Published December 8, 2025
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Appendix 1 in Integrative taxonomy supports the description of two new species of Macrobiotidae (Tardigrada, Eutardigrada) from Kristianstads Vattenrike Biosphere Reserve in Sweden
Authors/Creators
- 1. Department of Zoology, Swedish Museum of Natural History, Box 50007, 104 05 Stockholm, Sweden.
- 2. Department of Life Sciences, University of Modena and Reggio Emilia, 41124 Modena, Italy.
- 3. Department of Environmental Science, Kristianstad University, SE- 291 88 Kristianstad, Sweden.
Description
Appendix 1 (continued on next 14 pages). GenBank Accession numbers for COI, 18S, ITS, and 28S sequences of all specimens analysed in this study. New sequences are highlighted in red.
| Taxon | Specimen | COI | 18S | ITS | 28S | Taxon name in GenBank | Notes | Reference in GenBank |
|---|---|---|---|---|---|---|---|---|
| Diaforobiotus islandicus | IS.042 | MT808072 | MT812470 | MT812597 | MT812461 | Diaforobiotus islandicus | Stec et al. 2020d | |
| Macrobiotus aff. polonicus | S165.1 | MW593929 | MW588026 | MW588021 | MW588032 | Macrobiotus aff. polonicus S165 | Vecchi & Stec 2021 | |
| Macrobiotus aff. polonicus | S165.2 | MW593930 | MW588027 | MW588020 | MW588033 | Macrobiotus aff. polonicus S165 | Vecchi & Stec 2021 | |
| Macrobiotus almadai | ZYL123 | MW990245 | MW995180 | Macrobiotus almadai | Wang et al. (unpubl.) | |||
| Macrobiotus annewintersae | S207.1 | MW593927 | MW588025 | MW588019 | MW588030 | Macrobiotus annewintersae | Vecchi & Stec 2021 | |
| Macrobiotus annewintersae | S207.2 | MW593928 | MW588024 | MW588018 | MW588031 | Macrobiotus annewintersae | Vecchi & Stec 2021 | |
| Macrobiotus ariekammensis ariekammensis | NO.393.01 | MZ460999 | MZ463668 | MZ463657 | MZ463674 | Macrobiotus ariekammensis ariekammensis | Stec et al. 2022 | |
| Macrobiotus ariekammensis groenlandicus | GL.018.04 | MZ461006 | MZ463664 | MZ463654 | MZ463679 | Macrobiotus ariekammensis groenlandicus | Stec et al. 2022 | |
| Macrobiotus azzunae | C4218_T5 | MW698698 | Macrobiotus sp. 1 MC-2021 | temporary taxon name has not been updated in GenBank at time of writing | Marnissi et al. 2021 | |||
| Macrobiotus azzunae | C4218_V4 | MW698697 | MW695447 | MW695454 | MW695450 | Macrobiotus sp. 1 MC-2021 | temporary taxon name has not been updated in GenBank at time of writing | Marnissi et al. 2021 |
| Macrobiotus basiatus | USA /NEL/1 | MT502116 | MT498094 | MT505165 | MT488397 | Macrobiotus sp. 1 DG-2020 | temporary taxon name has not been updated in GenBank at time of writing | Nelson et al. 2020 |
| Macrobiotus birendrai | CN8.101 | MW656266 | MW680641 | MW680418 | MW680644 | Macrobiotus birendrai | Kayastha et al. 2021 | |
| Macrobiotus caelestis | KG.007 | MK737922 | MK737073 | MK737072 | MK737071 | Macrobiotus caelestis | Coughlan et al. 2019 | |
| Macrobiotus canaricus | ES.004_H1 | MH057765 | MH063925 | MH063928 | MH063934 | Macrobiotus canaricus | Stec et al. 2018c | |
| Macrobiotus canaricus | ES.004_H2 | MH057766 | Macrobiotus canaricus | Stec et al. 2018c | ||||
| Macrobiotus cf. hufelandi | C2959a | HQ876590 | OP596303 | Macrobiotus cf. hufelandi 1 MC-2011 | Bertolani et al. 2011 | |||
| Macrobiotus cf. muralis | C3251_2 | OP561785 | OP596299 | Macrobiotus cf. muralis | Bertolani et al. 2023 | |||
| Macrobiotus cf. muralis | C3251_3 | OP561786 | OP596300 | Macrobiotus cf. muralis | Bertolani et al. 2023 | |||
| Macrobiotus cf. muralis | C3251_4 | OP561788 | Macrobiotus cf. muralis | Bertolani et al. 2023 | ||||
| Macrobiotus cf. muralis | C3251_5 | OP561789 | Macrobiotus cf. muralis | Bertolani et al. 2023 | ||||
| Macrobiotus cf. muralis | C3251_FA | OP561787 | OP596301 | Macrobiotus cf. muralis | Bertolani et al. 2023 | |||
| Macrobiotus cf. nelsonae | 1 | HQ604965 | Macrobiotus | Bertolani et al. 2014 | ||||
