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Published July 15, 2025 | Version v1
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A new Maastrichtian hyposaurine dyrosaurid (Crocodylomorpha) from Namibe province, Angola

Authors/Creators

  • 1. GeoBioTec, Departamento de Ciências da Terra, FCT, Universidade Nova de Lisboa, Caparica 2829-516, Portugal
  • 2. Museo di Storia Naturale, Università di Pisa, Calci 56011, Italy
  • 3. Departamento de Ciencias de la Tierra, Facultad de Ciencias, Aragosaurus-IUCA Reconstrucciones Palaeoambientales, Universidad de Zaragoza, Zaragoza 50009, Spain
  • 4. GeoBioTec, Departamento de Ciências da Terra, FCT, Universidade Nova de Lisboa, Caparica 2829-516, Portugal & Departamento de Geodinámica, Estratigrafía y Palaeontología, Universidad Complutense de Madrid, Madrid 28040, Spain
  • 5. Departamento de Geologia, Faculdade de Ciências, Universidade Agostinho Neto, Luanda 374J + R8F, Angola

Description

Maréchal, Arthur, Rotatori, Filippo Maria, Merella, Marco, Puértolas-Pascual, Eduardo, Sequero, Cristina, Pereira, Ricardo, Nsungani, Pedro, Mateus, Octávio (2025): A new Maastrichtian hyposaurine dyrosaurid (Crocodylomorpha) from Namibe province, Angola. Zoological Journal of the Linnean Society 204 (4): 1-19, DOI: 10.1093/zoolinnean/zlaf092, URL: https://doi.org/10.1093/zoolinnean/zlaf092

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Identifiers

URL
https://www.checklistbank.org/dataset/312385
LSID
urn:lsid:plazi.org:pub:2D4DFFE3FFD85F67F709FF936D6B0A0D
URL
http://publication.plazi.org/id/2D4DFFE3FFD85F67F709FF936D6B0A0D

Has part
Taxonomic treatment: 10.5281/zenodo.17185176 (DOI)
Taxonomic treatment: http://treatment.plazi.org/id/D174879BFFDC5F6EF795FE2469A60E41 (URL)
Figure: 10.5281/zenodo.17185178 (DOI)
Figure: 10.5281/zenodo.17185180 (DOI)
Figure: 10.5281/zenodo.17185184 (DOI)
Figure: 10.5281/zenodo.17185190 (DOI)
Figure: 10.5281/zenodo.17185195 (DOI)
Figure: 10.5281/zenodo.17185197 (DOI)
Figure: 10.5281/zenodo.17185201 (DOI)
Figure: 10.5281/zenodo.17185203 (DOI)
Is source of
Dataset: https://www.gbif.org/dataset/d95c5d1d-e4fc-408e-ba62-2adf9ca69102 (URL)

