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From continents to islands: tracking the red deer (Cervus elaphus Linnaeus, 1758) and its "troublesome cousin", the wapiti (Cervus canadensis Erxleben, 1777), in Europe

  • 1. Laboratoire méditerranéen de Préhistoire Europe Afrique (Lampea), Aix-Marseille université, CNRS, ministère de la Culture, , 5 rue du Château de l'Horloge, CS 90412, F-13094 Aix-en-Provence cedex 2 (France) romancroitor@europe.com
  • 2. Muséum Requien, 67 rue Joseph Vernet, 84 000 Avignon (France) Laboratoire Travaux et Recherches archéologiques sur les Cultures, les Espaces et les Sociétés (TRACES) CNRS, ministère de la Culture, université Toulouse 2 Jean-Jaurès Maison de la Recherche, 5 allées Antonio Machado, F-31058 Toulouse cedex 9 (France) evelyne.cregut1313@orange.fr

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Croitor, Roman, Crégut-Bonnoure, Évelyne (2025): From continents to islands: tracking the red deer (Cervus elaphus Linnaeus, 1758) and its "troublesome cousin", the wapiti (Cervus canadensis Erxleben, 1777), in Europe. Anthropozoologica 60 (11): 143-163, DOI: 10.5252/anthropozoologica2025v60a11, URL: https://sciencepress.mnhn.fr/sites/default/files/articles/pdf/anthropozoologica2025v60a11.pdf

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  • ABBAZZI L. 1995. - Occurrence of palmated Cervus elaphus from Italian late Pleistocene localities. Rendiconti Lincei, Scienze fisiche e naturali, Serie IX 6 (3): 189-206. https://doi.org/10.1007/ BF03001667
  • AHLEN I. 1965. - Studies on the red deer, Cervus elaphus L., in Scandinavia. II, Taxonomy and osteology of prehistoric and recent populations. Swedish Wildlife 3 (2): 89-176.
  • ARNOLD M. L. 2015. - Divergence with Genetic Exchange. Oxford University Press, Oxford, 251 p.
  • AVERBOUH A. (ed.) 2016. - Multilingual Lexicon of Bone Industry (Version 2). APPAM, GDRE PREHISTOS, Aix-en-Provence, 109 p. (Archaeological Studies; II /Hors-serie a Prehistoires mediterraneennes).
  • AZZAROLI A. 1947. - I cervi fossili della Toscana.Palaeontographia Italica 43: 46-81.
  • AZZAROLI A. 1961. - Il nanismo nei cervi insulari.Palaeontographia Italica 56: 1-31.
  • AZZAROLI A. 1979. - Critical remarks on some giant deer (genus Megaceros Owen) from the Pleistocene of Europe.Palaeontographia Italica 41: 5-16.
  • AZZAROLI A. 1983. - Quaternary mammals and the "End-Villafranchian" dispersal event - a turning point in the history of Eurasia. Palaeogeography, Palaeoclimatology, Palaeoecology 44 (1-2): 113-139.https://doi.org/10.1016/0031-0182(83)90008-1
  • AZZAROLI A. 1992. - The cervid genus Pseudodama n.g. in the Villafranchian of Tuscany.Palaeontographia Italica 79: 1-41.
  • BANKS W. E., D'ERRICO F., PETERSON A. T., KAGEYAMA M. & COLOMBEAU G. 2008. - Reconstructing ecological niches and geographic distributions of caribou (Rangifer tarandus) and red deer (Cervus elaphus) during the Last Glacial Maximum. Quaternary Science Reviews 27 (27-28): 2568-2575. https://doi. org/10.1016/j.quascirev.2008.09.013
  • BELGRAND E. 1883. - La Seine. I, Le bassin parisien aux ages antehistoriques. Imprimerie nationale, Paris, 35 p.
  • BENINDE J. 1937. - Uber die edelhirschformen von Mosbach, Mauer und Steinheim ad Murr. Palaontologische Zeitschrift 19: 79-116.
  • BOESKOROV G. G. 2005. - Taxonomic position of the Red Deer Cervus elaphus L. (Cervidae, Artiodactyla, Mammalia) from the Neopleistocene of Northeastern Asia. Paleontological Journal 39 (5): 73-84.