| Macrobiotus cf. recens | ES.006 haplotype 1 | MH057768 | MH063927 | MH063932 | MH063936 | Macrobiotus cf. recens DS-2018 | Stec et al. 2018c | |
| Macrobiotus cf. sapiens | S12.Mac.1 | OK662997 | OK663226 | OK663215 | OK663237 | Macrobiotus cf. sapiens | Vecchi et al. 2022c | |
| Macrobiotus cf. shonaicus | MM1-2018 | LC431582 | LC431591 | Macrobiotus cf. shonaicus | Matsumoto & Sugiura (unpubl.) | |||
| Macrobiotus crustulus | GF.271 | MT260371 | MT261912 | MT261907 | MT261903 | Macrobiotus crustulus | Stec et al. 2020a | |
| Macrobiotus dolosus | C3209_2 | OP561772 | OP596290 | Macrobiotus dolosus | Bertolani et al. 2023 | |||
| Macrobiotus dolosus | C3209_4 | OP561774 | Macrobiotus dolosus | Bertolani et al. 2023 | ||||
| Macrobiotus dolosus | C3209_ US 2 | OP561773 | Macrobiotus dolosus | Bertolani et al. 2023 | ||||
| Macrobiotus dolosus | C3581_V6 | OP561775 | OP596292 | Macrobiotus dolosus | Bertolani et al. 2023 | |||
| Macrobiotus engbergi | haplotype 1 | MN444824 | MN443039 | MN443036 | MN443034 | Macrobiotus engbergi | Vecchi et al. 2024b | |
| Macrobiotus engbergi | haplotype 2 | MN444825 | MN443037 | Macrobiotus engbergi | Vecchi et al. 2024b | |||
| Macrobiotus fontourai | C2861_5 | OP561783 | OP596295 | Macrobiotus fontourai | Bertolani et al. 2023 | |||
| Macrobiotus fontourai | C2861_ US 1 | OP561784 | OP596296 | Macrobiotus fontourai | Bertolani et al. 2023 | |||
| Macrobiotus glebkai | UA.003 | MW246134 | MW247177 | MW247180 | MW247176 | Macrobiotus glebkai | Kiosya et al. 2021 | |
| Macrobiotus hannae | PL.010 | MH057764 | MH063922 | MH063923 | MH063924 | Macrobiotus hannae | Nowak & Stec 2018 | |
| Macrobiotus hufelandi | C2953a | HQ876585 | OP596302 | Macrobiotus hufelandi | Bertolani et al. 2023 | |||
| Macrobiotus hufelandi | S605.Mac.1 | OK662992 | OK663221 | OK663210 | OK663232 | Macrobiotus hufelandi | Bertolani et al. 2023 | |
| Macrobiotus joannae | 1 | HQ604974 | Macrobiotus joannae | Bertolani et al. 2014 | ||||
| Macrobiotus kamilae | IN.030 haplotype 1 | MK737920 | MK737070 | MK737067 | MK737064 | Macrobiotus kamilae | Coughlan & Stec 2019 | |
| Macrobiotus kathyae | US.006.01 | PP386934 | PP391285 | PP391289 | PP391287 | Macrobiotus sp. MV2024 | temporary taxon name has not been updated in GenBank at time of writing | Massa & Vecchi 2024 |
| Macrobiotus kathyae | US.006.02 | PP386935 | PP391286 | PP391290 | PP391288 | Macrobiotus sp. MV2024 | temporary taxon name has not been updated in GenBank at time of writing | Massa & Vecchi 2024 |
| Macrobiotus kathyae | US.006.03 | PP386932 | Macrobiotus sp. MV2024 | temporary taxon name has not been updated in GenBank at time of writing | Massa & Vecchi 2024 | |||
| Macrobiotus kathyae | US.006.04 | PP386933 | Macrobiotus sp. MV2024 | temporary taxon name has not been updated in GenBank at time of writing | Massa & Vecchi 2024 | |||
| Macrobiotus kirghizicus | KG.062.01 | MZ461002 | MZ463666 | MZ463659 | MZ463672 | Macrobiotus kirghizicus | Stec et al. 2022 | |
| Macrobiotus kosmali | M8.1 | OP141639 | OP142472 | OP153786 | OP143765 | Macrobiotus kosmali | Kayastha et al. 2023a | |
| Macrobiotus kristenseni | C3291_A02_V2 | KC193573 | KC193577 | Macrobiotus kristenseni | Guidetti et al. 2013 | |||
| Macrobiotus kyoukenus | C4313_ US 3 | ON809462 | ON818314 | ON818301 | ON818309 | Macrobiotus kyoukenus | Cesari et al. 2022 | |
| Macrobiotus macrocalix | PL.110 | MH057767 | MH063926 | MH063931 | MH063935 | Macrobiotus macrocalix | Stec et al. 2018c | |
| Macrobiotus margoae | US.057 | MN888315 | MN888368 | MN888340 | MN888354 | Macrobiotus margoae | Vecchi & Stec 2021 | |
| Macrobiotus margoae | US.057.1 | MT807927 | MT809072 | MT809098 | MT809084 | Macrobiotus margoae | Stec et al. 2021b | |
| Macrobiotus margoae | US.057.3 | MT807928 | MT809099 | Macrobiotus margoae | Stec et al. 2021b | |||
