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Published November 19, 2025 | Version v1
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Four new species of Cleidogona Cook & Collins, 1895 (Diplopoda: Chordeumatida: Cleidogonidae) from the Great Smoky Mountains National Park, USA

Authors/Creators

  • 1. Department of Biological and Environmental Sciences, Georgia College & State University, Milledgeville, GA, USA 31061
  • 2. Professor Emeritus, Department of Biology, Hampden-Sydney College, Hampden-Sydney VA 23943 USA, current address: 1950 Price Drive, Farmville, VA 23901 USA

Description

Snyder, Bruce A., Shear, William A. (2025): Four new species of Cleidogona Cook & Collins, 1895 (Diplopoda: Chordeumatida: Cleidogonidae) from the Great Smoky Mountains National Park, USA. Zootaxa 5722 (2): 249-264, DOI: 10.11646/zootaxa.5722.2.5, URL: https://doi.org/10.11646/zootaxa.5722.2.5

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Additional details

Identifiers

URL
https://www.checklistbank.org/dataset/313986
LSID
urn:lsid:zoobank.org:pub:CEF8EFA8-D23F-45B3-82F9-36B61E345EBE
LSID
urn:lsid:plazi.org:pub:FFE4EE01FFD9FFA6FF8F43345907A146
URL
http://publication.plazi.org/id/FFE4EE01FFD9FFA6FF8F43345907A146

Cites
Publication: 10.5962/bhl.part.18578 (DOI)
Publication: 10.1111/j.1749-6632.1896.tb55430.x (DOI)
Publication: 10.1656/1528-7092(2007)6[215:RADIFA]2.0.CO;2 (DOI)
Publication: 10.5962/bhl.title.3182 (DOI)
Publication: 10.3897/zookeys.1166.103516 (DOI)
Publication: 10.11646/zootaxa.5397.2.9 (DOI)
Publication: 10.2307/3494274 (DOI)
Publication: 10.11646/zootaxa.1856.1.2 (DOI)
Publication: 10.3157/021.121.0103 (DOI)
Publication: 10.1007/s10530-010-9826-4 (DOI)
Publication: 10.2307/1005211 (DOI)
Has part
Taxonomic treatment: 10.5281/zenodo.17892820 (DOI)
Taxonomic treatment: http://treatment.plazi.org/id/03DD9679FFDBFFA4FF18409A5B5AA43F (URL)
Taxonomic treatment: 10.5281/zenodo.17892822 (DOI)
Taxonomic treatment: http://treatment.plazi.org/id/03DD9679FFDBFFA5FF1846FE5870A2AB (URL)
Taxonomic treatment: 10.5281/zenodo.18406558 (DOI)
Taxonomic treatment: http://treatment.plazi.org/id/03DD9679FFDAFFA5FF18470A5B11A4FD (URL)
Taxonomic treatment: 10.5281/zenodo.17892824 (DOI)
Taxonomic treatment: http://treatment.plazi.org/id/03DD9679FFDAFFA3FF1845395C7DA477 (URL)
Taxonomic treatment: 10.5281/zenodo.17892826 (DOI)
Taxonomic treatment: http://treatment.plazi.org/id/03DD9679FFDCFFAEFF1846B65879A4DB (URL)
Taxonomic treatment: 10.5281/zenodo.17892828 (DOI)
Taxonomic treatment: http://treatment.plazi.org/id/03DD9679FFD1FFACFF1846DA5A8CA51F (URL)
Taxonomic treatment: 10.5281/zenodo.17892832 (DOI)
Taxonomic treatment: http://treatment.plazi.org/id/03DD9679FFD3FFAAFF18479E5ADEA717 (URL)
Figure: 10.5281/zenodo.17892045 (DOI)
Figure: 10.5281/zenodo.17892051 (DOI)
Figure: 10.5281/zenodo.17892055 (DOI)
Figure: 10.5281/zenodo.17892059 (DOI)
Dataset: http://table.plazi.org/id/DF0B77E7FFD4FFABFF1841F95AB5A24D (URL)
Figure: 10.5281/zenodo.17892061 (DOI)
Is source of
Dataset: https://www.gbif.org/dataset/5c339d72-8393-4a0c-9c4d-08fe8a023284 (URL)