  • BOUCHUD J. 1966. - Essay sur le renne et climatologie du Paleolithique moyen et superieur. Imprimerie Magne, Perigueux, 297 p.
  • BOULE M. 1892. - Description geologique du Velay. Bulletin du Service de la carte geologique de la France 28: 1-259.
  • BREDA M. & LISTER A. M. 2013. - Dama roberti, a new species of deer from the early Middle Pleistocene of Europe, and the origins of modern fallow deer. Quaternary Science Reviews 69: 155-167. https://doi.org/10.1016/j.quascirev.2013.01.029
  • CABRERA A. 1914. - Fauna iberica: mamiferos. Museo Nacional de Ciencias Naturales, Madrid, 441 p.
  • CALOI L. & PALOMBO M. R. 1995. - Functional aspects and ecologicalimplications in Pleistocene endemic cervids of Sardinia, Sicily and Crete.Geobios 28 (2): 247-258.https://doi.org/10.1016/ S0016-6995(95)80231-2
  • CHERIN M., BREDA M., ESATTORE B., HART V., TUREK J., PORCIELLO F., ANGELI G., HOLPIN S. & IURINO D. A. 2022. - A Pleistocene Fight Club revealed by the palaeobiological study of the Dama-like deer record from Pantalla (Italy). Scientific Reports 12 (13898).https://doi.org/10.1038/s41598-022-18091-1
  • COVALENCO S. & CROITOR R. 2022. - On the origin of the red deer upper canine beads from the multilayer Late Paleolithic site of CosauTi (Moldova), in DIACONU V. (ed.), Ab lapide ad metallum: studia in honorem Professoris Marin Carciumaru. Istros, Braila; Constantin Matasa, Piatra-NeamT: 53-63 (Bibliotheca Memoriae Antiquitatis; xliv).
  • CREGUT-BONNOURE E. 2006. - European Ovibovini, Ovini and Caprini (Caprinae, Mammalia) from the Plio-Pleistocene: new interpretations, in KAHLKE R.-D., MAUL L. C. & MAZZA P. P. (eds), Late Neogene and Quaternary Biodiversity and Evolution: Regional Developments and Interregional Correlations. Proceedings of the 18th International Senckenberg Conference (VI International Palaeontological Colloquium in Weimar). Vol. I. E. Schweizerbart'schVerlagbuchshandlung (Nagele. Obermiller), Stuttgart: 139-158. (Courier Forschungsinstitut Senckenberg; 256).
  • CREGUT-BONNOURE E. 2007. - Apport des Caprinae et Antilopinae (Mammalia, Bovidae) a la biostratigraphie du Pliocene terminal et du Pleistocene d'Europe, in BRUGAL J.-P., LIMONDIN-LOZOUET N. & SEMAH F. (eds), Q5 Le Quaternaire, Limites et specificites, Paris, 1-3 Fevrier 2006. Quaternaire 18 (1): 73-97. https://doi.org/10.4000/quaternaire.996
  • CREGUT-BONNOURE E. 2020. - Les Ovibovini, Caprini et Ovini (Mammalia, Artiodactyla, Bovidae, Caprinae) du Plio-Pleistocene d'Europe : systematique, evolution et biochronologie. BAR Publishing, Oxford, 2 vol.: 351 + 306 p. (BAR International Series; 2975).
  • CROITOR R. 2006. - Early Pleistocene small-sized deer of Europe. Hellenic Journal of Geosciences 41: 89-117.
  • CROITOR R. 2011. - Early Pleistocene deer Cervus nestii (Cervidae, Mammalia) and the earliest dispersal of the genus Cervus (sensu stricto) in Western Eurasia, in TODERAS I. (ed.), Actual Problems of Protection and Sustainable Use of Animal World Diversity. International Conference of Zoologists. Academy of Sciences of Moldova, Department of Nature and Life Sciences, Institute of Zoology, Chisinau: 203-204.
