Scytodes joniae sp. nov.

Figs 71–78, 95–98, 136

Type material. Holotype: ♂, DOMINICAN REPUBLIC: La Altagracia Province: Occidental, San Rafael de Yuma, Cueva de Linea, Parque Nacional los Haitises, 19.07796ºN, 69.46355ºW, 7 mts., 12 June 2012, Team CarBio leg., USNM782341, DR008 (NMNH). Paratypes: DOMINICAN REPUBLIC: La Altagracia Province: 1 ♀, same collection data as for holotype, USNM785538 (NMNH); 1 ♀, same collection locality as for previous specimen, Cueva La Arena, 19.0800ºN, 69.4650ºW, 17 mts., 12 June 2012, Team CarBio leg., USNM782490, DR007 (NMNH); 1 ♀, same collection data as for previous specimen, USNM785252 (NMNH); 1 ♀, same collection data as for previous specimen, USNM782496 (NMNH); 1 ♀, same collection data as for previous specimen, USNM784365 (NMNH); 1 ♀, same collection data as for previous specimen, USNM785799 (NMNH).

Etymology. The specific name is a matronym honoring the Canadian singer-songwriter Joni Mitchell, known for her influential work in folk, jazz, and pop music, and for her complex compositions and poetic lyrics.

Diagnosis. Males of S. joniae sp. nov. resemble those of S. cubensis (Figs 16–18, 61, 62), S. mingusi sp. nov. (Figs 89, 90, 100, 101), S. noeli (Alayón, 1992: fig. 3), S. reinhardti sp. nov. (Figs 112, 113, 128, 129) and S. robertoi (Alayón, 1977: fig. 2C) by the palp with a piriform bulb and a slender, tapering embolus. They are distinguished from S. robertoi by the bulb with a median constriction (vs. lacking in S. robertoi); from S. mingusi sp. nov. by the filiform embolus (vs. thicker at base in S. mingusi sp. nov.); from S. noeli by the straight embolus (vs. curved in S. noeli); and from S. reinhardti sp. nov. by the shorter cymbium, roughly 1.5 times tibia length, and bulb, roughly as long as embolus (vs. cymbium with more than 1.8 times tibia length and bulb 1.6 times longer than embolus in S. reinhardti sp. nov.). Females resemble those of S. coltrainei sp. nov. (Figs 12, 13, 36), S. cubensis (Figs 21, 22, 63–70), and S. jimmorrisoni sp. nov. (Figs 59, 60, 99) by the vulva with S-shaped IS. They are distinguished from S. coltrainei sp. nov. by the shorter IS, approximately three times longer than the OS (Figs 78, 95, 96) (vs. more than four times longer in S. coltrainei sp. nov.); from S. cubensis by the PS simple and untwisted (vs. twisted and coiled in S. cubensis); and from S. jimmorrisoni sp. nov. by the IS with the same with throughout its entire length (vs. widened at base in S. jimmorrisoni sp. nov.).

Description. Male (USNM782341, holotype): prosoma pale golden yellow with brown markings; chelicerae pale yellow; legs pale yellow with brown markings; labium grayish brown; endites pale yellow with brown outlines; sternum pale yellow with brown dotted markings; opisthosoma grayish yellow with scattered brown markings (Figs 71, 72). Total length 3.8; prosoma: 2.0 long, 1.6 wide; sternum: 1.1 long, 0.8 wide; labium: 0.22 long, 0.25 wide; opisthosoma: 1.7 long, 1.3 wide; eye diameters: PME 0.15, ALE.014, PLE 0.13; legs: I – (3.8, rest of leg absent); all other legs absent. Palp: cymbium with two apical spines; embolus long and filiform, slightly curving at tip (Figs 73, 74, 97, 98). Measurements: tibia 0.39; cymbium 0.67; bulb 0.68.

Female (paratype, USNM785252): prosoma golden brown, with dark brown markings; chelicerae pale yellow with brown margins; legs golden yellow with brown ring-marks; labium dark brown with two oval-shaped paleyellow spots; endites dark brown with golden yellow bases; sternum golden yellow covered with brown markings; opisthosoma grayish yellow with scattered brown markings (Figs 75, 76). Measurements: total length 6.3; prosoma: 2.4 long, 1.9 wide; sternum: 1.1 long, 0.9 wide; labium: 0.28 long, 0.3 wide; opisthosoma: 3.9 long, 2.0 wide; eye diameters: PME 0.11, ALE 0.11, PLE 0.10; legs: I: 9.6 (3.0, 0.5, 3.0, 2.15, 0.95); II: − (2.3, 0.5, 2.25, 2.2, absent); III: 5.85 (1.6, 0.5, 1.55, 1.5, 0.7); IV: 8.1 (2.3, 0.5, 2.25, 2.3, 0.75). Female genitalia: positioning ridges postero-laterad, hood-like, four times wider than long, separated from each other by twice their width (Fig. 77). Vulva: IS slightly widened at tip; OS roughly five times longer than wide, with the same width throughout its length and apically rounded; antero-laterad; PS with mediad loop (Fig. 78).

Variation. Females (n=6): total length 4.8–6.3; prosoma length 2.3–2.6; femur I length 2.9–3.0.

Distribution. Known from Parque Nacional los Haitises, Dominican Republic (Fig. 136).