Published August 9, 2021
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Table 2 in Between sand, rocks and branches: an integrative taxonomic revision of Angolan Hemidactylus Goldfuss, 1820, with description of four new species
Authors/Creators
- 1. CIBIO Centro de Investigacao em Biodiversidade e Recurso Geneticos, Universidade do Porto, Rua Padre Armando Quintas, Campus de Vairao, 4485 - 661 Vairao, Portugal & Faculdade de Ciencias da Universidade do Porto, Porto, Portugal
- 2. Port Elizabeth Museum (Bayworld), P. O. Box 13147, Humewood 6013, South Africa & Department of Nature Conservation Management, Natural Resource Science and Management Cluster, Faculty of Science, George Campus, Nelson Mandela University, George, South Africa
- 3. Departament de Biologia (Genetica), Facultad de Ciencias, Universidad Autonoma de Madrid (UAM), c / Darwin 2, 28049, Madrid, Spain & Centro de Investigacion en Biodiversidad y Cambio Global CIBC-UAM, Facultad de Ciencias, Universidad Autonoma de Madrid, c / Darwin 2, 28049 - Madrid, Spain
- 4. Museum of Zoology, Senckenberg Natural History Collections Dresden, Ko ̈ nigsbru ̈ cker Landstr. 159, D- 01109, Dresden, Germany
- 5. Fundacao Kissama, Rua 60 Casa 560, Lar do Patriota, Luanda, Angola
- 6. CIBIO Centro de Investigacao em Biodiversidade e Recurso Geneticos, Universidade do Porto, Rua Padre Armando Quintas, Campus de Vairao, 4485 - 661 Vairao, Portugal & Faculdade de Ciencias da Universidade do Porto, Porto, Portugal & TwinLab CIBIO / ISCED, Instituto Superior de Ciencias da Educacao da Huila, Rua Sarmento Rodrigues s / n, Lubango, Angola
- 7. CIBIO Centro de Investigacao em Biodiversidade e Recurso Geneticos, Universidade do Porto, Rua Padre Armando Quintas, Campus de Vairao, 4485 - 661 Vairao, Portugal & Fundacao Kissama, Rua 60 Casa 560, Lar do Patriota, Luanda, Angola & TwinLab CIBIO / ISCED, Instituto Superior de Ciencias da Educacao da Huila, Rua Sarmento Rodrigues s / n, Lubango, Angola
Description
Table 2. Comparison between the different Angolan Hemidactylus (excluding H. mabouia and including H. gramineus from DRC). Measurements are represented in mm (average and standard deviation), based on newly collected material for this work with exception of H. nzingae, H. gramineus and H. paivae, where morphological measurements were compiled from information available in Ceríaco et al. (2020a, 2021) and information collected for this work. For abbreviations see Material and Methods section.