References

  • Amoudji YZ, Guinot G, Hautier L et al. New data on the Dyrosauridae (Crocodylomorpha) from the Paleocene of Togo.Annales de Paleontologie 2021;107:102472.
  • Andrade RCLP, de Sayao JM. Paleohistology and lifestyle inferences of a dyrosaurid (Archosauria: Crocodylomorpha) from Paraiba Basin (Northeastern Brazil). PLoS One 2014;9:e102189.
  • Andrade MB, Edmonds R, Benton MJ et al. A new Berriasian species of Goniopholis (Mesoeucrocodylia, Neosuchia) from England, and a review of the genus. Zoological Journal of the Linnean Society 2011; 163:S66-S108.
  • Antunes MT, Cappetta H. Selaciens du Cretace (Albien-Maastrichtien) d'Angola.Palaeontographica Abteilung A 2002;264:85-146.
  • Arambourg C. Note preliminaire sur les vertebres fossiles des phosphates du Maroc. Bulletin de la Societe geologique de France 1935;5:413-39.
  • Arambourg C. Les vertebres fossiles des gisements de phosphates (Maroc-Algerie-Tunisie). Notes et Memoires du Service geologique du Maroc 1952;92:1-372.
  • Arambourg C, Magnier P. Gisements de vertebres dans le bassin tertiaire de Syrte (Libye).Comptes rendus hebdomadaires des seances de l'Academie des sciences 1961;252:1181-1183.
  • Araujo R, Goncalves AO, Jacobs LL et al. New aristonectine elasmosaurid plesiosaur specimens from the Early Maastrichtian of Angola and comments on paedomorphism in plesiosaurs.Netherlands Journal of Geosciences -Geologie en Mijnbouw 2015;94:93-108.
  • Argollo J, Buffetaut E, Cappetta H et al.Decouverte de vertebres aquatiques presumes paleocenes dans les Andes septentrionales de Bolivie (Rio Suches, synclinorium de Putina).Geobios 1987;20:123-27.
  • Barbosa JA,Kellner AWA, Viana, MS,S. New dyrosaurid crocodylomorph and evidences for faunal turnover at the K -P transition in Brazil. Proceedings of the Royal Society B: Biological Sciences 2008;275: 1385-1391.
  • Benton MJ, Clark JM. Archosaur phylogeny and the relationships of the Crocodylia.In: MJ Benton (ed.),The Phylogeny and Classification of the Tetrapods. Vol. 1: Amphibians, Reptiles, Birds. Oxford: Clarendon Press, 1988, 295-338.
  • Bergounioux FM. Les crocodiliens fossiles des depots phosphates du Sud-Tunisien. Comptes rendus hebdomadaires de seances de l'Academie des sciences 1955;240:1917-8.
  • Bergounioux FM. Les reptiles fossiles des depots phosphates sud tunisiens. Annales des Mines et de la Geologie 1956;15:1-105.
  • Brochu CA. Phylogenetic Approaches Toward Crocodylian History. Annual Review of Earth and Planetary Sciences 2003;31:357-97.
  • Brochu CA, Bouare ML,Sissoko F et al.A dyrosaurid crocodyliform braincase from Mali. Journal of Paleontology 2002;76:1060-71.
  • Bronzati M, Montefeltro FC, Langer MC. Diversification events and the effects of mass extinctions on Crocodyliformes evolutionary history. Royal Society Open Science 2015;2:140385.
  • Buffetaut E. Osteologie et affinites de Trematochampsa taqueti (Crocodylia, Mesosuchia) du Senonien inferieur d'in Beceten (Republique du Niger).Geobios 1976;9:143-98.
  • Buffetaut E. A dyrosaurid (Crocodylia, Mesosuchia) from the Upper Eocene of Burma. Neues Jahrbuch fur Geologie und Palaontologie, Monatshefte, 1978a;273-81.
  • Buffetaut E. Les Dyrosauridae (Crocodylia, Mesosuchia) des phosphates de l'Eocene inferieur de Tunisie : Dyrosaurus,Rhabdognathus, Phosphatosaurus. Geologie Mediterraneenne 1978b;5:237-55.
  • Buffetaut E. Sokotosuchus ianwilsoni and the evolution of the dyrosaurid crocodilians.The Nigerian Field Monograph 1979a;1:31-41.