| Macrobiotus mileri | IL.001.01 | OR544397 | OR543312 | OR543316 | OR543320 | Macrobiotus mileri | Stec 2024 | |
| Macrobiotus mileri | IL.001.02 | OR544398 | OR543313 | OR543317 | OR543321 | Macrobiotus mileri | Stec 2024 | |
| Macrobiotus mileri | IL.001.03 | OR544399 | Macrobiotus mileri | Stec 2024 | ||||
| Macrobiotus mileri | IL.001.04 | OR544400 | Macrobiotus mileri | Stec 2024 | ||||
| Macrobiotus noongaris | AU.031 | MK737919 | MK737069 | MK737066 | MK737063 | Macrobiotus noongaris | Coughlan & Stec 2019 | |
| Macrobiotus ovovittatus | GL.001.01 | OR544395 | OR543311 | OR543314 | OR543318 | Macrobiotus ovovittatus | Stec 2024 | |
| Macrobiotus pallarii | IT.337.1 | MT807924 | MT809069 | MT809094 | MT809081 | Macrobiotus pallarii | Stec et al. 2021b | |
| Macrobiotus pallarii | IT.337.2 | MT807925 | MT809070 | MT809095 | MT809082 | Macrobiotus pallarii | Stec et al. 2021b | |
| Macrobiotus pallarii | IT.337.3 | MT807926 | MT809071 | MT809096 | MT809083 | Macrobiotus pallarii | Stec et al. 2021b | |
| Macrobiotus papei | TZ.027 | MH057763 | MH063881 | MH063921 | MH063880 | Macrobiotus papei | Stec et al. 2018b | |
| Macrobiotus paulinae | KT 951668 | KT935502 | KT935500 | Macrobiotus paulinae | Stec et al. 2015 | |||
| Macrobiotus scoticus | DK.056.1 | OK662989 | OK663217 | OK663206 | OK663228 | Macrobiotus scoticus | Vecchi et al. 2022c | |
| Macrobiotus shonaicus | haplotype 1 | MG757136 | MG757132 | MG757134 | MG757133 | Macrobiotus shonaicus | Stec et al. 2018a | |
| Macrobiotus siderophilus | C2796_2 | OP561776 | OP596293 | Macrobiotus siderophilus | Bertolani et al. 2023 | |||
| Macrobiotus siderophilus | C3282_UFK | OP561777 | Macrobiotus siderophilus | Bertolani et al. 2023 | ||||
| Macrobiotus siderophilus | C3282_UFL | OP561778 | Macrobiotus siderophilus | Bertolani et al. 2023 | ||||
| Macrobiotus siderophilus | C3282_UFM | OP561779 | Macrobiotus siderophilus | Bertolani et al. 2023 | ||||
| Macrobiotus siderophilus | C3282_UFN | OP561780 | OP596294 | Macrobiotus siderophilus | Bertolani et al. 2023 | |||
| Macrobiotus siderophilus | C3282_UFO | OP561781 | Macrobiotus siderophilus | Bertolani et al. 2023 | ||||
| Macrobiotus siderophilus | C3282_UFP | OP561782 | Macrobiotus siderophilus | Bertolani et al. 2023 | ||||
| Macrobiotus sottilei | PL.352 | MW246133 | MW247178 | MW247179 | MW247175 | Macrobiotus sottilei | Kiosya et al. 2021 | |
| Macrobiotus sp. | DI02 | OP684780 | OP696675 | Macrobiotus sp. | Amezcua-Martinez et al. (unpubl.) | |||
| Macrobiotus sp. 1 MC-2022d | UK 12 | OQ525891 | OQ525899 | OQ548104 | OQ525904 | Macrobiotus sp. 1 MC-2022d | Mioduchowsk et al. (unpubl.) | |
| Macrobiotus sp. 1 MC-2022d | UK 14 | OQ525890 | OQ525897 | OQ525905 | Macrobiotus sp. 1 MC-2022d | Mioduchowsk et al. (unpubl.) | ||
| Macrobiotus sp. 1 YW-2020 | BN140005 | MT863704 | MT872214 | Macrobiotus sp. 1 YW-2020 | Wang (unpubl.) | |||
| Macrobiotus sp. 1 ZY-2020 | HP1110201 | MW187003 | MW183923 | Macrobiotus sp. 1 ZY-2020 | Yuan et al. (unpubl.) | |||
| Macrobiotus sp. n. MM-2020 | MAD967 | MT246661 | MT241900 | MT241898 | Macrobiotus sp. n. MM-2020 | Kuzdrowska et al. (unpubl.) | ||
| Macrobiotus terminalis | C2868_N02 | JN673959 | OP596308 | Macrobiotus terminalis | Cesari et al. 2011 | |||
| Macrobiotus vattenrikense sp. nov | 22-003 | PX093655 | PX093663 | PX093644 | PX093649 | COI haplotype 3; 18S haplotype 1; ITS haplotype 1; 28S haplotype 1 | this study | |
| Macrobiotus vattenrikense sp. nov | 22-004-1 | PX093655 | COI haplotype 3 | this study | ||||
| Macrobiotus vattenrikense sp. nov | 22-017 | PX093653 | PX093644 | PX093649 | COI haplotype 1; ITS haplotype 1; 28S haplotype 1 | this study | ||
| Macrobiotus vattenrikense sp. nov | 22-101 | PX093654 | PX093664 | PX093644 | PX093650 | COI haplotype 2; 18S haplotype 2; ITS haplotype 1; 28S haplotype 2 | this study | |
