Fish-life in the Late Miocene Lake Pannon of the Vienna Basin – a review
Description
The long-lived brackish Late Miocene (Pannonian) Lake Pannon formed a large endorheic lake in Europe known for its explosive endemic speciation in mollusks. Unlike the many decades of intense research of fossil mollusks in Lake Pannon, relatively little research has been conducted into its fish fauna including otoliths. Here, I review the Pannonian otoliths from the westernmost outlier of Lake Pannon, the Vienna Basin in Austria, and some smaller samples from the Danubian and Pannonian basins in Hungary. In contrast to the richness and rapid adaptive evolution of mollusk taxa during the Pannonian, the otolith-based fish fauna is relatively lean in species, with 29 species including four new species and four in open nomenclature. Otoliths were found in brackish lake and fluviatile sediments, and freshwater lake deposits away from Lake Pannon. Sixteen species represent genuine lacustrine fishes, with the remainder stemming from freshwater environments or having been transported from freshwater environments into Lake Pannon. Half of the lake fish species are endemic to Lake Pannon. This compares to 36 otolith-based species identified in time-equivalent deposits in the restricted marine Eastern Paratethys, which were almost 100% endemic. The other 50% of the Lake Pannon fishes have roots in the restricted marine Middle Miocene Central Paratethys that preceded the Lake Pannon phase. Unlike mollusks, therefore, the fish fauna of Lake Pannon is characterized furthermore by a relative stasis in evolution with relatively long-ranging species. The paleoenvironmental affinities of the fishes identified from otoliths is also discussed. The new taxa described are Pannonigadus gen. nov. (Gadidae), Gadiculus weinfurteri sp. nov. (Gadidae), Eleotris pannonicus sp. nov. (Eleotridae), Ponticola planodorsalis sp. nov., and Toxopyge bradicae sp. nov. (both Gobiidae).
Files
ANHMW_article_168691.pdf
Files
(15.7 MB)
| Name | Size | Download all |
|---|---|---|
|
md5:11c1830c40a48177351c51b6e27305fb
|
15.7 MB | Preview Download |
System files
(373.8 kB)
| Name | Size | Download all |
|---|---|---|
|
md5:d9a75c1baca42a322d54300527a5337a
|
373.8 kB | Download |
Linked records
Additional details
References
- Assis CA (2005) The utricular otoliths, lapilli, of teleosts: their morphology and relevance for species identification and systematic studies. Scientia Marina 69(2): 259–273. https://doi.org/10.3989/scimar.2005.69n2259
- Bannikov AF, Schwarzhans W, Carnevale G (2018) Neogene Paratethyan croakers (Teleostei, Sciaenidae). Rivista Italiana di Paleontologia e Stratigrafia 124(3): 535–571. https://doi.org/10.13130/2039-4942/10696
- Böhme M (2002) Freshwater fishes from the Pannonian of the Vienna Basin with special reference to the locality Sandberg near Götzendorf, Lower Austria. Courier Forschungsinstitut Senckenberg 237: 151–173.