  • CROITOR R. 2014. - Deer from Late Miocene to Pleistocene of Western Palearctic: matching fossil record and molecular phylogeny data. Zitteliana Reihe B: Abhandlungen der Bayerischen Staatssammlung fur Palaontologie und Geologie (32): 115-153.
  • CROITOR R. 2016. - Systematical position and paleoecology of the endemic deer Megaceroides algericus Lydekker, 1890 (Cervidae, Mammalia) from the late Pleistocene - early Holocene of North Africa. Geobios 49 (4): 265-283. https://doi.org/10.1016/j.geobios.2016.05.002
  • CROITOR R. 2018. - Plio-Pleistocene deer of Western Palearctic: Taxonomy, Systematics, phylogeny. Institute of Zoology of the Academy of Sciences of Moldova, Chisinau: 140 p.
  • CROITOR R. 2020a. - A new form of wapiti Cervus canadensis Erxleben, 1777 (Cervidae, Mammalia) from the Late Pleistocene of France. Palaeoworld 29 (4): 789-806.https://doi.org/10.1016/j. palwor.2019.12.001
  • CROITOR R. 2020b. - On a discovery of musk ox Ovibos moschatus (Zimmermann, 1780) (Bovidae, Mammalia) from the Upper Paleolithic site of Rashkov 7. Revista Arheologica 16 (2): 100- 105. https://doi.org/10.5281/zenodo.4957959
  • CROITOR R. 2021. - Early evolutionary radiation and diversity of the Old World telemetacarpal deer (Capreolinae, Cervidae, Mammalia). Neues Jahrbuch fur Geologie und Palaontologie-Abhandlungen 300 (1): 33-67. https://doi.org/10.1127/njgpa/2021/0978
  • CROITOR R. 2022. - Paleobiogeography of crown deer.Earth 3 (4): 1138-1160. https://doi.org/10.3390/earth3040066
  • CROITOR R. & BONIFAY M.-F. 2001. - Etude preliminaire des cerfs du gisement Pleistocene inferieur de Ceyssaguet (Haute-Loire). Paleo 13: 129-144. https://doi.org/10.4000/paleo.1011
  • CROITOR R. & COJOCARU I. 2016. - An antlered skull of a subfossil red deer, Cervus elaphus L., 1758 (Mammalia: Cervidae), from Eastern Romania.Acta Zoologica Bulgarica 68 (3): 407-414.
  • CROITOR R. & OBADA T. 2018. -On the presence of Late Pleistocene wapiti, Cervus canadensis Erxleben, 1777 (Cervidae, Mammalia) in the Palaeolithic site ClimauTi II (Moldova). Contributions to Zoology 87 (1): 1-10.https://doi.org/10.1163/18759866-08701001
  • CROITOR R. & ROBINSON C. 2020. - A revision of "Cervus" punjabiensis Brown, 1926 (Cervidae, Mammalia) from the Upper Siwaliks of Chandigarh, India. Quaternary International 550: 147-158. https://doi.org/10.1016/j.quaint.2020.04.020
  • CROITOR R., STEFANIAK K., PAWLOWSKA K., RIDUSH B., WOJTAL P. & STACH M. 2014. - Giant deer Megaloceros giganteus Blumenbach, 1799 (Cervidae, Mammalia) from Palaeolithic of Eastern Europe. Quaternary International 326-327: 91-104. https://doi. org/10.1016/j.quaint.2013.10.068
  • CUVIER G. 1823. - Recherches sur les ossements fossiles. Tome quatrieme. Dufour et D'Ocagne, Paris, 514 p.
  • DANILKIN A. A. 1999. - Deer (Cervidae). Geos, Moscow, 552 p. [in Russian].
  • DAVID A. I. 1980. - Theriofauna of the Pleistocene of Moldova. Chisinau, Stiinta, 185 p. [in Russian].
  • DE STEFANO G. 1911. - I mammiferi preistorici dell'Imolese. Palaeontographia Italica 17: 49-139.
  • DI STEFANO G. & PETRONIO C. 1992. - Nuove osservazioni su Cervus elaphus acoronatus Beninde del Pleistocene europeo. Bollettino della Societa Paleontologica Italiana 31 (3): 295-315.