| Macrobiotus vladimiri | FI.068 | MN888327 | MN888375 | MN888347 | MN888360 | Macrobiotus vladimiri | Vecchi & Stec 2021 | |
| Mesobiotus aradasi | KPRI-MF-1 | MW727934 | MW751962 | Mesobiotus cf. furciger | see Vecchi et al. 2024a | Short et al. 2022 | ||
| Mesobiotus aradasi | S1982.Meb.1 | OQ928737 | OQ933000 | OQ940183 | OQ932999 | Mesobiotus aradasi | Vecchi et al. 2024a | |
| Mesobiotus bockebodicus | 22-044 | PQ365775 | PQ367888 | PQ367894 | Mesobiotus sp. 1 SA-2024a | temporary taxon name has not been updated in GenBank at time of writing | Atherton et al. 2025 | |
| Mesobiotus cf. barabanovi | KG.002 | MN313170 | MN310392 | MN310390 | MN310388 | Mesobiotus cf. barabanovi | Kaczmarek et al. 2020a | |
| Mesobiotus cf. furciger | AF01-MF-2 | MW727944 | MW751946 | Mesobiotus cf. furciger | Short et al. 2022 | |||
| Mesobiotus datanlanicus | VN.026 | MK578905 | MK584659 | MK584657 | MK584658 | Mesobiotus datanlanicus | Stec 2019 | |
| Mesobiotus diegoi | ZA.001.02 | OP143857 | OP142526 | OP142514 | OP142520 | Mesobiotus diegoi | Stec 2022a | |
| Mesobiotus dilimanensis | PH.006 | MN257047 | MN257048 | MN257050 | MN257049 | Mesobiotus dilimanensis | Itang et al. 2020 | |
| Mesobiotus efa | Lembolovo.DT262 | OR803035 | OR804457 | OR805169 | OR805135 | Mesobiotus sp. 1 DVT-2023b | temporary taxon name has not been updated in GenBank at time of writing | Tumanov et al. 2024 |
| Mesobiotus emiliae | 22-046 | PQ365768 | PQ367886 | PQ367892 | Mesobiotus emiliae | Atherton et al. 2025 | ||
| Mesobiotus ethiopicus | ET.004 | MF678794 | MF678793 | MN122776 | MF678792 | Mesobiotus ethiopicus | Stec & Kristensen 2017 | |
| Mesobiotus fiedleri | MG.001 | MH676056 | MH681585 | MH681724 | MH681693 | Mesobiotus fiedleri | Kaczmarek et al. 2020a | |
| Mesobiotus furciger | Macro_06_313 | MW727960 | MW751939 | Mesobiotus furciger | Short et al. 2022 | |||
| Mesobiotus harmsworthi | haplotype 1 | MH195150 | MH197146 | MH197154 | MH197264 | Mesobiotus harmsworthi | Kaczmarek et al. 2018 | |
| Mesobiotus hilariae | Novolazarevskaya_ C_1 | KT226108 | KT226071 | Mesobiotus hilariae | Vecchi et al. 2016 | |||
| Mesobiotus huecoensis | S2027.4 | OQ756247 | OQ756248 | Mesobiotus huecoensis | Vecchi et al. 2023 | |||
| Mesobiotus imperialis | VN.061.01 | OL311514 | OL257855 | OL257867 | Mesobiotus imperialis | Stec 2021 | ||
| Mesobiotus insanis | MF 441491 | MF441488 | MF441490 | MF441489 | Mesobiotus insanis | Mapalo et al. 2017 | ||
| Mesobiotus maklowiczi | ZA.002.01 | OP143855 | OP142525 | OP142518 | Mesobiotus maklowiczi | Stec 2022a | ||
| Mesobiotus mandalori | 2.18.1.2 | OP825090 | OP829143 | OP829058 | OP829056 | Mesobiotus sp. m MM-2022 | temporary taxon name has not been updated in GenBank at time of writing | Erdmann et al. 2024 |
| Mesobiotus mandalori | 22-053 | PQ365769 | PQ367887 | PQ367893 | Mesobiotus sp. m MM-2022 | temporary taxon name has not been updated in GenBank at time of writing | Atherton et al. 2025 | |
| Mesobiotus marmoreus | VN.055.01 | OL311516 | OL257856 | OL257861 | OL257868 | Mesobiotus marmoreus | Stec 2022a | |
| Mesobiotus occultatus | DT97 | OR803042 | OR794157 | OR805249 | OR794158 | Mesobiotus occultatus | Tumanov et al. 2024 | |
| Mesobiotus peterseni | GL.002.01 | OP143859 | OP142528 | OP142516 | OP142522 | Mesobiotus peterseni | Stec 2022a | |
| Mesobiotus philippinicus | KX 129796 | KX129793 | KX129795 | KX129794 | Mesobiotus philippinicus | Mapalo et al. 2016 | ||
| Mesobiotus radiatus | KE.008 | MH195147 | MH197153 | MH197267 | MH197152 | Mesobiotus radiatus | Stec et al. 2018d | |
| Mesobiotus romani | EC.002 | MH195149 | MH197158 | MH197150 | MH197151 | Mesobiotus romani | Roszkowska et al. 2018 | |
| Mesobiotus skanensis | 22-153 | PQ365777 | PQ367889 | PQ367895 | Mesobiotus sp. 2 SA-2024a | temporary taxon name has not been updated in GenBank at time of writing | Atherton et al. 2025 | |