- Böhme M (2004) Migration history of air-breathing fishes reveals Neogene atmospheric circulation patterns. Geology 32(5): 393–396. https://doi.org/10.1130/G20316.1
- Bosnakoff M, Katona LT (2012) A fish otolith assemblage from the Late Miocene (Pannonian) deposits of Lake Pannon (Doba, NW Hungary). Central European Geology 55(2): 135–146. https://doi.org/10.1556/CEuGeol.55.2012.2.2
- Bradić-Milinović K, Ahnelt H, Rundić Lj, Schwarzhans W (2019) The lost freshwater goby fish fauna (Teleostei, Gobiidae) from the early Miocene of Klinci (Serbia). Swiss Journal of Palaeontology 138: 285–315. https://doi.org/10.1007/s13358-019-00194-4
- Bradić-Milinović K, Rundić Lj, Schwarzhans W (2021) Middle Miocene otoliths of freshwater fishes from the Vračević Lake (Serbian Lake System). Geološki Anali Balkanskoga Poluostrva 82(2): 1–24. https://doi.org/10.2298/GABP210616006B
- Bratishko A, Schwarzhans W, Reichenbacher B, Vernyhorova Y, Ćorić S (2015) Fish otoliths from the Konkian (Miocene, early Serravallian) of Mangyshlak (Kazakhstan): testimony to an early endemic evolution in the Eastern Paratethys. Paläontologische Zeitschrift 89: 839–889. https://doi.org/10.1007/s12542-015-0274-4
- Bratishko A, Kovalchuk O, Schwarzhans W (2017) Bessarabian (Tortonian, Late Miocene) fish otoliths from a transitional freshwater-brackish environment of Mykhailivka, southern Ukraine. Palaeontologia Electronica 20.3.44A: 1–13. https://doi.org/10.26879/769
- Bratishko A, Schwarzhans W, Vernyhorova Y (2023) The endemic marine fish fauna from the Eastern Paratethys reconstructed from otoliths from the Miocene (middle Sarmatian s.l.; Bessarabian) of Jurkine (Kerch Peninsula, Crimea). Rivista Italiana di Paleontologia e Stratigrafia 129(1): 111–184. https://doi.org/10.54103/2039-4942/18877
- Brzobohatý R (1969) Die Fischfauna des südmährischen Unter-Miozäns. Folia Facultatis Scientiarum Naturalium Universitatis Purkynianae Brunensis, Geologia 17(1): 3–49.
- Brzobohatý R (1992) Otolithen aus dem Obermiozän, Pontien, des Wiener Beckens (Götzendorf und Stixneusiedl, NÖ). Annalen des Naturhistorischen Museums in Wien Serie A 94: 1–6. https://www.jstor.org/stable/41701918
- Brzobohatý R, Pana I (1985) Die Fischfauna des Pannonien. In: Papp A, Jâmbor Ä, Steininger FF (Eds) M6, Pannonien (Slovenien und Serbien). Chronostratigraphie und Neostratotypen 7: 427–439.
- Carnevale G, Schwarzhans W (2022) Marine life in the Mediterranean during the Messinian Salinity Crisis: a paleoichthyological perspective. Rivista Italiana de Paleontologia e Stratigrafia 128(2): 283–324. https://doi.org/10.54103/2039-4942/15964
- Chalupova B (2008) Sarmatská rybia fauna z vrtu TPM-23B Smolenice (dunajská panva, Slovensko). Mineralia Slovaca 40: 53–58. https://www.geology.sk/wp-content/uploads/2019/10/Chalup.pdf
- Chao L (1986) Sciaenidae. In: Whitehead PJP, Bauchot M-L, Hureau J-C, Nielsen J, Tortonese E (Eds) Fishes of the North-eastern Atlantic and the Mediterranean. 2: 865–874. https://doi.org/10.2307/1444931
- Clarke MR (1978) The cephalopod statolith – an introduction to its form. Journal of the Marine Biological Association UK 58: 701–712. https://doi.org/10.1017/S0025315400041345
- Clarke MR, Maddock L (1988) Statoliths from living species of cephalopods and evolution. The Mollusca, Paleontology and Neontology of Cephalopods 12: 169–184. https://doi.org/10.1016/B978-0-12-751412-3.50016-4
- Cope ED (1871) Contribution to the ichthyology of the Lesser Antilles. Transactions of the American Philosophical Society 14(3): 445–483. https://doi.org/10.2307/1005256
- Cope ED (1887) The orders of fishes are defined by modifications of the skeleton of lesser significance than those included in statements II. and III. In: General notes: geology and palaeontology. IV. American Naturalist 21(11): 1016–1019. https://doi.org/10.1086/274597
- Fricke R, Eschmeyer WN, Van der Laan R [Eds] (2023) Echmeyer's catalog of fishes: Genera, species, references. http://researcharchive.calacademy.org/research/ichthyology/catalog/fishcatmain.asp [Electronic version accessed July 2024]
- Froese R, Pauly D [Eds] (2024) FishBase. www.fishbase.org [Version (02/2024)]
- Fuchs T (1871) Ueber die Fischfauna der Congerien-Schichten. Verhandlungen der kaiserlich-königlichen geologischen Reichsanstalt 1871(13): 227–228. https://www.zobodat.at/pdf/VerhGeolBundesanstalt_1871_0227-0228.pdf
- Gaudant J (2005) L'ichthyofaune du Badenien supérieur diatomitique de Várpalota (comté de Veszprém, Hongrie): Signification paléoécologique. Földtani Közlöny 135(1): 1–30.