  • DI STEFANO G. & PETRONIO C. 1998. - Origin and relationships among Dama-like cervids in Europe. Neues Jahrbuch fur Geologie und Palaontologie - Abhandlungen 207: 37-55
  • DI STEFANO G. & PETRONIO C. 2002. - Systematics and evolution of the Eurasian Plio-Pleistocene tribe Cervini (Artiodactyla, Mammalia). Geologica Romana 36: 311-334.
  • DI STEPHANO G. & PETRONIO C. 2021. - Importance of the morphological plasticity of Cervus elaphus in the biochronology of the Middle and Late Pleistocene of the Italian peninsula. The Science of Nature 108 (5): 40.
  • DOAN K., ZACHOS F. E., WILKENS B., VIGNE J. D., PIOTROWSKA N., STANKOVIc A., JeDRZEJEWSKA B., STEFANIAK K. & NIEDZIALKOWSKA M. 2017. - Phylogeography of the Tyrrhenian red deer (Cervus elaphus corsicanus) resolved using ancient DNA of radiocarbon-dated subfossils.Scientific Reports 7 (2331). https://doi.org/10.1038/s41598-017-02359-y
  • DRUCKER D. G., BRIDAULT A., CUPILLARD C., HUJIC A. & BOCHERENS H. 2011. - Evolution of habitat and environment of red deer (Cervus elaphus) during the Late-glacial and early Holocene in eastern France (French Jura and the western Alps) using multi-isotope analysis (δ13C, δ15N, δ18O, δ34S) of archaeological remains. Quaternary International 245 (2): 268-278.https://doi. org/10.1016/j.quaint.2011.07.019
  • ELLERMAN J. R. & MORRISON-SCOTT T. C. S. 1951. - Checklist of Palaearctic and Indian Mammals 1758 to 1946. Second Edition. Trustees of the British Museum (Natural History), London, 810 p.
  • FLEROV K. K. 1952. - Musk Deer and Deer. House of Academy of Sciences, Moscow, Leningrad, 256 p. (Fauna of USSR. Mammals; 1 [2]) [in Russian].
  • FRIANT M. 1952. - Le Strongyloceros spelaeus Owen (Cerf geant des Cavernes). Comptes rendus hebdomadaires des seances de l'Academie des sciences 234: 1398-1400.
  • FRIANT M. 1957. - Le Strongyloceros spelaeus Owen de Kent's Hole (Angleterre). Annales de la Societe geologique du Nord 77: 57-74.
  • GAUDRY A. 1872. - Sur les ossements d'animaux quaternaires que M. l'abbe David a recueillis en Chine. Bulletin de la Societe Geologique de France Serie 2 29: 177-179.
  • GAUDRY A. 1876. - Le couloir de Louverne.Materiaux pour l'histoire des temps quaternaires 1: 34-50.
  • GAUDRY A. 1880. - L'existence des saigas en France a l'epoque Quaternaire. Materiaux pour l'histoire des temps quaternaires 2: 65-82.
  • GEIST V. 1987. - On the evolution of the Caprinae, in HIROAKI SOMA (ed.),The Biology and Management of Capricornis and Related Mountain Antelopes. Springer Netherlands, Dordrecht: 3-40.
  • GEIST V. 1998. - Deer of the World: Their Evolution, Behaviour, and Ecology. Stackpole Books, Mechanicsburg, PA, 421 p.
  • GERVAIS P. 1861. - Notes sur differentes especes de vertebres fossiles observes pour la plupart dans le midi de la France. Annales des Sciences naturelles. Quatrieme serie, Zoologie 16: 286-302.
  • GERVAIS P. 1872. - Rapport sur les decouvertes faites dans la Grotte de Loubeau, pres Melle.Archives des missions scientifiques et litteraires 7s: 531-536.
  • GLIOZZI E., MALATESTA A. & SCALONE E. 1993. - Revision of Cervus elaphus siciliae Pohlig, 1893, Late Pleistocene endemic seer of the Siculo-Maltese District. Geologica Romana 29: 307-353.
  • GOSS R. J. 1983. - Deer Antlers: Regeneration, Function and Evolution. Academic Press, New York, 315 p.
