Published October 9, 2025 | Version v1

Nyctimus kinabaluensis Benjamin & Dhiya'ulhaq 2025, sp. nov.

  • 1. National Institute of Fundamental Studies, Hantana Road, Kandy 20000, Sri Lanka
  • 2. Animal Ecology, J. F. Blumenbach Institute of Zoology and Anthropology, University of Göttingen, Untere Karspüle 2, Göttingen 37073, Germany & Species Obscura, Jl. Duren 1 No. 2, Depok, Jawa Barat 16434, Indonesia
  • 3. Netherlands
  • 4. Department of Plant Protection, Faculty of Agriculture, IPB University, Kampus IPB Dramaga Bogor, Jl. Raya Dramaga, West Java 16680, Indonesia
  • 5. Animal Ecology, J. F. Blumenbach Institute of Zoology and Anthropology, University of Göttingen, Untere Karspüle 2, Göttingen 37073, Germany
  • 6. Animal Ecology, J. F. Blumenbach Institute of Zoology and Anthropology, University of Göttingen, Untere Karspüle 2, Göttingen 37073, Germany & Centre for Transdisciplinary and Sustainability Sciences, IPB University, Kampus IPB Baranangsiang, Jl. Raya Pajajaran No. 27, West Java 16127, Indonesia

Description

Nyctimus kinabaluensis Benjamin & Dhiya’ulhaq sp. nov.

Figs 5, 6, 7

Etymology.

The specific epithet is derived from the name of the type locality.

Type material.

Holotype. Malaysia – Sabah State • ♂; Kinabalu National Park, Poring Hot Springs; 05°59'N, 116°42'E; elev. 600–700 m; 27 Feb. 1996; A. Floren leg.; canopy fogging on Barringtonia tree, in primary forest; RMNH RMNH.ARA.17862.

Other material examined.

Brunei – Tutong District • 1 ♀; Lamunin, Bukit Sulang; 1982; N. Stork leg.; canopy fogging; RMNH RMNH.ARA.17859. Indonesia – Sumatra, Jambi Province • 1 ♀; Sarolangun, Bukit Duabelas National Park; 01°58'55.2"S, 102°45'02.6"E; elev. 73 m; 7 Oct. 2013; J. Drescher leg.; canopy fogging in rainforest; GOET 2013_BF 2.1_AraThom 004 N_001 (to be transferred to MZB).

Diagnosis.

Males can be easily distinguished from all other congeners by the embolic base abruptly bent 90 ° towards the retrolateral side of the cymbium and the tapered, bent embolus (Fig. 7 A, B). Females are distinguishable by the long, oval spermatheca (globular or short-oval in all other congeners; Figs 6 C, D, 7 C, D).

Description.

Male (holotype RMNH.ARA.17862; Figs 5 A, B, 7 A, B). Total length 3.24. Prosoma length 1.61; width 1.50. Opisthosoma length 1.63; width 1.31. Diameter of eyes: AME 0.11; ALE 0.20; PLE 0.17; PME 0.08. Interdistances between eyes: AME – AME 0.44; AME – ALE 0.11; ALE – ALE 0.81; PME – PME 0.45; PME – PLE 0.36; ALE – PLE 0.21; AME – PME 0.17; PLE – PLE 1.15. Leg measurements: leg I 5.0 (1.5, 0.6, 1.3, 1.0, 0.6). Prosoma cuboid, almost as long as wide, dark reddish brown; surface weakly granulated; laterally adorned with long macrosetae, each arising from a small tubercle. Chelicerae and sternum coloured as prosoma. Endites brown with white tip. Front legs brown; back legs brown except for the white femora. Opisthosoma oval, greyish brown with white spots (most dense in the anterior half) and paired white bars in the middle; anteriorly with a white border; five large sigillae present, arranged in a triangular formation.

Palp (Fig. 7 A, B): cymbium oval. Embolus elongated, tapering, with a 150 ° bend in the middle. Tegular hood small. RTA bilobed; ventral lobed very large, claw-shaped; dorsal lobe much smaller than ventral lobed, roughly triangular with an obtuse tip and a small triangular projection; VTA hook-shaped, small, attached medially on the RTA.

Female (RMNH.ARA.17859; Figs 5 C, 7 C, D). Total length: 3.38. Prosoma length 1.48; width 1.27. Opisthosoma length 1.90; width 1.70. Diameter of eyes: AME 0.07; ALE 0.17; PLE 0.14; PME 0.06. Interdistances between eyes: AME – AME 0.38; AME – ALE 0.12; ALE – ALE 0.74; PME – PME 0.39; PME – PLE 0.33; ALE – PLE 0.22; AME – PME 0.17; PLE – PLE 0.98. Leg measurements: leg I 4.6 (1.3, 0.4, 1.1, 1.0, 0.8). Habitus as in male.

Epigynum (Fig. 7 C, D): exterior plate large, roughly square with rounded sides. Atrium oval. CO diagonally oriented, anteriorly facing. CD arched. Spermatheca long oval, with a slight bump anteriorly.

Distribution.

Brunei; Malaysia (Borneo: Sabah); Indonesia (Sumatra: Jambi) (Fig. 19).

Notes

Published as part of Benjamin, Suresh P., Dhiya'ulhaq, Naufal Urfi, Deeleman-Reinhold, Christa, Buchori, Damayanti, Hidayat, Purnama, Scheu, Stefan & Drescher, Jochen, 2025, Taxonomic notes on the crab-spider genera Nyctimus Thorell, 1877 and Zametopina Simon, 1909 (Araneae, Thomisidae) with descriptions of six new species from Southeast Asia, pp. 95-125 in ZooKeys 1255 on pages 95-125, DOI: 10.3897/zookeys.1255.158380

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Linked records

Additional details

Biodiversity

Collection code
MZB , RMNH
Material sample ID
RMNH.ARA.17859 , RMNH.ARA.17862
Event date
1996-02-27 , 2013-10-07
Verbatim event date
1996-02-27 , 2013-10-07
Scientific name authorship
Benjamin & Dhiya'ulhaq
Kingdom
Animalia
Phylum
Arthropoda
Order
Araneae
Family
Thomisidae
Genus
Nyctimus
Species
kinabaluensis
Taxon rank
species
Taxonomic status
sp. nov.
Type status
holotype
Taxonomic concept label
Nyctimus kinabaluensis Benjamin & Dhiya'ulhaq, 2025