- Gayet M, Meunier FJ (1991) First discovery of Polypteridae (Pisces, Cladistia, Polypteriformes) outside of Africa. Geobios 24(4): 463–466. https://doi.org/10.1016/S0016-6995(06)80246-9
- Geary DH, Magyar I, Müller P (2000) Ancient Lake Pannon and its endemic molluscan fauna (Central Europe; Mio-Pliocene). Advances in Ecological Research 31: 463–482. https://doi.org/10.1016/S0065-2504(00)31025-X
- Gierl C, Reichenbacher B, Gaudant J, Erpenbeck D, Pharisat A (2013) An extraordinary gobioid fish fossil from southern France. PLoS ONE 8(5): e64117. https://doi.org/10.1371/journal.pone.0064117
- Goodrich ES (1930) Studies on the structure & development of vertebrates. Macmillan and Co., London, 837 pp. https://doi.org/10.5962/bhl.title.82144
- Gulyas S (2001) The paleogeography of Lake Pannon during deposition of the Congeria rhomboidea Beds. Geologia Croatica 54(1): 15–26. https://doi.org/10.4154/GC.2001.02
- Harzhauser M, Mandic O (2010) Neogene dreissenids in Central Europe: evolutionary shifts and diversity changes. In: van der Velde G, Rajagopal S, bij de Vaate A (Eds) The Zebra Mussel in Europe. Backhuys Publishers, Leiden, 11–28.
- Harzhauser M, Tempfer PM (2004) Late Pannonian wetland ecology of the Vienna Basin based on molluscs and lower vertebrate assemblages (Late Miocene, MN 9, Austria). CFS Courier Forschungsinstitut Senckenberg 246: 55–68.
- Harzhauser M, Piller WE, Steininger FF (2002) Circum-Mediterranean Oligo-Miocene biogeographic evolution – the gastropods' point of view. Palaeogeography, Palaeoclimatology, Palaeoecology 183: 103–133. https://doi.org/10.1016/S0031-0182(01)00464-3
- Harzhauser M, Daxner-Höck G, Piller WE (2004) An integrated stratigraphy of the Pannonian (Late Miocene) in the Vienna Basin. Austrian Journal of Earth Sciences 95/96: 6–19. https://www.ajes.at/images/AJES/archive/Band%2095-96/harzhauser_daxner-hoeck_piller_ajes_v95_96.pdf
- Harzhauser M, Peresson M, Benold C, Mandic O, Ćorić S, De Lange GJ (2022) Environmental shifts in and around Lake Pannon during the Tortonian Thermal Maximum based on a multi-proxy record from the Vienna Basin (Austria, Late Miocene, Tortonian). Palaeogeography, Palaeoclimatology, Palaeoecology 610: 113322. https://doi.org/10.1016/j.palaeo.2022.111332
- Harzhauser M, Peresson M, Benold C, Mandic O (2024) Potentially astronomically forced climate sensitivity of Lake Pannon during the Tortonian Thermal Maximum (Late Miocene, Vienna Basin, Austria). Newsletters on Stratigraphy 57: 89–107. https://doi.org/10.1127/nos/2023/0763
- Herbert Mainero A, Vasilyan D, Reichenbacher B (2024) Two new genera of killifsh (Cyprinodontiformes) from the Middle Miocene of the Bugojno Basn, Bosnia and Herzegowina: insghts into the lost diversity of Valenciidae. Journal of Systematic Palaeontology 22(1): 2412539. https://doi.org/10.1080/14772019.2024.2412539
- Jovanović G, Knežević S, Đurić D, Bosnakoff M, Paunović G (2010) Upper Miocene fauna of Orešac near Smederevo (Serbia). Bulletin of the Natural History Museum in Belgrade 3: 67–93.
- Koken E (1884) Über Fisch-Otolithen, insbesondere über diejenigen der norddeutschen Oligocän-Ablagerungen. Zeitschrift der Deutschen Geologischen Gesellschaft 36: 500–565.
