyaróvár, Hungary.
  • 2. Department of Morphology, Systematics and Phylogeny of Animals, Faculty of Biology, University of Belgrade, Studentski trg 16, 11000 Belgrade, Serbia.
  • 3. SM Sehgal Foundation Center for Biodiversity and Conservation, Ashoka Trust for Research in Ecology and the Environment (ATREE), Royal Enclave, Sriramapura, Jakkur PO., Bangalore 560064, India. & Manipal Academy for Higher Education, Madhav Nagar, Manipal 576104, India. & Yenepoya Research Center, University Road, Derlakatte, Mangalore, India.

Description

Dicharax khasiacus (Godwin-Austen, 1871)

Fig. 68

Alycaeus Khasiacus Godwin-Austen, 1871: 90, pl. 3 fig. 4.

Alycaeus duorugosus Godwin-Austen, 1914: 391. Syn. nov.

Alycaeus khasiacus – Sowerby 1877: pl. 1, species 8. — Godwin-Austen 1914: 356, 376–377, 393, pl. 143 figs 7, 7a–b.

Alycaeus (Dicharax) khasiacus – Kobelt 1902: 372–373. — Gude 1921: 257–258.

Alycaeus khasiacus var. – Godwin-Austen 1914: 356.

Alycaeus (Dicharax) duorugosus – Gude 1921: 249.

Chamalycaeus (Dicharax) duorugosus – Ramakrishna et al. 2010: 59.

Chamalycaeus (Dicharax) khasiacus – Ramakrishna et al. 2010: 62.

Dicharax (?) khasiacus – Páll-Gergely et al. 2020: 90.

Dicharax (?) duorugosus – Páll-Gergely et al. 2020: 86.

Diagnosis

Dicharax khasiacus is characterized by the rounded outline, the short, smooth R2, and the two, anteriorly situated R3 swellings.

Type material examined

INDIA • 29 syntypes of A. khasiacus (Fig. 68A–E); Lailangkote, Khasi Hills; NHMUK 1903.7.1.2650 • 1 syntype of A. duorugosus (Fig. 68F–J); Burrail; Godwin-Austen coll.; NHMUK 1903.7.1.2771.

Additional material examined

INDIA • 4 shells; Jawai, Assam; Oberwimmer coll.; NHMW • 2 shells; same data as for preceding; NHMW 14711 • 4 shells; Jawai, India; Dr. Leo P. Rušnov coll., ex coll. Dr W. Blume ex coll. W.F. Webb; NHMW 15168.

Type localities

“On the highest parts of the Khasi and Jiantia Hills” (A. khasiacus); “Burrail Range, Naga ” “Also Angaoluo Trigonometrical Station, No. 2572; South Barak, No. 2629, and Munipur, No. 2654 B.M.” (A. duorugosus).

Differential diagnosis

Dicharax lahupaensis has a ribbed R2 (smooth in D. khasiacus) and a thicker peristome. Dicharax akhaensis is larger, has a shorter R3 swelling, and a less fringed aperture.

Remarks

The single examined syntype of A. duorugosus fits the morphological variability of the 29 syntypes of A. khasiacus; therefore the former is a junior synonym of the latter.

Notes

Published as part of Páll-Gergely, Barna, Gojšina, Vukašin & Aravind, Neelavar Ananthram, 2025, A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae), pp. 1-158 in European Journal of Taxonomy 1029 on pages 115-117, DOI: 10.5852/ejt.2025.1029.3131, http://zenodo.org/record/17864673

Files

Files (2.7 kB)

Name Size Download all
md5:06e85f0e890b51ee0bb9ceaa61664843
2.7 kB Download

System files (30.9 kB)

Name Size Download all
md5:4051153477d6d4ae9b718b29c61b5fa8
30.9 kB Download

Linked records

Additional details

Biodiversity

References

  • Godwin-Austen H. H. 1871. Descriptions of the species of Alycaeinae known to inhabit the Khasi Hill ranges. Journal of the Asiatic Society of Bengal 40: 87-93. Available from https://www.biodiversitylibrary.org/page/35630903 [accessed 1 May 2025].
  • Sowerby G. B. II 1877. Monograph of the genus Alycaeus. In: Conchologia iconica, or Illustrations of the Shells of Molluscous Animals Vol. 20: pls 1-6 and unpaginated text. L. A. Reeve, London. https://doi.org/10.5962/bhl.title.8129
  • Kobelt W. 1902. Das Tierreich. Eine Zusammenstellung und Kennzeichnung der rezenten Tierformen. In Verbindung mit der Deutschen Zoologischen Gesellschaft herausgegeben von der Koniglich Preussischen Akademie der Wissenschaften zu Berlin. Lfg 16. Mollusca: Cyclophoridae. R. Friedlander und Sohn, Berlin. Available from https://www.biodiversitylibrary.org/page/1000211 [accessed 1 May 2025].
  • Gude G. K. 1921. The Fauna of British India including Ceylon and Burma. Mollusca. III. Land operculates (Cyclophoridae, Truncatellidae, Assimineidae, Helicinidae). Taylor and Francis, London. https://doi.org/10.5962/bhl.title.12890
  • Ramakrishna, Mitra S. C. & Dey A. 2010. Annotated Checklist of Indian Land Molluscs. Zoological Survey of India, Kolkata. Available from http://faunaofindia.nic.in/PDFVolumes/occpapers/306/index.pdf [accessed 1 May 2025].
  • Pall-Gergely B., Sajan S., Tripathy B., Meng K., Asami T. & Ablett J. D. 2020. Genus-level revision of the Alycaeidae (Gastropoda, Cyclophoroidea), with an annotated species catalogue. ZooKeys 981: 1-220. https://doi.org/10.3897/zookeys.981.53583