References

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  • Chamberlin, R.V. (1928) Some chilopods and diplopods from Missouri. Entomological News, 39, 153-155.
  • Cook, O.F. (1896a) Note on the classification of the Diplopoda. American Naturalist, 30, 681-684.
  • Cook, O.F. (1896b) On recent diplopod names. Brandtia, 2, 5-8.
  • Cook, O.F. & Collins, G.N. (1895) The Craspedosomatidae of North America. Annals of the New York Academy of Science, 9, 1-100. https://doi.org/10.1111/j.1749-6632.1896.tb55430.x
  • Cope, E.D. (1869) Synopsis of the extinct Mammalia…with observations on some Myriapoda. Proceedings of the American Philosophical Society, 11, 171-192.
  • Hoffman, R.L. (1950a) Records and descriptions of Diplopods from the southern Appalachians. Journal of the Elisha Mitchell Scientific Society, 66, 11-33.
  • Hoffman, R.L. (1950b) A preliminary list of the cleidogonid millipeds, with descriptions of a new genus from Guatemala and a new species from Virginia. Journal of the Washington Academy of Sciences, 40, 87-92.
  • Lessard, J-P., Dunn, R.R., Parker, C.R. & Sanders, N.J. (2007) Rarity and diversity in forest ant assemblages of Great Smoky Mountains National Park. Southeastern Naturalist, 6 (Sp2), 215-228. https://doi.org/10.1656/1528-7092(2007)6[215:RADIFA]2.0.CO;2
  • Loomis, H.F. (1964) The millipeds of Panama (Diplopoda). Fieldiana, Zoology, 47 (1), 1-134. https://doi.org/10.5962/bhl.title.3182
  • Olliff, M. (2021) Soil macroinvertebrate responses to wildfires in the Blue Ridge Mountains, USA. MS Thesis, Georgia College & State University, Milledgeville, Georgia, 59 pp.
  • Parker, C. & Bernard, E. (2006) The science approach to the Smokies ATBI. The George Wright Forum, 23, 26-36.
  • QGIS Development Team. (2025) QGIS geographic information system. QGIS Association, Lyon. [program]
  • Recuero, E. & Caterino, M.S. (2023) A second species of the pill millipede genus Nearctomeris Wesener, 2012 (Diplopoda, Glomerida) from the Great Smoky Mountains, USA. ZooKeys, 1166, 333-349. https://doi.org/10.3897/zookeys.1166.103516
  • Schubart, O. (1953) Um novo genero de Ascospermophora de Portugal (Diplopoda, Nematophora). Publicacoes do Instituto de Zoologia "Dr. Augusto Nobre" Faculdade de Ciencias do Porto, 45, 80-90.
  • Shear, W.A. (1971) The milliped family Conotylidae in North America, with a Description of the new family Adritylidae (Diplopoda, Chordeumida). Bulletin of the Museum of Comparative Zoology, 141 (2), 55-97.
  • Shear, W.A. (1972) Studies in the milliped order Chordeumida (Diplopoda): A revision of the family Cleidogonidae and a reclassification of the order Chordeumida in the New World. Bulletin of the Museum of Comparative Zoology, 144, 151- 352.
  • Shear, W.A. (1982) Cabraca unigon, n gen., n. sp., a remarkable new cleidogonid milliped from Mexico (Chordeumatida: Cleidogonidae). Myriapodologica, 1 (10), 1-6.
  • Shear, W.A. (2024) Lineagona, a new millipede genus (Diplopoda, Chordeumatida, Cleidogonidae) from Texas, USA, and Nuevo Leon, Mexico. Zootaxa, 5397 (2), 283-290. https://doi.org/10.11646/zootaxa.5397.2.9
  • Shelley, R.M. (1976) A new Diplopod of the genus Cleidogona from North Carolina (Chordeumida: Cleidogonidae). Florida Entomologist, 58 (3), 325-327. https://doi.org/10.2307/3494274
  • Sierwald, P., Decker, P. & Spelda, J. (2025) MilliBase. Cleidogona Cook & Collins, 1895. Available from: https://www. millibase.org/aphia.php/1zqjmv'%22()%7B%7D%3Cx%3E:1zqjmv%3B9/aphia.php?p=taxdetails&id=891296 (accessed 7 June 2025)
  • Snyder, B.A. (2008) A preliminary checklist of the millipedes (Diplopoda) of the Great Smoky Mountains National Park, USA. Zootaxa, 1856 (1), 16-32. https://doi.org/10.11646/zootaxa.1856.1.2
  • Snyder, B.A. & Seewagen, C. (2010) First record of the millipede Cleidogona nantahala Shear, 1972 (Chordeumatida: Cleidogonidae) for New York State, USA. Entomological News, 121, 104-105. https://doi.org/10.3157/021.121.0103
  • Snyder, B.A., Callaham Jr., M.A. & Hendrix, P.F. (2011) Spatial variability of an invasive earthworm (Amynthas agrestis) population and potential impacts on soil characteristics and millipedes in the Great Smoky Mountains National Park, USA. Biological Invasions, 13, 349-358. https://doi.org/10.1007/s10530-010-9826-4
  • Straube, D. (2013) Development and application of molecular techniques to assess feeding interactions in a forest invaded by the Asian earthworm Amynthas agrestis. Doctoral Thesis, University of Innsbruck, Innsbruck, 102 pp.
  • Verhoeff, K.W. (1899) Beitrage zur Kenntniss palaarktischer Myriopoden. VIII. Aufsatz: Zur vergleichenden Morphologie, Phylogenie, Gruppen- und Art-Systematik der Chordeumiden. Archiv fur Naturgeschichte, 65 (1), 95-154.
  • Verhoeff, K.W. (1914) Ascospermophora aus Japan (Uber Diplopoden 69. Aufsatz). Zoologischer Anzeiger, 43, 342-370.
  • Wood, H.C. (1865) The Myriapoda of North America. Transactions of the American Philosophical Society, 13, 137-248. https://doi.org/10.2307/1005211