  • GUADELLI J. L. 1997. - Les cerfs du Wurm ancien en Aquitaine. Paleo 8: 99-108. https://doi.org/10.3406/pal.1996.908
  • HEINTZ E. 1970. - Les Cervides villafranchiens de France et d'Espagne : Volumes 1 & 2. Museum national d'Histoire naturelle, Paris, 206 p. (Memoires du Museum national d'Histoire naturelle, Serie C - Sciences de la Terre (1950-1992); 22).
  • HEPTNER V. G. & ZALKIN V. I. 1947. - Deer of the USSR (Systematics and Zoogeography). Transactions on Study of Fauna and Flora of the USSR, the Moscow Society of Naturalists 10 (25): 1-176 [in Russian].
  • HU P., SHAO Y., XU J., WANG T., LI Y., LIU H., RONG M., SU W., CHEN B., CUI S. & CUI X. 2019. - Genome-wide study on genetic diversity and phylogeny of five species in the genus Cervus. BMC Genomics 20 (384). https://doi.org/10.1186/ s12864-019-5785-z
  • JANIS C. M. & LISTER A. M. 1985. - The morphology of the lower fourth premolar as a taxonomic character in the Ruminantia (Mammalia; Artiodactyla), and the systematic position of Triceromeryx. Journal of Paleontology 59 (2): 405-410.
  • JOHNSTON H. 1903. - British Mammals: An Attempt to Describe and Illustrate the Mammalian Fauna of the British Islands from the Commencement of the Pleistocene Period Down to the Present Day. Hutchinson & Co., London, 340 p.
  • KAHLKE H.-D. 2001. - Neufunde fon Cerviden-resten aus dem Unterpleistozan von Untermassfeld, in KAHLKE H. D., Das Pleistozan von Untermassfeld bei Meningen (Thuringen). Teil 2. Propylaeum, Heidelberg: 461-482. (Monographien des RomischGermanischen Zentralmuseums Mainz; 40.2). https://doi. org/10.11588/propylaeum.559
  • KAISER T. M. & CROITOR R. 2004. - Ecological interpretations of early Pleistocene deer (Mammalia, Cervidae) from Ceyssaguet (Haute-Loire, France). Geodiversitas 26 (4): 661-674.
  • KUWAYAMA R. & OZAWA T. 2000. - Phylogenetic relationships among European Red Deer, Wapiti, and Sika Deer inferred from mitochondrial DNA sequences. Molecular Phylogenetics and Evolution 15 (1): 115-123. https://doi.org/10.1006/mpev.1999.0731
  • LISTER A. 1987a. - Diversity and evolution of antler form in Quaternary deer, in WEMMER C. M. (ed.), Biology and Management of the Cervidae. Smithsonian, Washington: 81-98. https://doi. org/10.13140/2.1.2517.6961
  • LISTER A. 1987b. - Giant deer and giant red deer from Kent's Cavern, and the status of Strongyloceros spelaeus Owen. Transactions and Proceedings of the Torquay Natural History Society 19: 189-198.
  • LISTER A. 1989. - Rapid dwarfing of red deer on Jersey in the last interglacial. Nature 342: 539-542.https://doi.org/10.1038/342539a0
  • LISTER A. 1990. - Critical reappraisal of the middle Pleistocene deer "Cervus" elaphoides Kahlke. Quaternaire 1 (3-4): 175-192. https://doi.org/10.3406/quate.1990.1935
  • LISTER A. 1999. - The Pliocene deer of the Red Crag Nodule Bed (UK), in REUMER J. W. F. & DE VOS J. (eds.), Elephants Have a Snorkel! Papers in Honour of Paul Y. Sondaar. Deinsea 7 (1): 215-222.
  • LONNBERG E. 1906. - On the geographic races of red deer in Scandinavia. Arkiv for Zoologi 3 (9): 1-19.
