Pterostichus (Cryobius) kirilli Sundukov, sp. nov.

(Figs 6–10)

Type locality. RUSSIA, Khabarovsky Krai, Badzhal Ridge, source of the Omot-Makit River, 50.505559 N, 134.273428 E.

Type material. Holotype, ♂ with labels: “ Khabarovsky Krai, Badzhal Ridge, source of the Omot-Makit River, 1800–2100 m, 7–19.VII.1997, leg. Yu. Sundukov (in Russian); “HOLOTYPUS Pterostichus (Cryobius) kirilli sp. n. det. Yu. Sundukov 2025” [FEB]. Paratypes, 5♂, 5♀, “ Khabarovsky Krai, Badzhal Ridge, source of the Omot-Makit River, 1800–2100 m, 7–19.VII.1997, leg. Yu. Sundukov (in Russian); “PARATYPUS Pterostichus (Cryobius) kirilli sp. n. det. Yu. Sundukov 2025” [FEB].

Description. Body slightly convex. Length 7.00– 7.70 mm, width 2.35–2.65 mm.

Black; protarsi, palps, mandibles and partially antennae reddish.

Dorsal side of head almost smooth, with very weak isodiametric meshes on the occiput; pronotum almost glossy, with areas of very weak transverse meshes; elytra with weak short transverse meshes, more distinct in females.

Measurements: HW 1.33–1.40 (M 1.36); HL 0.90–0.95 (M 0.94); PA 1.28–1.38 (M 1.33); PW 1.78–1.93 (M 1.87); PB 1.30–1.43 (M 1.36); PLt 1.50–1.75 (M 1.64); PLm 1.43–1.65 (M 1.54); EW 2.35–2.65 (M 2.54); EL 3.80–4.38 (M 4.14); Ls 6.20–7.00 (M 6.73); L 7.00–7.70 (M 7.49).

Head small, convex, rather short, HW/HL 1.42–1.47, M 1.45. Eyes small, convex. Temples rather long, more than 1/2 longitudinal diameter of eye. Frontal grooves short, moderately deep, widely groove-shaped, almost parallel, starting from anterior supraorbital seta, indistinct on clypeus. Frontoclypeal suture thin, distinct or weak. Clypeus trapezoidal, with slightly concave anterior margin and single seta on each side. Labrum transverse, with 6 setae along anterior margin. Mandibles short, strong. Tooth of mentum short, a half as long as lateral lobes, wide, truncated-concave at apex. Antennae relatively thin, moderately long, in males they extend beyond the base of pronotum by 3 apical segments, in females—by 2.5 segments.

Pronotum cordate or slightly elongate-cordate, relatively small (PW/HW 1.34–1.40, M 1.37; EW/PW 1.32– 1.40, M 1.36), slightly wider than long (PW/PLt 1.08–1.18, M 1.14), with maximum width 3/5 from base. Anterior margin barely concave, anterior angles barely protruding, rounded at apex. Sides rather strongly or moderately convex, moderately or slightly concave before posterior angles. Base narrow (PW/PB 1.32–1.43, M 1.38; PB/PA 1.02–1.04, M 1.02), rectilinear or very slightly concave. Posterior angles acute, pointed at apex. Disc slightly or moderately convex to narrow lateral edges. Median line thin, distinct. Anterior transverse impression distinct, deep. Two basal foveae on each side: inner deep, wide, parallel or slightly arcuate, running on about basal 1/2–2/5, outer small, separated from inner by thin, slightly raised keel. Puncturation present only in basal foveae. Two setae on each side: in front of maximum width and in posterior angles (holotype has two anterior setae on left side of pronotum).

Elytra moderately convex, elongated-oval (EL/EW 1.60–1.68, M 1.63; EL/PLt 2.44–2.61, M 2.52), widest two thirds from base. Disc uniformly, moderately or weakly convex. Humeri broadly rounded, not protruding, with small tooth; basal ridge rectilinear or weakly concave. Striae complete, straight, moderately deep, indistinctly punctate. Intervals 1–5 moderately convex, outer ones flattened. Parascutellar striole absent or barely indicated by weak stroke at apex of 2nd interval. Parascutellar setigerous pores present—at base of stria 2 or in 2nd interval. 3rd interval with 2 small discal pores in apical half of elytron, located at 2nd stria; anterior pores absent. Umbilical seta series consisting of 10–11 setae: 6 in humeral group and 4–5 at apex. Preapical notches very weak. Apterous.

Legs long and slender. Middle and hind femora with 2 setae along posterior margin. Hind coxae with 3 setae. Trochanters with 1 seta. Metepisterum short: 1.10–1.15 times longer along outer margin than width along anterior margin. Tarsomere 5 with thin ventral setae in apical half.

Pro-, meso-, metepisternum, mesoventrite/mesosternum and sides of 1st and 2nd visible abdominal sternites finely punctate. Intercoxal process of prosternum not bordered, slightly pointed at apex. Anal sternite in male with 2 setae at apex, in female—with 4 setae.

Aedeagus (Figs 7–10). Median lobe rather slender, moderately curved ventrally in basal part and weakly before apex. Apical lamella short, with rounded apex; its length about 3 times less than diameter of aedeagus in basal part.

Etymology. The name given after famous Russian entomologist Kirill Vladimirovich Makarov.

Distribution. Known only from the type locality.

Habitat. The beetles live in the alpine zone: on rocky watersheds and northern slopes, as well as among stones at streamheads. They live together with P. badzhaliensis sp. nov.

Differential diagnosis. Pterostichus kirilli sp. nov. is closest to the arctic-boreal P. longipes Poppius, 1906, different in the predominantly red or blackish-red coloration of the body appendages, the sides of the pronotum rectilinear and short before the posterior angles, longer and narrower meshes of the elytral microsculpture, impunctate striae, and the absence of parascutellar setigerous pores of the elytra in the Far Eastern specimens.