  • Buffetaut E. Presence du crocodilien Phosphatosaurus (Mesosuchia, Dyrosauridae) dans le paleocene du Niger et du Mali. Palaontologische Zeitschrift 1979b;53:323-33.
  • Buffetaut E. Les crocodiliens paleogenes du Tilemsi (Mali): un apercu systematique. In: Palaeovertebrata, Memoire Jubilaire Rene Lavocat. Montpellier: Laboratoire de Paleontologie des Vertebres de l'Ecole Pratique des Hautes Etudes, 1980, 15-35.
  • Buffetaut E. Radiation evolutive, paleoecologie et biogeographie des crocodiliens mesosuchiens. Memoires de la Societe geologique de France 1982;142:1-88.
  • Buffetaut E, Wouters G. Atlantosuchus coupatezi n.g., n.sp., un nouveau Dyrosauride (Crocodylia, Mesosuchia) des phosphates Montiens du Maroc. Bulletin trimestriel de la Societe Geologique de Normandie et des Amis du Museum du Havre 1979;66:85-90.
  • Buffetaut E, Bussert R, Brinkmann W. A new nonmarine vertebrate fauna in the Upper Cretaceous of northern Sudan.Berliner Geowissenschaftliche Abhandlungen 1990;120:183-202.
  • Callahan WR, Pellegrini R, Schein JP et al.A nearly complete specimen of Hyposaurus rogersii (Crocodylomorpha, Dyrosauridae) from the Late Cretaceous -Early Paleogene of New Jersey. Journal of Vertebrate Paleontology, Program and Abstracts, 2015;2015:101.
  • Case GR, Cappetta H. A new Selachian fauna from the late Maastrichtian of Texas (Upper Cretaceous/Navarroan;Kemp Formation).Munchner Geowissenschaftliche Abhandlungen A 1997;34:131-89.
  • Chiarenza AA, Farnsworth A, Mannion PD et al.Asteroid impact, not volcanism, caused the end-Cretaceous dinosaur extinction.Proceedings of the National Academy of Sciences of the United States of America 2020;117: 17084-93.
  • Churcher CS, Russell DA. Terrestrial vertebrates from Campanian strata in Wadi El-Gedid (Kharga and Dahkleh Oases), Western Desert of Egypt. Journal of Vertebrate Paleontology 1992;12:23A.
  • Cooper MR. The Mid-Cretaceous (Albian/Turonien) Biostratigraphy of Angola. Annales du Museum d'Histoire Naturelle de Nice 1976;4: XVI.1-.22.
  • Cope ED. A contribution to the vertebrate paleontology of Brazil. Paleontological Bulletin 1885;40:1-20.
  • Costa B, Fialho P, Balbino A. Occurrence of odontaspidid sharks (Chondrichthyes,Neoselachii) from the Cretaceous of Bentiaba,Angola.Paleo-Fall Meeting 2019;2019:1.
  • Denton RK, Dobie JL, Parris DC. The Marine Crocodilian Hyposaurus in North America. In: JM Callaway, EL Nicholls (eds), Ancient Marine Reptiles. San Diego: Academic Press, 1997, 375-397.
  • Dollo L.Sur la decouverte de teleosauriens tertiaires au Congo.Bulletin de l'Academie royale de Belgique 1914;7:288-98.
  • Dubicka Z, Wierny W, Bojanowski MJ et al. Multi-proxy record of the mid-Maastrichtian event in the European Chalk Sea: Paleoceanographic implications. Gondwana Research 2024;129:1-22.
  • Ehret D, Hastings A. An Eocene occurrence of a dyrosaurid (Crocodylomorpha, Mesoeucrocodylia) from Alabama,USA.Journal of Vertebrate Paleontology, Program and Abstracts 2013;2013:121.
  • Erb A, Turner AH. Braincase anatomy of the Paleocene crocodyliform Rhabdognathus revealed through high resolution computed tomography.PeerJ 2021;9:e11253.
  • Fernandes AE,Mateus O, Andres B et al.Pterosaurs from the late cretaceous of Angola. Diversity 2022;14:741.
  • Foret T, Aubier P, Jouve S et al. Biotic and abiotic factors and the phylogenetic structure of extinction in the evolution of tethysuchia.Paleobiology 2024;50:285-307.
  • Fortier D, Perea D, Schultz C.Redescription and phylogenetic relationships of Meridiosaurus vallisparadisi, a pholidosaurid from the Late Jurassic of Uruguay. Zoological Journal of the Linnean Society 2011;163: S257-S272.
