Published December 4, 2025 | Version v1

Neusticosaurus pusillus

  • 1. Department of Paleobiology, National Museum of Natural History, MRC 121, Smithsonian Institution, Washington, DC 20560, USA
  • 2. Staatliches Museum für Naturkunde Stuttgart, Rosenstein 1, 70191 Stuttgart, Germany & Institut für Biologie, Universität Hohenheim, Garbenstrasse 30, 70599 Stuttgart, Germany

Description

Neusticosaurus pusillus (E. Fraas, 1881)

Holotype.

NHMUK PV R 53, nearly complete skeleton.

Type locality.

Kugelberg at Ludwigsburg-Hoheneck, Stuttgart district, Baden-Württemberg.

Type horizon.

Hoheneck-Kalk, Erfurt Formation, Lower Keuper Subgroup. Age: Middle Triassic (Ladinian: Longobardian).

Referred material.

For a detailed list see Sander (1989). Sander provided a thorough description of this species based primarily on numerous excellently preserved skeletons from the Ladinian-age Cava Inferiore Beds of the Meride Limestone at Monte San Giorgio (Southern Alps of Switzerland).

Diagnosis.

Distinguished by the following combination of features: small size (total body length of up to 52 cm); skull table with parallel lateral margins; orbit proportionately small; supratemporal fossa oval; 22–24 dorsal vertebrae; atlas lacking ribs; and two carpal bones present in adults (Sander 1989).

Comments.

Sander (1989) considered Neusticosaurus pygmaeus E. Fraas, 1896, based on SMNS 53932, a group of three complete and nine partial skeletons from the Hoheneck-Kalk (Fig. 12 A), a subjective junior synonym of N. pusillus.

References.

Seeley (1882), E. Fraas (1896), Sander (1989), Rieppel (2000).

Notes

Published as part of Sues, Hans-Dieter & Schoch, Rainer R., 2025, Synopsis of the Triassic reptiles from Germany, pp. 411-483 in Fossil Record 28 (2) on pages 411-483, DOI: 10.3897/fr.28.164405

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Linked records

Additional details

Biodiversity

Collection code
NHMUK, PV
Material sample ID
R 53
Scientific name authorship
E. Fraas
Kingdom
Animalia
Phylum
Chordata
Order
Eosauropterygia
Family
Pachypleurosauridae
Genus
Neusticosaurus
Species
pusillus
Taxon rank
species
Type status
holotype
Taxonomic concept label
Neusticosaurus pusillus (Fraas, 1881) sec. Sues & Schoch, 2025

References

  • Fraas O (1881) Simosaurus pusillus aus der Lettenkohle von Hoheneck. Jahreshefte des Vereins fur vaterlandische Naturkunde 37: 319-324.
  • Fraas E (1896) Die schwa ¨ bischen Trias-Saurier, nach dem Material der Kgl. Naturalien-Sammlung in Stuttgart zusammengestellt. E. Schweizerbart'sche Verlagsbuchhandlung, Stuttgart, 1 - 18.
  • Seeley HG (1882) On Neusticosaurus pusillus (Fraas), an amphibious reptile having affinities with the terrestrial Nothosauria and with the marine Plesiosauria. Quarterly Journal of the Geological Society of London 38: 350–366. https://doi.org/10.1144/GSL.JGS.1882.038.01-04.39
  • Fraas E (1896) Die schwa ̈ bischen Trias-Saurier, nach dem Material der Kgl. Naturalien-Sammlung in Stuttgart zusammengestellt. E. Schweizerbart'sche Verlagsbuchhandlung, Stuttgart, 1–18.
  • Rieppel O (2000) Sauropterygia I. Placodontia, Pachypleurosauria, Nothosauroidea, Pistosauroidea. Handbuch der Paläoherpetologie Part 12 A. Verlag Dr. Friedrich Pfeil, Munich, 1–134.