- Kovalchuk O, Otero O, Barkaszi Z, Murray AM, Divay JD (2024) A new species of Lates (Perciformes, Latidae) from the Late Miocene of Ukraine and notes on the latest records of lates perches in the Eastern Paratethys. Journal of Vertebrate Paleontology 43(4): e2299314. https://doi.org/10.1080/02724634.2023.2299314
- Lin Z, Strauss H, Peckmann J, Roberts AP, Lu Y, Sun X, Chen T, Harzhauser M (2023) Seawater sulphate heritage governed early Late Miocene methane consumption in the long-lived Lake Pannon. Communications Earth & Environment 4: 207. https://doi.org/10.1038/s43247-023-00879-2
- Lombarte A, Chic Ò, Parisi-Baradad V, Olivella R, Piera J, García-Ladona E (2006) A web-based environment from shape analysis of fish otoliths. The AFORO database. Scientia Marina 70: 147–152. https://doi.org/10.3989/scimar.2006.70n1147
- Lörenthey J (1906) Beiträge zur Fauna und stratigraphischen Lage der pannonischen Schichten in der Umgebung des Balatonsees. Resultate der wissenschaftlichen Erforschung des Balatonsees 1(1): 1–216. https://www.zobodat.at/pdf/Resultate-wiss-Erforschung-Balatonsee_1_4_0001-0215.pdf
- Magyar I (2021) Chronostratigraphy of clinothem-filled non-marine basins: datig the Pannonian Stage. Global and Planetary Change 205: 103609. https://doi.org/10.1016/j.gloplacha.2021.103609
- Magyar I, Geary DH, Müller P (1999) Palaeogeographic evolution of the Late Miocene Lake Pannon in Central Europe. Palaeogeography, Palaeoclimatology, Palaeoecology 147: 151–167. https://doi.org/10.1016/S0031-0182(98)00155-2
- Magyar I, Radivojević S, Sztanó O, Synak R, Ujszászi K, Pócsik M (2013) Progradation of the paleo-Danube shelf margin across the Pannonian Basin during the Late Miocene and Early Pliocene. Global and Planetary Change 103: 168–173. https://doi.org/10.1016/j.gloplacha.2012.06.007
- Malz H (1978) Vergleichend-morphologische Untersuchungen an aquitanen Fisch-Otolithen aus dem Untergrund von Frankfurt am Main. Senckenbergiana Lethaea 59(4–6): 441–481.
- Menzel H, Becker-Platen JD (1981) Otolithen aus dem Tertiär der Türkei (Känozoikum und Braunkohlen der Türkei 24). Geologisches Jahrbuch B 42: 5–91.
- Müller P, Geary DH, Magyar I (2007) The endemic molluscs of the Late Miocene Lake Pannon: their origin, evolution, and family-level taxonomy. Lethaia 32(1): 47–60. https://doi.org/10.1111/j.1502-3931.1999.tb00580.x
- Muzek K, Mandic O, Tadesse VH, Harzhauser M, Kovačić M, Kurečić T, Pezelj Đ (2024) Tracing the origin of Lago Mare biota: ostracods and mollusks from the late Neogene of the Slavonian mountains in the southern Pannonian Basin (NE Croatia). Palaeogeography, Palaeoclimatology, Palaeoecology 632: 111847. https://doi.org/10.1016/j.palaeo.2023.111847
- Near TJ, Thacker C (2024) Phylogenetic classification of living and fossil ray-finned fishes (Actinopterygii). Bulletin of the Peabody Museum of Natural History 65(1): 3–302. https://doi.org/10.3374/014.065.0101
- Neubauer TA, Harzhauser M, Kroh A (2013) Phenotypic evolution in a fossil gastropod species lineage: evidence for adaptive radiation? Palaeogeography, Palaeoclimatology, Palaeoecology 370: 117–126. https://doi.org/10.1016/j.palaeo.2012.11.025
- Neubauer TA, Harzhauser M, Mandic O, Kroh A, Georgopoulou E (2016) Evolution, turnovers and spatial variation of the gastropod fauna of the Late Miocene biodiversity hotspot Lake Pannon. Palaeogeography, Palaeoclimatology, Palaeoecology 442: 84–95. https://doi.org/10.1016/j.palaeo.2015.11.016
- Nolf D (1981) Révision des Types d'Otolithes de Poissons Fossiles décrits par R. Schubert. Verhandlungen der Geologischen Bundesanstalt-Anstalt 1981(2): 133–183. https://www.zobodat.at/pdf/VerhGeolBundesanstalt_1981_0133-0183.pdf
- Nolf D (2013) The diversity of fish otoliths, past and present. Royal Belgian Institute of Natural Sciences, Brussels, 222 pp.
