Published November 12, 2025 | Version v1

Capoeta coadi Alwan, Zareian & Esmaeili 2016

Description

Capoeta coadi

Common name. Karun scraper.

Diagnosis: Distinguished from other species of Capoeta in Karkheh, Karun, Zohreh and endorheic Zayandeh drainages by: ○ usually 9½ branched dorsal rays / ○ one pair of barbels / ○ flank beige, golden or brown without small black spots, juveniles and some adults individuals with large black blotches / ○ last unbranched dorsal ray weakly ossified and serrated / ○ 14–18 gill rakers, 10–13 gill rakers on lower limb / ○ 65–84 total scales along lateral line / ○ 12–17 scales between lateral line and dorsal origin / ○ 9–11 scales between lateral line and pelvic origin / ○ 25–32 circumpeduncular scales / ○ a deep groove in front of nares. Size up to 280 mm SL.

Distribution. Iran: Karun drainage and Esfahan basin.

Habitat. Moderate or fast-flowing rivers, usually with gravel substrate and clear water.

Biology. Lives up to 10 years. Males mature at 2 years, females at 3−6 years. Spawns May−June. Feeds on periphyton.

Conservation status. LC.

Remarks. Capoeta birunii from the Zayandeh drainage is a synonym.

Further reading. Alwan 2010 (morphology, phylogeny); Alwan et al. 2016a (description); Jouladeh-Roudbar et al. 2016a (phylogeny); Jouladeh-Roudbar et al. 2017b (phylogeny, morphology); Zareian & Esmaeili 2017 (description of C. birunii); Jouladeh-Roudbar et al. 2020 (distribution, C. birunii as a synonym of C. coadi); Coad 2021a (biology, morphology); Jouladeh-Roudbar 2022 (C. birunii as a synonym of C. coadi).

Notes

Published as part of Freyhof, Jörg, Yoğurtçuoğlu, Baran, Jouladeh-Roudbar, Arash & Kaya, Cüneyt, 2025, Handbook of Freshwater Fishes of West Asia, GmbH, Berlin / Boston :De Gruyter on page 126, DOI: 10.1515/9783111677811, http://zenodo.org/record/17881367

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Additional details

References

  • Alwan, N. 2010. Systematics, taxonomy, phylogeny and zoogeography of the Capoeta damascina species complex (Pisces: Teleostei: Cyprinidae) inferred from comparative morphology and molecular markers. PhD Thesis, Johann Wolfgang Goethe-Universitat, Frankfurt.
  • Alwan, N. H., H. Zareian & H. R. Esmaeili. 2016 a. Capoeta coadi, a new species of cyprinid fish from the Karun River drainage, Iran based on morphological and molecular evidences (Teleostei, Cyprinidae). ZooKeys 572: 155-180. https://doi.org/10.3897/zookeys.572.7377
  • Jouladeh-Roudbar, A., S. Eagderi, H. R. Ghanavi & I. Doadrio. 2016 a. Taxonomic review of the genus Capoeta Valenciennes, 1842 (Actinopterygii, Cyprinidae) from central Iran with the description of a new species. FishTaxa 1: 166-175.
  • Jouladeh-Roudbar, A., S. Eagderi, H. R. Ghanavi & I. Doadrio. 2017 b. A new species of the genus Capoeta Valenciennes, 1842 from the Caspian Sea basin in Iran (Teleostei, Cyprinidae). ZooKeys 682: 137-155. https://doi.org/10.3897/zookeys.682.12670
  • Zareian, H. & H. R. Esmaeili. 2017. Mitochondrial phylogeny and taxonomic status of the Capoeta damascina species group (Actinopterygii: Cyprinidae) in Iran with description of a new species. Iranian Journal of Ichthyology 4: 231-269. https://doi.org/10.22034/iji.v4i3.239
  • Jouladeh-Roudbar, A., H. R. Ghanavi & I. Doadrio. 2020. Ichthyofauna From Iranian Freshwater: Annotated checklist, diagnosis, taxonomy, distribution and conservation assessment. Zoological Studies 59: e 21. https://doi.org/10.6620/ZS.2020.59-21
  • Coad, B. W. 2021 a. Carps and Minnows of Iran (Families Cyprinidae and Leuciscidae) Volume I: General introduction and Carps (Family Cyprinidae). Canadian Museum of Nature, Ottawa, Ontario, Canada, 1 - 1263. Available at: http://www.briancoad.com/species%20accounts/Carps%20of%20Iran%2010Sept2opt1.pdf (Accessed January 2023).
  • Jouladeh-Roudbar, A. 2022. Molecular phylogeny of Capoeta Valenciennes, 1842 (Teleostei, Cyprinidae) species in Iran, using the cytochrome b gene. Modern Genetic journal 17: 171-183. [in Farsi, English summary].