| Species | H. faustus sp. nov. | H. pfindaensis sp. nov. | H. carivoensis sp. nov. | H. benguellensis | H. cinganji sp. nov. | H. longicephalus LINEAGE 1 | H. longicephalus LINEAGE 2 | H. paivae | H. bayonii | H. vernayi | H. gramineus | H. nzingae |
| Specimens | n=10 | n=4 | n=9 | n=12 | n=2 | n=6 | n=9 | n=18 | n=6 | n=25 | n=2 | n=12 |
| SVL | 39.38 ± 3.35 | 45.54 ± 3.12 | 38,18 ± 5,16 | 49.66 ± 8.12 | 48.72 ± 5.19 | 46.21 ± 8.76 | 47.30 ± 3.09 | 57.31 ± 2.19 | 36.29 ± 3.46 | 32.52 ± 4.07 | 36.65 ± 4.03 | 44.28 ± 3.88 |
| TL | - | 32.93 | 36,8 ± 7,64 | 40.23 ± 8.65 | 42.77 ± 0 | 44.20 ± 6.94 | 48.52 ± 1.94 | 63.05 ± 7.14 | 30.06 ± 11.11 | 25.92 ± 4.19 | 34.40 ± 0.99 | 40.33 ± 3.48 |
| TRL | 16.83 ± 2.51 | 17.85 ± 3.76 | 15,82 ± 1,92 | 21.94 ± 5.93 | 21.33 ± 4.13 | 19.25 ± 4.29 | 18.98 ± 0.09 | 21.45 ± 2.89 | 15.02 ± 2.62 | 13.95 ± 1.88 | 15.85 ± 2.33 | 19.41 ± 2.58 |
| BW | 7.44 ± 0.93 | 7.43 ± 0.41 | 6,99 ± 1,92 | 10.056 ± 1.83 | 10.14 ± 0.50 | 8.28 ± 3.26 | 8.5 ± 0.81 | 9.81 ± 0.47 | 6.69 ± 0.78 | 6.12 ± 1.26 | 5 ± 0.14 | 7.27 ± 1.29 |
| FL | 9.10 ± 1.36 | 6.99 ± 1.35 | 5,79 ± 0,98 | 7.32 ± 1.81 | 7.86 ± 1.30 | 6.24 ± 1.53 | 8.57 ± 0.12 | 7.12 ± 1.14 | 5.57 ± 0.81 | 5.89 ± 1.23 | 4.70 ± 0.28 | 6.06 ± 0.77 |
| CL | 6.32 ± 0.54 | 8.03 ± 1.73 | 6,75 ± 1,18 | 7.74 ± 1.47 | 8.39 ± 1.98 | 7.11 ± 0.76 | 5.47 ± 0.92 | 7.21 ± 1.47 | 6.30 ± 0.63 | 4.90 ± 1.18 | 5.55 ± 0.21 | 7.03 ± 0.68 |
| HL | 11.62 ± 1.09 | 13.19 ± 1.40 | 11,10 ± 0,69 | 14.25 ± 2.86 | 12.99 ± 2.62 | 13.12 ± 2.09 | 14.02 ± 0.66 | 14.55 ± 0.59 | 10.13 ± 1.49 | 9.59 ± 0.79 | 10.5 ± 0.71 | 12.18 ± 0.27 |
| HW | 8.22 ± 0.79 | 8.65 ± 0.85 | 6,81 ± 1,10 | 9.72 ± 1.73 | 9.57 ± 0.84 | 8.60 ± 1.55 | 9.38 ± 0.52 | 10.02 ± 0.62 | 6.75 ± 0.56 | 6.01 ± 0.98 | 6.35 ± 0.49 | 8.11 ± 0.57 |
| HH | 4.95 ± 0.71 | 5.26 ± 0.28 | 4,21 ± 0,69 | 5.52 ± 0.97 | 6.32 ± 0.58 | 4.93 ± 0.99 | 5.65 ± 0.27 | 6.12 ± 0.28 | 4.37 ± 0.49 | 4.04 ± 0.52 | 4.25 ± 0.49 | 5.08 ± 0.55 |
| OD | 2.87 ± 0.29 | 2.81 ± 0.18 | 2,68 ± 0,36 | 2.97 ± 0.39 | 3.14 ± 0.13 | 2.97 ± 0.48 | 3.02 ± 0.11 | 3.21 ± 0.18 | 2.42 ± 0.14 | 2.25 ± 0.29 | 2.10 ± 0.42 | 2.81 ± 0.34 |
| NE | 3.39 ± 0.49 | 3.81 ± 0.47 | 3,38 ± 0,51 | 4.52 ± 0.78 | 4.18 ± 0.20 | 4.13 ± 0.64 | 4.45 ± 0.36 | 4.54 ± 0.57 | 2.88 ± 0.67 | 2.68 ± 0.35 | 2.95 ± 0.35 | 4.26 ± 0.68 |
| SE | 4.6325 ± 0.31 | 5.32 ± 0.21 | 4,46 ± 0,70 | 6.03 ± 1.13 | 5.38 ± 0.61 | 5.29 ± 0.79 | 6.09 ± 0.85 | 5.80 ± 0.51 | 4.02 ± 0.71 | 3.58 ± 0.45 | 4.25 ± 0.07 | 5.21 ± 0.34 |
| EE | 3.29 ± 0.36 | 3.53 ± 0.21 | 2,91 ± 0,39 | 3.89 ± 0.68 | 3.85 ± 0.48 | 3.21 ± 0.63 | 3.66 ± 0.46 | 3.83 ± 0.27 | 2.9 ± 0.46 | 2.75 ± 0.39 | 2.75 ± 0.21 | 3.56 ± 0.23 |