- Nolf D (2018) Otoliths of the fishes of the North Sea and the English Cannel. Fauna of Belgium, Royal Belgian Institute of Natural Sciences, Brussels, 277 pp.
- Nolf D, De Potter H, Lafond-Grellety J (2009) Hommage à Joseph Chaine et Jean Duvergier. Diversité et variabilité des otolithes des poissons. Palaeo Publishing & Library Special Publication 1, 208 pp.
- Otero O, Likius A, Vignaud P, Brunet M (2006) A new polypterid fish: Polypterus faraou sp. nov. (Cladistia, Polypteridae) from the Late Miocene, Toros-Menalla, Chad. Zoological Journal of the Linnean Society 146: 227–237. https://doi.org/10.1111/j.1096-3642.2006.00201.x
- Pana I (1977) Problems arisen by the appearance of the new otoliths species and their stratigraphical implications. Examples of Dacian otoliths from S. Dobrudja. Analele Universitatii Bucuresti, Geologie 26: 105–124.
- Pana I (1982) Débris de poissons dans les dépôts pannoniens du Bassins de Beius (Roumanie). Revue Roumaine de Géologie, Géophysique et Géographie, Géologie 26: 69–92.
- Pana I (1995) Otolithes daciens. In: Marinescu F, Papaianopol I (Eds) PL1, Dacien. Chronostratigraphie und Neostratotypen, Neogen der Zentralen Paratethys 9: 427–445.
- Papp A (1951) Das Pannon des Wiener Beckens. Mitteilungen der Geologischen Gesellschaft Wien 39–41: 99–193. https://www.zobodat.at/pdf/MittGeolGes_39_41_0099-0193.pdf
- Papp A, Thenius E (1954) Vösendorf – ein Lebensbild aus dem Pannon des Wiener Beckens. Mitteilungen der Geologischen Gesellschaft Wien 46: 1–109. https://www.zobodat.at/pdf/MittGeolGes_46_Sonderband_0001-0109.pdf
- Pindakiewicz MK, Hryniewicz K, Janiszewska K, Kaim A (2022) First Cretaceous cephalopod statoliths fill the gap between Jurassic and Cenozoic forms. Comptes Rendus Paleovol 21(36): 801–813. https://doi.org/10.5852/cr-palevol2022v21a36
- Přikryl T, Brzobohatý R, Carnevale G (2021) Skeletal remains with otoliths in situ of the Miocene croaker Trewasciaena cf. kokeni (Teleostei, Sciaenidae) from the Pannonian of the Vienna Basin. Bulletin of Geosciences 96(1): 19–28. http://www.geology.cz/bulletin/fulltext/1813_Prikryl_210110.pdf
- Purnell MA, Darras LPG (2016) 3D tooth microwear texture analysis in fishes as a test of dietary hypotheses of durophagy. Surface Topography: Metrology and Properties 4(1): 014006. https://doi.org/10.1088/2051-672X/4/1/014006
- Radwańska U (1984) Some new fish otoliths from the Korytnica Clays (Middle Miocene; Holy Cross Mountains, Central Poland). Acta Geologica Polonica 34(3–4): 299–323. https://geojournals.pgi.gov.pl/agp/article/download/9501/8048
- Radwańska U (1992) Fish otoliths in the Middle Miocene (Badenian) deposits of southern Poland. Acta Geologica Polonica 42: 141–328. https://geojournals.pgi.gov.pl/agp/article/download/9343/7892
- Reichenbacher B (1988) Die Fischfauna der Kirchberger Schichten (Unter-Miozän) an der Typuslokalität Illerkirchberg bei Ulm. Stuttgarter Beiträge zur Naturkunde Serie B 139: 1–53.