  • LORENZINI R. & GAROFALO L. 2015. - Insights into the evolutionary history of Cervus (Cervidae, tribe Cervini) based on Bayesian analysis of mitochondrial marker sequences, with first indications for a new species. Journal of Zoological Systematics and Evolutionary Research 53 (4): 340-349.https://doi.org/10.1111/jzs.12104
  • LUDT C. J., SCHROEDER W., ROTTMANN O. & KUEHN R. 2004. - Mitochondrial DNA phylogeography of red deer (Cervus elaphus). Molecular Phylogenetics and Evolution 31 (3): 1064-1083.https:// doi.org/10.1016/j.ympev.2003.10.003
  • LYDEKKER R. 1876. - Indian Tertiary and post-Tertiary Vertebrata. Part II, Molar teeth and other remains of Mammalia. Palaeontologia Indica, Memoires of Geological Survey of India Series X 1: 19-87.
  • LYDEKKER R. 1896. - On apparently New Deer from North China, in the menagerie of the Duke of Bedford at Woburn Abbey. Proceedings of the Zoological Society of London 1896: 930-934.
  • LYDEKKER R. 1898. - Deer of All Lands: A History of the Family Cervidae Living and Extinct. Rowland Ward, London, 329 p.
  • LYDEKKER R. 1915. - Catalogue of the Ungulate Mammals in the British Museum (Natural History). Vol. IV, Artiodactyla, Families Cervidae (Deer), Tragulidae (Chevrotains), Camelidae (Camels and Llamas), Suidae (Pigs and Peccaries), and Hippopotamidae (Hippopotamuses). Trustees of the British Museum, London, 438 p.
  • MAGNIEZ P., MOIGNE A.-M., TESTU A. & LUMLEY H. DE 2013. - Biochronologie des mammiferes quaternaires. Apport des Cervidae du site Pleistocene moyen de la Caune de l'Arago (Tautavel, Pyrenees-Orientales, France). Quaternaire 24 (4): 477-502. https://doi.org/10.4000/quaternaire.6830
  • MARTIN D. 1893. - Le bassin du Pignon et sa faune : note sur la vallee du Buech.Bulletin de la Societe d'Etudes - Gap 6 (2): 5-19.
  • MARTINEZ-NAVARRO B. 2010. - Early Pleistocene faunas of Eurasia and hominid dispersals, in FLEAGLE J. G., SHEA J. J., GRINE F. E., BADEN A. L. & LEAKEY R. E. (eds), Out of Africa. Vol. I, The First Hominin Colonization of Eurasia. Springer, New York: 207-224. (Vertebrate Paleobiology and Paleoanthropology Series). https:// doi.org/10.1007/978-90-481-9036-2_13
  • MAYR E. 1942. - Systematics and the Origin of Species from the Viewpoint of a Zoologist. Columbia University Press, New York, 334 p.
  • MEIRI M., LISTER A. M., HIGHAM T. F., STEWART J. R., STRAUS L. G., OBERMAIER H., GONZALEZ MORALES M.R., MARIN-ARROYO A.B. & BARNES I. 2013. - Late-glacial recolonization and phylogeography of European red deer (Cervus elaphus L.). Molecular Ecology 22 (18): 4711-4722. https://doi.org/10.1111/mec.12420
  • MEIRI M., KOSINTSEV P., CONROY K., MEIRI S., BARNES I. & LISTER A. 2018. - Subspecies dynamics in space and time: a study of the red deer complex using ancient and modern DNA and morphology. Journal of Biogeography 45 (2): 367-380. https:// doi.org/10.1111/jbi.13124
  • NARAYAN S., SRIVASTAVA T., SREEDHARAN G., PANDA B. K., HAMEED J., VASUDEVAN K. & REDDY P. A. 2024. - Genetic insights to assist management of the Critically Endangered hangul Cervus hanglu hanglu in the Kashmir Himalaya. Oryx 58 (3): 396-404.https://doi.org/10.1017/S0030605323001266
  • NELSON E. W. 1902. - A new species of elk from Arizona.Bulletin of the American Museum of Natural History 16 (1): 1-12.
  • OSHIRO I. & NOGARA T. 2000. - Distribution of Pleistocene terrestrial vertebrates and their migration to the Ryukyus. Tropics 10 (1): 41-50. https://doi.org/10.3759/tropics.10.41
  • OGILBY W. 1840. - List of Mammals exhibited for the first time at the Menagerie. Reports of the Council and Auditors of the Zoological Society of London 1840: 21-22.