  • Gayet M, Marshall LG, Sempere T.The Mesozoic and Paleocene vertebrates of Bolivia and their stratigraphic context:a review.Revista Tecnica Yacimientos Petroliferos Fiscales Bolivianos Santa Cruz, Bolivia 1991;12:393-433.
  • Gayet M, Sempre T, Cappetta H et al. La presence de fossiles marins dans le Cretace terminal des Andes centrales et ses consequences paleogeographiques.Palaeogeography,Palaeoclimatology,Palaeoecology 1993;102: 283-319.
  • Granot R, Dyment J.The Cretaceous opening of the South Atlantic Ocean. Earth and Planetary Science Letters 2015;414:156-63.
  • Guinot G,Condamine FL.Global impact and selectivity of the Cretaceous - Paleogene mass extinction among sharks, skates, and rays.Science (New York, N.Y.) 2023;379:802-6.
  • Guinot G, Hautier L, Sambou BS et al.The upper cretaceous elasmobranch fauna from Senegal. Cretaceous Research 2023;146:105480.
  • Goloboff PA, Farris JS, Nixon KC. TNT, a free program for phylogenetic analysis. Cladistics 2008;24:774-86.
  • Halstead LB. Sokotosuchus ianwilsoni n.g., n.sp., a new teleosaur crocodile from the upper cretaceous of Nigeria. Journal of Mining and Geology 1975;11:101-3.
  • Hastings AK, Bloch JI, Cadena EA et al. A new small short-snouted dyrosaurid (Crocodylomorpha, Mesoeucrocodylia) from the Paleocene of northeastern Colombia. Journal of Vertebrate Paleontology 2010;30: 139-62.
  • Hastings AK, Bloch JI,Jaramillo CA.A new longirostrine dyrosaurid (Crocodylomorpha,Mesoeucrocodylia) from the Paleocene of northeastern Colombia:biogeographic and behavioural implications for New-World Dyrosauridae. Palaeontology 2011;54:1095-116.
  • Hastings AK, Bloch JI, Jaramillo CA. A new blunt-snouted dyrosaurid, Anthracosuchus balrogus gen.et sp. nov.(Crocodylomorpha, Mesoeucrocodylia), from the Palaeocene of Colombia. Historical Biology 2015;27:998-1020.
  • Hay OP, 1930. Second Bibliography and Catalogue of the Fossil Vertebrata of North America, Vol. 2. Washington, DC: Carnegie Institution of Washington.
  • Hill RV, McCartney JA, Roberts E et al. Dyrosaurid (Crocodyliformes: Mesoeucrocodylia) fossils from the Upper Cretaceous and Paleogene of Mali: implications for phylogeny and survivorship across the K/T boundary.American Museum Novitates 2008;3631:1-19.
  • Jouve S.A new description of the skull of Dyrosaurus phosphaticus (Thomas, 1893) (Mesoeucrocodylia: Dyrosauridae) from the Lower Eocene of North Africa.Canadian Journal of Earth Sciences 2005;42:323-37.
  • Jouve S. Taxonomic revision of the dyrosaurid assemblage (Crocodyliformes: Mesoeucrocodylia) from the Paleocene of the Iullemmeden Basin, West Africa.Journal of Paleontology 2007;81:163-75.
  • Jouve S. Differential diversification through the K -Pg boundary, and post-crisis opportunism in longirostrine crocodyliforms. Gondwana Research 2021;99:110-30.
  • Jouve S, Jalil NE.Paleocene resurrection of a crocodylomorph taxon:biotic crises, climatic and sea-level fluctuations. Gondwana Research 2020;85: 1-18.
  • Jouve S, Schwarz D.Congosaurus bequaerti, a Paleocene dyrosaurid (Crocodyliformes; Mesoeucrocodylia) from Landana (Angola). Bulletin.de l'Institut Royal des Sciences Naturelles de Belgique, Sciences de la Terre 2004;74:129-46.
  • Jouve S, Bouya B, Amaghzaz M. A short-snouted dyrosaurid (Crocodyliformes, Mesoeucrocodylia) from the Paleocene of Morocco. Palaeontology 2005a;48:359-69.
  • Jouve S, Iarochene M, Bouya B, A, M. A new dyrosaurid crocodyliform from the Paleocene of Morocco and a phylogenetic analysis of Dyrosauridae. Acta Palaeontologica Polonica 2005b;50:581-94.