| EH | 0.99 ± 0.15 | 0.84 ± 0.25 | 1,73 ± 2,67 | 1.25 ± 0.55 | 1.17 ± 0.42 | 0.92 ± 0.25 | 0.98 ± 0.09 | 0.95 ± 0.37 | 0.69 ± 0.19 | 0.57 ± 0.12 | 0.7 ± 0.0 | 1.07 ± 0.17 |
| IN | 1.56 ± 0.24 | 1.36 ± 0.05 | 1,21 ± 0,20 | 3.75 ± 6.35 | 1.42 ± 0.12 | 1.41 ± 0.21 | 1.59 ± 0.15 | 1.63 ± 0.18 | 1.27 ± 0.12 | 1.22 ± 0.18 | - | - |
| OI | 3.97 ± 0.40 | 3.9 ± 1.41 | 3,51 ± 0,61 | 4.20 ± 0.66 | 4.81 ± 0.04 | 3.47 ± 0.72 | 4.68 ± 0.17 | 4.85 ± 0.04 | 3.3 ± 0.35 | 2.98 ± 0.49 | - | - |
| Precloacal-Femoral Pores | 17-19 | 8 | 15-22 | 23-33 | 26-28 | 5-11 | 6 | 6-8 | 4-9 | 4-6 | 8 | 7-8 |
| Post-cloacal (tail) spurs | 1/1 | 1/1 | 1/1 | 1-4/1-4 | 1-2/1-2 | 0-1/0-1 | 1-1/1-1 | - | 1/1 | 1-2/1-2 | - | - |
| Scales across belly | 29-32 | 28-30 | 32-38 | 33-41 | 30-32 | 28-37 | 28-36 | 28-34 | 25-32 | 32-39 | 23-25 | 22-27 |
| Dorsal tubercles row | 15-17 | 11-12 | 12-18 | 13-18 | 15 | 13-17 | 13-16 | 13-16 | 15-16 | 11-16 | 12-14 | 16-21 |
| Supralabials (right/left) | 9/9 | 10/10 | 11/10-11 | 10-11/10-11 | 9/9 | 10-12 | 10-12 | 10-12/10-12 | 8-9/9-10 | 9-10/9-10 | 10/9-0 | 10-11/10-11 |
| Infralabials (right/left) | 8/8 | 8-10/9-10 | 9/9-10 | 9-10/8-10 | 7-9/8 | 9-10 | 9-10 | 9-10/8-10 | 7-9/7-8 | 7-8/7-8 | 8-9/8-9 | 8-9/8-9 |
| Subcaudals size | Small (<1⁄4 tail width) | Small (<1⁄4 tail width) | Small (<1⁄4 tail width) | Small (<1⁄4 tail width) | Small (<1⁄4 tail width) | Small (<1⁄4 tail width) | Small (<1⁄4 tail width) | Small-Medium | Small (<1⁄4 tail width) | Small (<1⁄4 tail width) | Small (<1⁄4 tail width) | Small (<1⁄4 tail width) |
| # Granular scales between tubercles | 2-3 | 4-5 | 2-3 | 2-5 | 2-3 | 3-6 | 3-4 | 3-9 | 2-4 | 2-4 | 2-3 | 2-3 |
| Lamellae 1st finger | 5-6 | 6-5 | 5 | 6 | 5-6 | 5-6 | 6 | 5-8 | 5-6 | 5-6 | 5 | 5-7 |
| Lamellae 4th finger | 7 | 7 | 6-7 | 6-8 | 7 | 7-8 | 7-8 | 8-9 | 7 | 7 | 7 | 6-8 |
| Lamellae 1st toe | 5-6 | 5-6 | 5 | 5-7 | 6 | 5-7 | 5-6 | 6-7 | 6 | 5-6 | 5 | 5-7 |
| Lamellae 4th toe | 9-8 | 7-8 | 9 | 8-9 | 8 | 8-10 | 8-19 | 9-11 | 8-10 | 7-8 | 7 | 9 |
Notes
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Additional details
Identifiers
References
- Ceriaco, LM, Agarwal, I, Marques, MP, Bauer, AM, 2020a. A review of the genus Hemidactylus Goldfuss, 1820 (Squamata: Gekkonidae) from Angola, with the description of two new species. Zootaxa 4746 (1): 1 - 71, DOI: https://doi.org/10.11646/zootaxa.4746.1.1
- Ceriaco, L. M., Bauer, AM, Kusamba, C., Agarwal, I., Greenbaum, E., 2021. . A New Species of Ground-Dwelling Hemidactylus (Squamata: Gekkonidae) from Southwestern Democratic Republic of the Congo. Journal of Herpetology, 55(2), 105-111. https://doi.org/10.1670/20-094