- Reichenbacher B (1993) Mikrofaunen, Paläogeographie und Biostratigraphie der miozänen Brack- und Süsswassermolasse in der westlichen Paratethys unter besonderer Berücksichtigung der Fisch-Otolithen. Senckenbergiana Lethaea 73(2): 277–374.
- Reichenbacher B, Bannikov AF (2022) Diversity of gobioid fishes in the late Middle Miocene of northern Moldova, Eastern Paratethys – part I: an extinct clade of Lesueurigobius look-alikes. PalZ 96: 67–112. https://doi.org/10.1007/s12542-021-00573-8
- Reichenbacher B, Bannikov AF (2023) Diversity of gobioid fishes in the late Middle Miocene of northern Moldova, Eastern Paratethys – part II: description of †Moldavigobius helenae gen. et sp. nov. PalZ 97: 365–381. https://doi.org/10.1007/s12542-022-00639-1
- Rückert-Ülkümen N (1992) Zur Stratigraphie, Palökologie und Otolithenfauna der Braunkohleschichten (Oligo-Miozän) von Küçük Doanca Köyü (Thrakien, Türkei). Mitteilungen der Bayerischen Staatsammlung für Paläontologie und Historische Geologie 32: 93–114. https://www.zobodat.at/pdf/Mitt-Bayer-Staatsslg-Pal-hist-Geol_32_0093-0114.pdf
- Rückert-Ülkümen N (1993) Neue Beiträge zur Tertiär-Stratigraphie und Otolithenfauna der Umgebung von Istanbul (Küçukçekmece - und Büyükçekmece See). Mitteilungen der Bayerischen Staatsammlung für Paläontologie und Historische Geologie 33: 51–89. https://www.zobodat.at/pdf/Mitt-Bayer-Staatsslg-Pal-hist-Geol_33_0051-0089.pdf
- Schubert RJ (1902) Die Fischotolithen des österreichisch-ungarischen Tertiärs. I. Die Sciaeniden. Jahrbuch der kaiserlichen und königlichen Geologischen Reichsanstalt 51: 302–316. https://www.zobodat.at/pdf/JbGeolReichsanst_051_0301-0316.pdf
- Schubert RJ (1906) Die Fischotolithen des österreichisch-ungarischen Tertiärs. III. Jahrbuch der kaiserlichen und königlichen Geologischen Reichs-Anstalt 56: 623–706. https://www.zobodat.at/pdf/JbGeolReichsanst_056_0623-0706.pdf
- Schultz O (2013) Catalogus Fossilium Austriae. 3. Pisces. Verlag der Österreichischen Akademie der Wissenschaften, Wien, 576 pp. https://doi.org/10.2307/j.ctt1vw0qw3
- Schulz-Mirbach T, Reichenbacher B (2006) Reconstruction of Oligocene and Neogene freshwater fish faunas—an actualistic study on cypriniform otoliths. Acta Palaeontologica Polonica 51: 283–304. https://www.app.pan.pl/archive/published/app51/app51-283.pdf
- Schwarzhans W (1978) Otolith-morphology and its usage for higher systematical units with special reference to the Myctophiformes s.l. Mededelingen van de Werkgroep voor Tertiaire en Kwartaire Geologie 15: 167–185. https://natuurtijdschriften.nl/pub/521340/MWTKG1978015004009.pdf
- Schwarzhans W (1993) A comparative morphological treatise of recent and fossil otoliths of the family Sciaenidae. Piscium Catalogus, Otolithi Piscium 1: 1–245.
- Schwarzhans W (2010) The otoliths from the Miocene of the North Sea Basin. Backhuys Publishers, Leiden, 352 pp.
- Schwarzhans W (2014) Otoliths from the Middle Miocene (Serravallian) of the karaman Basin, Turkey. Cainozoic Research 14(1): 35–69.