| Mesobiotus skorackii | CN8 | MW656257 | MW680643 | MW680636 | Mesobiotus skorackii | Kayastha et al. 2021 | ||
| Mesobiotus sp. | BI02 | OP684777 | OP696672 | Mesobiotus sp. | Amezcua-Martinez et al. (unpubl.) | |||
| Mesobiotus sp. | NO.234 | MH195153 | MH197148 | MH197156 | MH197265 | Mesobiotus furciger group sp. | Kaczmarek et al. 2018 | |
| Mesobiotus sp. | RU.017 | MH195154 | MH197149 | MH197157 | MH197266 | Mesobiotus harmsworthi group sp. | Kaczmarek et al. 2018 | |
| Mesobiotus sp. 1 | DVT-2020 | MT904513 | MT903468 | MT903470 | MT903612 | Mesobiotus sp. 1 | Tumanov 2020 | |
| Mesobiotus sp. Fl | Mes_sp_S16_3 | OQ968323 | OQ974689 | OQ974696 | OQ974704 | Mesobiotus sp. Fl | Vecchi et al. 2024b | |
| Mesobiotus sp. VN | Mes_sp_ VN 036_04 | OQ968314 | OQ974691 | Mesobiotus sp. VN | Vecchi et al. 2024b | |||
| Mesobiotus vulpinus | DT553 | OR803040 | OR804461 | OR805172 | OR805140 | Mesobiotus sp. 2 DVT-2023b | temporary taxon name has not been updated in GenBank at time of writing | Tumanov et al. 2024 |
| Mesobiotus zelmae | 22-239 | PQ365781 | PQ367891 | PQ367896 | Mesobiotus sp. 3 SA-2024a | temporary taxon name has not been updated in GenBank at time of writing | Atherton et al. 2025 | |
| Mesobiotus cf. intermedius | DT270 | OP013287 | OP035718 | OP035707 | OP035798 | Minibiotus cf. intermedius | Tumanov et al. 2022 | |
| Minibiotus citlalium | AI04 | OP684765 | OP696660 | Minibiotus citlalium | Amezcua-Martinez et al. (unpubl.) | |||
| Minibiotus furcatus | Tar527 | FJ435802 | FJ435746 | FJ435760 | Minibiotus furcatus | Guil & Giribet 2012 | ||
| Minibiotus gumersindoi | Tar710 | FJ435803 | FJ435748 | FJ435761 | Minibiotus gumersindoi | Guil & Giribet 2012 | ||
| Minibiotus intermedius | Min4 GR | ON005160 | ON005189 | ON005195 | Minibiotus intermedius | Kaczmarek et al. 2022 | ||
| Minibiotus ioculator | ZA.274 | MT023412 | MT023998 | MT024000 | MT024041 | Minibiotus ioculator | Stec et al. 2020b | |
| Minibiotus pentannulatus | TZ.027 haplotype 1 MT023413 | MT023999 | MT024001 | MT024042 | Minibiotus pentannulatus | Stec et al. 2020b | ||
| Minibiotus sidereus | AI12 | OP684770 | OP696665 | Minibiotus sidereus | Amezcua-Martinez et al. (unpubl.) | |||
| Minibiotus sp. A MV2020 | S69.Min.01 | MW306859 | OK663227 | OK663216 | OK663238 | Minibiotus sp. A MV2020 | Vecchi et al. 2022b | |
| Paramacrobiotus marchelmoni sp. nov. | 22-004 | PX093656 | PX093665 | PX093646 | PX093651 | COI haplotype 1; 18S haplotype 1; ITS haplotype 2; 28S haplotype 1 | this study | |
| Paramacrobiotus marchelmoni sp. nov. | 22-006 | PX093656 | PX093665 | PX093646 | COI haplotype 1; 18S haplotype 1; ITS haplotype 2 | this study | ||
| Paramacrobiotus marchelmoni sp. nov. | 22-007 | PX093656 | PX093646 | COI haplotype 1; ITS haplotype 2 | this study | |||
| Paramacrobiotus marchelmoni sp. nov. | 22-019 | PX093656 | PX093665 | PX093646 | PX093651 | COI haplotype 1; 18S haplotype 1; ITS haplotype 2; 28S haplotype 1 | this study | |
| Paramacrobiotus marchelmoni sp. nov. | 22-026 | PX093656 | COI haplotype 1 | this study | ||||
| Paramacrobiotus marchelmoni sp. nov. | 22-029 | PX093656 | PX093645 | PX093651 | COI haplotype 1; 18S haplotype 1; 28S haplotype 1 | this study | ||
| Paramacrobiotus marchelmoni sp. nov. | 22-034 | PX093656 | PX093665 | PX093646 | COI haplotype 1; 18S haplotype 1; ITS haplotype 2 | this study | ||
| Paramacrobiotus marchelmoni sp. nov. | 22-102 | PX093656 | PX093645 | COI haplotype 1; ITS haplotype 1 | this study | |||
| Paramacrobiotus marchelmoni sp. nov. | 22-106 | PX093657 | COI haplotype 2 | this study | ||||
| Paramacrobiotus marchelmoni sp. nov. | 22-108 | PX093656 | COI haplotype 1 | this study | ||||
| Paramacrobiotus marchelmoni sp. nov. | 22-110 | PX093657 | PX093665 | PX093646 | PX093651 | COI haplotype 2; 18S haplotype 1; ITS haplotype 2; 28S haplotype 1 | this study | |