  • OWEN R. 1846. - A History of British Fossil Mammals, and Birds. John van Voorst, London, 560 p.
  • PAVELKOVA RICANKOVA V., ROBOVSKY J. & RIEGERT J. 2014. - Ecological structure of recent and Last Glacial Mammalian faunas in northern Eurasia: the case of Altai-Sayan Refugium. PLoS ONE 9 (1): e85056. https://doi.org/10.1371/journal.pone.0085056
  • PETRONIO C. 1979. - Dama nestii eurygonos Azz. di Capena (Roma). Geologica Romana 18: 105-125.
  • PETRONIO C., BELUCCI L. & DI STEFANO G. 2013. - Axis eurygonos from Pirro Nord (Apricena, Southern Italy). Palaeontographica Abteilung A 298 (1-6): 169-181.https://doi.org/10.1127/ pala/298/2013/169
  • PFEIFER T. 1997. - Dama (Pseudodama) reichenaiui (Kahlke 1996) (Artyodactyla: Cervidae, Cervini) aus den Mosbach-Sanden (Wiesbaden-Biebrich). Mainzer naturwissenschaftliches Archiv 35: 31-59.
  • PITRA C., FICKEL J., MEIJAARD E. & GROVES C. 2004. - Evolution and phylogeny of old world deer. Molecular Phylogenetics and Evolution 33 (3): 880-895.
  • POCOCK R. I. 1912. - On a rare stag (Cervus wallichii) from Nepal recently presented to the Zoological Society by His Majesty King George. Proceedings of the Zoological Society of London 3-4: 505-913.
  • POLZIEHN R. O. & STROBECK C. 2002. - A phylogenetic comparison of red deer and wapiti using mitochondrial DNA. Molecular Phylogenetics and Evolution 22 (3): 342-356. https:// doi.org/10.1006/mpev.2001.1065
  • POMEL M. 1853. - Catalogue methodique et descriptif des vertebres fossiles decouverts dans le bassin hydrographique de la Loire. Chez J.-B. Bailliere, Paris, 193 p.
  • PRAT F. & SUIRE C. 1971. - Remarques sur les cerfs contemporains des deux premiers stades wurmiens. Bulletin de la Societe prehistorique francaise. Comptes rendus des seances mensuelles 68 (3): 75-79. https://doi.org/10.3406/bspf.1971.4300
  • RANCE C. E. DE 1888. - Notes on the Vale of Clwyd Caves.Proceedings of the Yorkshire Geological and Polytechnic Society 11 (1): 1-20.
  • RIVIERE E. 1873. - Decouverte d'un squelette humain de l'epoque paleolithique dans les cavernes des Baousse-Rousse dites Grottes de Menton. J.-B. Bailliere et fils, Paris, 64 p.
  • RIVIERE E. 1905. - Les parois gravees et peintes de la grotte de la Mouthe (Dordogne). Bulletins et Memoires de la Societe d'Anthropologie de Paris 4 (1): 191-196.
  • SCHONEWALD C. 1994. - Cervus canadensis and C. elaphus: North American subspecies and evaluation of clinal extremes. Acta Theriologica 39 (4): 431-452.
  • SCHREBER J. C. D. VON 1836. - Der Saugetiere in Abbildungen nach der Natur, mit Beschreibungen. Abteilung 5. Erlangen, Leipzig, i-vi + 939-1839.
  • SERRES M. DE 1838. - Essai sur les cavernes a ossements et sur les causes qui les y ont accumules [Troisieme edition]. Bailliere, Paris, 412 p.
  • SEVERTZOFF N. A. 1873. - Altitudinal and latitudinal distribution of Turkestan animals.Proceedings of the Imperial Society of Naturalists, Anthropologists, and Ethnographists 8 (2): 1-157 [in Russian].
  • SEVERTZOFF N. A. 1876. - The mammals of Turkestan.The Annals and Magazine of Natural History, including Zoology, Botany, and Geology 18 (4): 377-388.
  • SIMPSON G. G. 1945. - The principles of classification and a classification of mammals. Bulletin of the American Museum of Natural History 85 (16): 1-350.