- Schwarzhans W, Carnevale G, Bannikov AF, Japundžić S, Bradić K (2017a) Otoliths in situ from Sarmatian (Middle Miocene) fishes of the Paratethys. Part I: Atherina suchovi Switchenska, 1973. Swiss Journal of Palaeontology 136(1): 7–17. https://doi.org/10.1007/s13358-015-0111-0
- Schwarzhans W, Carnevale G, Bratishko A, Japundžić S, Bradić K (2017b) Otoliths in situ from Sarmatian (Middle Miocene) fishes of the Paratethys. Part II: Gadidae and Lotidae. Swiss Journal of Palaeontology 136(1): 19–43. https://doi.org/10.1007/s13358-016-0114-5
- Schwarzhans W, Ahnelt H, Carnevale G, Japundžić S, Bradić K, Bratishko A (2017c) Otoliths in situ from Sarmatian (Middle Miocene) fishes of the Paratethys. Part III: tales from the cradle of the Ponto-Caspian gobies. Swiss Journal of Palaeontology 136(1): 45–92. https://doi.org/10.1007/s13358-016-0120-7
- Schwarzhans W, Carnevale G, Japundžić S, Bradić-Milinović K (2017d) Otoliths in situ from Sarmatian (Middle Miocene) fishes of the Paratethys. Part IV: Scorpaenidae, Labridae, and Gobiesocidae. Swiss Journal of Palaeontology 136(1): 93–108. https://doi.org/10.1007/s13358-017-0124-y
- Schwarzhans W, Carnevale G, Japundžić S, Bradić-Milinović K (2017e) Otoliths in situ from Sarmatian (Middle Miocene) fishes of the Paratethys. Part V: Bothidae and Soleidae. Swiss Journal of Palaeontology 136(1): 109–127. https://doi.org/10.1007/s13358-017-0128-7
- Schwarzhans W, Agiadi K, Carnevale G (2020) Late Miocene-Early Pliocene evolution of Mediterranean gobies and their environmental and biogeographic significance. Rivista Italiana di Paleontologia e Stratigrafia 126(3): 657–724. https://doi.org/10.13130/2039-4942/14185
- Schwarzhans W, Klots O, Ryabokon T, Kovalchuk O (2022) A rare window into a back-reef fish community from the Middle Miocene (late Badenian) Medobory Hills barrier reef in western Ukraine, reconstructed mostly by means of otoliths. Swiss Journal of Palaeontology 141: 18. https://doi.org/10.1186/s13358-022-00261-3
- Schwarzhans W, Klots O, Kovalchuk O, Dubikovska A, Ryabokon T, KovalenkoV (2024) Life on a Miocene barrier reef - fish communities and environments in the Medobory backreef. Palaeontologia Electronica 27(3): 27.3.a46. https://doi.org/10.26879/1429
- Trewavas E (1977) The sciaenid fishes (croakers or drums) of the Indo-West-Pacific. Transactions of the Zoological Sciety London 33: 253–541. https://doi.org/10.1111/j.1096-3642.1977.tb00052.x
- Weiler W (1943) Die Otolithen aus dem Jungtertiär Süd-Rumäniens, 1. Buglow und Sarmat. Senckenbergiana Lethaea 26: 87–115.
- Weiler W (1949) Die Otolithen aus dem Jungtertiär Süd-Rumäniens, 1b. Ergänzende Tafeln zu den Otolithen des Buglow und Sarmat. Senckenbergiana Lethaea 30: 291–293.
- Weinfurter E (1950) Die operpannonische Fischfauna vom Eichkogel bei Mödling. Sitzungsberichte der österreichischen Akademie der Wissenschaften, mathematisch-naturwissenschaftliche Klasse 1 159: 37–50. https://www.zobodat.at/pdf/SBAWW_159_0037-0050.pdf
- Weinfurter E (1954) Pisces. In: Papp A, Thenius E (Eds) Vösendorf – ein Lebensbild aus dem Pannon des Wiener Beckens. Mitteilungen der Geologischen Gesellschaft Wien 46: 30–41. https://www.zobodat.at/pdf/MittGeolGes_46_Sonderband_0001-0109.pdf
- Woodward AS (1891) Catalogue of the fossil fishes in the British Museum (Natural History). Part II. British Museum (Natural History), London, 567 pp. https://doi.org/10.5962/bhl.title.61854
- Ziv E, Milgram J, Davis J, Soares A, Wilde F, Zaslansky P, Shahar R (2020) Neither cortical nor trabercular: an unusual type of bone in the heavy-load-bearing lower pharyngeal jaw of the black drum (Pogonias cromis). Acta Biomaterialia 104: 28–38. https://doi.org/10.1016/j.actbio.2020.01.001