| Paramacrobiotus marchelmoni sp. nov. | 22-113 | PX093657 | PX093665 | PX093646 | COI haplotype 2; 18S haplotype 1; ITS haplotype 2 | this study | ||
| Paramacrobiotus marchelmoni sp. nov. | 23-015 | PX093658 | COI haplotype 3 | this study | ||||
| Paramacrobiotus marchelmoni sp. nov. | 23-041 | PX093657 | COI haplotype 2 | this study | ||||
| Paramacrobiotus aff. arduus | PRODO-3 | MK041022 | Paramacrobiotus aff. arduus | Guidetti et al. 2019a | ||||
| Paramacrobiotus aff. spatialis OLBIA-1 | MK041002 | MK041026 | Paramacrobiotus aff. spatialis | Guidetti et al. 2019a | ||||
| Paramacrobiotus arduus | PRODO-1 | MK041020 | MK041032 | Paramacrobiotus arduus | Guidetti et al. 2019a | |||
| Paramacrobiotus arduus | PRODO-2 | MK041021 | Paramacrobiotus arduus | Guidetti et al. 2019a | ||||
| Paramacrobiotus areolatus | NO.385 | MH675998 | MH664931 | MH666080 | MH664948 | Paramacrobiotus areolatus | Stec et al. 2020c | |
| Paramacrobiotus bifrons | C4497_ US 1 | PP236542 | PP240910 | Paramacrobiotus sp. 1 MC-2024a | temporary taxon name has not been updated in GenBank at time of writing | Brandoli et al. 2024 | ||
| Paramacrobiotus bifrons | C4497_ US 2 | PP236543 | PP240911 | Paramacrobiotus sp. 1 MC-2024a | temporary taxon name has not been updated in GenBank at time of writing | Brandoli et al. 2024 | ||
| Paramacrobiotus bifrons | C4497_ US 7 | PP236544 | PP240912 | Paramacrobiotus sp. 1 MC-2024a | temporary taxon name has not been updated in GenBank at time of writing | Brandoli et al. 2024 | ||
| Paramacrobiotus bifrons | C4497_ US 8 | PP236545 | PP240913 | Paramacrobiotus sp. 1 MC-2024a | temporary taxon name has not been updated in GenBank at time of writing | Brandoli et al. 2024 | ||
| Paramacrobiotus bifrons | C5025_ US 6 | PP236551 | PP240918 | Paramacrobiotus sp. 1 MC-2024a | temporary taxon name has not been updated in GenBank at time of writing | Brandoli et al. 2024 | ||
| Paramacrobiotus bifrons | C5062_ US 2 | PP236553 | PP240919 | Paramacrobiotus sp. 1 MC-2024a | temporary taxon name has not been updated in GenBank at time of writing | Brandoli et al. 2024 | ||
| Paramacrobiotus bifrons | C5062_ US 3 | PP236554 | PP240920 | Paramacrobiotus sp. 1 MC-2024a | temporary taxon name has not been updated in GenBank at time of writing | Brandoli et al. 2024 | ||
| Paramacrobiotus bifrons | IT.048 | MH676007 | MH664937 | MH666085 | MH664955 | Paramacrobiotus sp. | see Brandoli et al. 2024 | Stec et al. 2020c |
| Paramacrobiotus bifrons | Par_cf_kly_ S108_2 | OQ968322 | OQ974697 | Paramacrobiotus cf. klymenki | see Brandoli et al. 2024 | Vecchi et al. 2024b | ||
| Paramacrobiotus celsus | PIANE_DI_ MOCOGNO-1 | MK041017 | MK041031 | Paramacrobiotus celsus | Guidetti et al. 2019a | |||
| Paramacrobiotus celsus | PIANE_DI_ MOCOGNO-2 | MK041018 | Paramacrobiotus celsus | Guidetti et al. 2019a | ||||
| Paramacrobiotus celsus | PIANE_DI_ MOCOGNO-3 | MK041019 | Paramacrobiotus celsus | Guidetti et al. 2019a | ||||
| Paramacrobiotus danielae | GY26001 | MZ081363 | MZ081364 | Paramacrobiotus danielae | Wang et al. (unpubl.) | |||
| Paramacrobiotus depressus | ANDALO-1 | MK041015 | Paramacrobiotus depressus | Guidetti et al. 2019a | ||||
| Paramacrobiotus depressus | OSPITALETTO_ A-1 | MK041016 | Paramacrobiotus depressus | Guidetti et al. 2019a | ||||
| Paramacrobiotus depressus | PASSO_ BALLINO-1 | MK041012 | MK041030 | Paramacrobiotus depressus | Guidetti et al. 2019a | |||
| Paramacrobiotus depressus | PASSO_ BALLINO-2 | MK041013 | Paramacrobiotus depressus | Guidetti et al. 2019a | ||||
| Paramacrobiotus depressus | PASSO_ BALLINO-3 | MK041014 | Paramacrobiotus depressus | Guidetti et al. 2019a | ||||
| Paramacrobiotus experimentalis | MAD-TAR11 | MN097837 | MN073464 | MN073465 | Paramacrobiotus experimentalis | Kaczmarek et al. 2020c | ||
| Paramacrobiotus fairbanksi | 22-423 | PX093660 | PX093666 | Paramacrobiotus fairbanksi | this study | |||