  • SMITH J. A. 1881. - Notice of remains of the Red Deer (Cervus elaphus, Linn.) found in the bed of an old loch near Dundas Castle, Linlithgowshire; with notes of remains of Red Deer found in different localities in the South of Scotland. Proceedings of the Society of Antiquaries of Scotland 15: 37-63.
  • SOMMER R. S. & NADACHOWSKI A. 2006. - Glacial refugia of mammals in Europe: evidence from fossil records. Mammal Review 36 (4): 251-265.
  • SOMMER R. S., ZACHOS F. E., STREET M., JORIS O., SKOG A. & BENECKE N. 2008. - Late Quaternary distribution dynamics and phylogeography of the red deer (Cervus elaphus) in Europe. Quaternary Science Reviews 27 (7-8): 714-733.
  • SONDAAR P. Y. 1977. - Insularity and its effect on mammal evolution, in HECHT M. K., GOODY P. C. & HECHT B. M. (eds), Major Patterns in Vertebrate Evolution. Springer US, Boston, MA: 671-707.https://doi.org/10.1007/978-1-4684-8851-7_23
  • SOKOLOV I. I. 1959. - Hoofed Animals (Orders Perissodactyla and Artiodactyla). Academy of Sciences of USSR, Moscow, Leningrad, 640 p. (Fauna of the USSR. Mammals 1.3) [in Russian].
  • SPENCER L. 1995. - Morphological correlates of dietary resource partitioning in the African Bovidae. Journal of Mammalogy 76 (2): 448-471. https://doi.org/10.2307/1382355
  • STANKOVIC A., DOAN K., MACKIEWICZ P., RIDUSH B., BACA M., GROMADKA R., SOCHA P., WEGLENSKI P., NADACHOWSKI A. & STEFANIAK K. 2011. - First ancient DNA sequences of the Late Pleistocene red deer (Cervus elaphus) from the Crimea, Ukraine. Quaternary International 245 (2): 262-267. https:// doi.org/10.1016/j.quaint.2011.05.023
  • STEPANOVA V. V. & ARGUNOV A. V. 2016. - Morphometry of antlers of red deer (Cervus elaphus L., 1758) of Yakutia. Bulletin of Udmurd University, Series Biology, Earth Sciences 26 (1): 126- 132 [in Russian].
  • STONEHOUSE B. 1968. -Thermoregulatory Function of Growing Antlers. Nature 218 (1): 870-872.https://doi.org/10.1038/218870a0
  • TAKAKUWA Y., NAKADA H. & HASEGAWA Y. 1999. - Reconstruction of a Miyako roe deer skeleton,Capreolus miyakoensis (Artiodactyla: Cervidae). Gunma Prefectural Museum of Natural History 3: 81-84
  • TARASOV P. P. 1956. - Some peculiarities of the morphology of the reindeer as a tundra animal. Proceedings of the Moscow Society of Naturalists (new series), Biological Section 61 (4): 80-82 [in Russian].
  • VAN VALEN L. 1976. - Ecological species, multispecies, and oaks. Taxon 25 (2/3): 233-239. https://doi.org/10.2307/1219444
  • VISLOBOKOVA I. A. 1990. - Fossil deer of Eurasia. Transactions of Paleontological Institute 240: 1-208 [in Russian].
  • WARD R. 1892. - Horn Measurements and Weights of the Great Game of the World. Rowland Ward & Co., London, 264 p.
  • WEBB S. D. 2000. - Evolutionary history of New World Cervidae, in VRBA E. S. & SCHALLER G. B. (eds), Antelopes, Deer, and Relatives: Fossil Record, Behavioral Ecology, Systematics, and Conservation. Yale University Press, New York, NY: 38-64.
  • ZDANSKY O. 1925. - Fossile Hirsche Chinas.Palaeontologia Sinica, Series C 2 (3): 1-94.
  • ZEUNER F. E. 1946. - Cervus elaphus jerseyensis, and other fauna in the 25-ft beach of Belle Hougue Cave, Jersey, C. I. Bulletin annuel de la Societe jersiaise 14: 238-254.