| Paramacrobiotus fairbanksi | 23-022 | PX093661 | Paramacrobiotus fairbanksi | this study | ||||
| Paramacrobiotus fairbanksi | 23-041 | PX093659 | PX093666 | PX093647 | PX093648 | Paramacrobiotus fairbanksi | this study | |
| Paramacrobiotus fairbanksi | Alb1 | ON911918 | Paramacrobiotus fairbanksi | Mioduchowsk et al. (unpubl.) | ||||
| Paramacrobiotus fairbanksi | Alb4 | ON911917 | Paramacrobiotus fairbanksi | Mioduchowsk et al. (unpubl.) | ||||
| Paramacrobiotus fairbanksi | C2345-MR_2 | AY598778 | Macrobiotus richtersi | Guidetti et al. 2005 | ||||
| Paramacrobiotus fairbanksi | C2345-MR_3 | AY598779 | Macrobiotus richtersi | Guidetti et al. 2005 | ||||
| Paramacrobiotus sp. | Kenya 3 | EU244599 | Paramacrobiotus sp. ‘ richtersi ’ group | Schill et al. 2010 | ||||
| Paramacrobiotus sp. | MG.002 | MH676008 | MH664938 | MH666086 | MH664956 | Paramacrobiotus sp. | Stec et al. 2020c | |
| Paramacrobiotus sp. | NO.386 | MH676009 | MH664939 | MH666088 | MH664957 | Paramacrobiotus sp. | Stec et al. 2020c | |
| Paramacrobiotus sp. | NZ.001 | MH676010 | MH664940 | MH666089 | MH664958 | Paramacrobiotus sp. | Stec et al. 2020c | |
| Paramacrobiotus sp. | Par_sp_S175_2 | OQ968321 | Paramacrobiotus sp. | Vecchi et al. 2024b | ||||
| Paramacrobiotus sp. | PT.006 | MH676013 | MH664943 | MH666092 | MH664960 | Paramacrobiotus sp. | Stec et al. 2020c | |
| Paramacrobiotus sp. | PT.048 haplotype 1 | MH676014 | MH664944 | MH666093 | MH664961 | Paramacrobiotus sp. | Stec et al. 2020c | |
| Paramacrobiotus sp. | PT.048 haplotype 2 | MH676015 | Paramacrobiotus sp. | Stec et al. 2020c | ||||
| Paramacrobiotus sp. | T3 | OP531839 | ON923868 | Paramacrobiotus sp. | Basu et al. 2023 | |||
| Paramacrobiotus sp. | Tar407 | FJ435807 | FJ435741 | FJ435756 | Macrobiotus pallarii | see Discussion | Guil & Giribet 2012 | |
| Paramacrobiotus sp. | TN.014 | MH676016 | MH664945 | MH666094 | MH664962 | Paramacrobiotus sp. | Stec et al. 2020c | |
| Paramacrobiotus sp. | TZ.018 | MH676017 | MH664933 | MH666095 | MH664951 | Paramacrobiotus sp. | Stec et al. 2020c | |
| Paramacrobiotus sp. DLS-2024a | T82_S135 | PP414782 | PP416751 | PP416753 | PP416752 | Paramacrobiotus sp. DLS-2024a | López-Sandoval et al. 2025 | |
| Paramacrobiotus sp. MAC-2014 | T1 | KF788251 | Paramacrobiotus richtersi group sp. 1 MAC-2014 | Caicedo et al. 2017 | ||||
| Paramacrobiotus sp. MAC-2014 | T2 | KF788252 | Paramacrobiotus richtersi group sp. 1 MAC-2014 | Caicedo et al. 2017 | ||||
| Paramacrobiotus sp. MAC-2014 | T3 | KF788253 | Paramacrobiotus richtersi group sp. 1 MAC-2014 | Caicedo et al. 2017 | ||||
| Paramacrobiotus sp. MAC-2014 | T4 | KF788254 | Paramacrobiotus richtersi group sp. 1 MAC-2014 | Caicedo et al. 2017 | ||||
| Paramacrobiotus sp. MAC-2014 | T6 | KF788255 | Paramacrobiotus richtersi group sp. 1 MAC-2014 | Caicedo et al. 2017 | ||||
| Paramacrobiotus sp. MAC-2014 | T7 | KF788256 | Paramacrobiotus richtersi group sp. 1 MAC-2014 | Caicedo et al. 2017 | ||||
| Paramacrobiotus sp. MAC-2014 | T8 | KF788257 | Paramacrobiotus richtersi group sp. 1 MAC-2014 | Caicedo et al. 2017 | ||||
| Paramacrobiotus sp. ZY-2020 | hp1100404 | MT731035 | MT723894 | Paramacrobiotus sp. ZY-2020 | Yuan et al. (unpubl.) | |||
| Paramacrobiotus sp. ZY-2022c | N 3 | OP870172 | ON819001 | Paramacrobiotus sp. ZY-2022c | Yuan et al. (unpubl.) | |||
| Paramacrobiotus spatialis | FORMIGINE-1 | MK040995 | MK041024 | Paramacrobiotus spatialis | Guidetti et al. 2009 | |||
| Paramacrobiotus spatialis | FORMIGINE-2 | MK040996 | Paramacrobiotus spatialis | Guidetti et al. 2019a | ||||
| Paramacrobiotus spatialis | FORMIGINE-3 | MK040997 | Paramacrobiotus spatialis | Guidetti et al. 2019a | ||||
| Paramacrobiotus spatialis | FORMIGINE-4 | MK040998 | Paramacrobiotus spatialis | Guidetti et al. 2019a | ||||
| Paramacrobiotus spatialis | GAGGIO-1 | MK040999 | Paramacrobiotus spatialis | Guidetti et al. 2019a | ||||
| Paramacrobiotus spatialis | GAGGIO-2 | MK041000 | Paramacrobiotus spatialis | Guidetti et al. 2019a | ||||
| Paramacrobiotus spatialis | GAGGIO-3 | MK041001 | MK041025 | Paramacrobiotus spatialis | Guidetti et al. 2019a | |||
| Paramacrobiotus spatialis | S107.Par | OK662996 | OK663225 | OK663214 | OK663236 | Paramacrobiotus spatialis | Vecchi et al. 2022c | |
| Paramacrobiotus tonollii | US.035 | MH676018 | MH664946 | MH666096 | MH664963 | Paramacrobiotus tonollii | Stec et al. 2020c | |
| Richtersius coronifer | NO.385 | MH676053 | MH681760 | MH681763 | MH681757 | Richtersius coronifer | Stec et al. 2020c | |
| Sisubiotus hakaiensis | Sis.hak.S1911 | OM523181 | OM523054 | OM523057 | OM523059 | Sisubiotus hakaiensis | Vecchi et al. 2022a | |
| Sisubiotus spectabilis | FI.067 | MN888322 | MN888371 | MN888331 | MN888357 | Sisubiotus spectabilis | Stec et al. 2021a | |
| Tenuibiotus cf. ciprianoi | ES.086 | MN888328 | MN888376 | MN888348 | MN888361 | Tenuibiotus cf. ciprianoi | Stec et al. 2021a | |
| Tenuibiotus danilovi | KG.128 | MN888329 | MN888377 | MN888349 | MN888362 | Tenuibiotus danilovi | Stec et al. 2021a | |
| Tenuibiotus sp. | 22-109 | PX093662 | PX093667 | PX093652 | Tenuibiotus sp. | this study | ||
| Tenuibiotus tenuiformis | KG.140 | MN888330 | MN888378 | MN888350 | MN888363 | Tenuibiotus tenuiformis | Stec et al. 2021a | |
| Tenuibiotus voronkovi | haplotype 1 | KX810042 | KX810045 | KX810046 | KX810049 | Tenuibiotus voronkovi | Zawierucha et al. 2016 | |
| Tenuibiotus yeliseii | DT422 | OR145334 | OR142418 | OR142424 | OR142426 | Tenuibiotus sp. 1 DVT-2023a | temporary taxon name has not been updated in GenBank at time of writing | Tsvetkova & Tumanov 2024 |
| Tenuibiotus zandrae | MN 444827 | MN443040 | MN443038 | MN443035 | Tenuibiotus zandrae | Stec et al. 2020c | ||
| Xerobiotus arenosum | C4411_V4 | OR397039 | OR398024 | OR398050 | Xerobiotus arenosum | Vincenzi et al. 2024 | ||
| Xerobiotus cf. reductus | PL3177_1 | OR397050 | OR398033 | OR398056 | Xerobiotus cf. reductus | Vincenzi et al. 2024 | ||
| Xerobiotus euxinus | check 1.4 | PP469626 | PP439986 | PP470593 | PP440001 | Macrobiotus euxinus | see Vincenzi et al. 2024 | Polishchuk et al. 2024 |
| Xerobiotus euxinus | C3834_V4 | OR397005 | OR398005 | OR398119 | Macrobiotus euxinus | see Vincenzi et al. 2024 | Vincenzi et al. 2024 | |
| Xerobiotus gretae | C4341 G | MW581665 | MW588434 | MW588431 | MW588438 | Macrobiotus gretae | see Vincenzi et al. 2024 | Massa et al. 2021 |
| Xerobiotus litus | C4383_2 | OR397000 | OR398002 | OR398118 | Xerobiotus litus | Vincenzi et al. 2024 | ||
| Xerobiotus naginae | S226.Mac.1 | OK662990 | OK663219 | OK663230 | Marobiotus naginae | see Vincenzi et al. 2024 | Vecchi et al. 2022c | |
| Xerobiotus pseudohufelandi | C2358-XP1 | AY598776 | HQ604989 | Xerobiotus pseudohufelandi | Guidetti et al. 2005 | |||
| Xerobiotus reductus | PL_3176_1 | OR397048 | OR398028 | OR398130 | Xerobiotus reductus | Vincenzi et al. 2024 | ||
| Xerobiotus sp. | PL.360 | MN888325 | MN888373 | MN888345 | MN888358 | Xerobiotus sp. | Vecchi & Stec 2021 | |
| Xerobiotus sp. | ZA.373 | MN888326 | MN888374 | MN888346 | MN888359 | Xerobiotus sp. | Vecchi & Stec 2021 | |
| Xerobiotus sp. 1 MC-2023a | C3306_V1 | OR397025 | OR398020 | OR398123 | Xerobiotus sp. 1 MC-2023a | Vincenzi et al. 2024 | ||
Notes
Published as part of Hulterström, Jens, Guidetti, Roberto, Jönsson, K. Ingemar & Atherton, Sarah, 2025, Integrative taxonomy supports the description of two new species of Macrobiotidae (Tardigrada, Eutardigrada) from Kristianstads Vattenrike Biosphere Reserve in Sweden, pp. 38-52 in European Journal of Taxonomy 1030 on page 38, DOI: 10.5852/ejt.2025.1030.3135, http://zenodo.org/record/17962351
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