  • Jouve S, Iarochene M, Bouya B et al. A new species of Dyrosaurus (Crocodylomorpha, Dyrosauridae) from the Early Eocene of Morocco: phylogenetic implications.Zoological Journal of the Linnean Society 2006;148: 603-56.
  • Jouve S, Bouya B, Amaghzaz M. A long-snouted dyrosaurid (Crocodyliformes, Mesoeucrocodylia) from the Paleocene of Morocco: Phylogenetic and palaeobiogeographic implications. Palaeontology 2008;51: 281-94.
  • Jouve S, de Muizon C, Cespedes-Paz R et al. The longirostrine crocodyliforms from Bolivia and their evolution through the Cretaceous -Palaeogene boundary. Zoological Journal of the Linnean Society 2021;192: 475-509.
  • Jouve S, Rodriguez-Jimenez JV. The youngest known South American dyrosaurid (Late Paleocene of Colombia), and evolution of Dyrosauridae (Crocodyliformes: Tethysuchia). Geodiversitas 2023;46: 931-53.
  • von Koken E. Die Dinosaurier. Crocodiliden und Sauropterygier des norddeutschen Wealden. Palaeontologische Abhandlungen 1887;3:309-419.
  • Khosla A, Sertich JJ, Prasad GV et al. Dyrosaurid remains from the intertrappean beds of India and the Late Cretaceous distribution of Dyrosauridae. Journal of Vertebrate Paleontology 2009;29:1321-6.
  • Kuhn O.Die Reptilien:System und Stammesgeschichte. Krailing bei Munchen: Oeben, 1966, 154.
  • Lamanna MC, Smith JB, Attia YS et al. From dinosaurs to dyrosaurids (Crocodyliformes): removal of the post-Cenomanian (Late Cretaceous) record of Ornithischia from Africa.Journal of Vertebrate Paleontology 2004;24:764-8.
  • Langston W. Dyrosaurs (Crocodilia, Mesosuchia) from the Paleocene Umm Himar Formation,Kingdom of Saudi Arabia.Paleocene Vertebrates from Jabal Umm Himar, Kingdom of Saudi Arabia: US Geological Survey Bulletin 1995;2093-F:F1-F36.
  • Lewis PO. A likelihood approach to estimating phylogeny from discrete morphological character data.Systematic Biology 2001;50:913-25.
  • Martin JE, Amiot R, Lecuyer C et al. Sea surface temperature contributes to marine crocodylomorph evolution.Nature Communications 2014;5: 4658.
  • Martin JE, Sarr R, Hautier L. A dyrosaurid from the Paleocene of Senegal. Journal of Paleontology 2019;93:343-58.
  • Marx MP,Mateus O, Polcyn MJ et al.The cranial anatomy and relationships of Cardiocorax mukulu (Plesiosauria: Elasmosauridae) from Bentiaba, Angola.PLoS One 2021;16:e0255773.
  • Mateus O, Callapez PM, Polcyn MJ, Schulp AS, Goncalves AO, Jacobs LL. The fossil record of biodiversity in Angola through time:a paleontological perspective. In: Biodiversity of Angola: Science and Conservation: A Modern Synthesis. Cham, Switzerland: Springer, 2019, 53-76.
  • Moody R, TJ,, Buffetaut E. Notes on the systematics and palaeoecology of the crocodiles and turtles of the Metlaoui Phosphates (Eocene) of southern Tunisia. Tertiary Research 1981;3:125-40.
  • Morales ME, Goloboff PA.New TNT routines for parallel computing with MPI. Molecular Phylogenetics and Evolution 2023;178:107643.
  • Nopcsa BF. The genera of reptiles. Palaeobiologica 1928;1:163-88.
  • Norell MA,Storrs GW. Catalogue and review of the type fossil crocodilians in the Yale Peabody Museum. Postilla 1989;N203:1-28.
  • O'Leary MA, Bouare ML, Claeson KM et al.Stratigraphy and paleobiology of the Upper Cretaceous -Lower Paleogene sediments from the trans-Saharan seaway in Mali. Bulletin of the American Museum of Natural History 2019;2019:1-183.
  • Ortega F, Gasparini ZB, Buscalioni AD et al. A new species of Araripesuchus (Crocodylomorpha, Mesoeucrocodylia) from the Lower Cretaceous of Patagonia (Argentina).Journal of Vertebrate Paleontology 2000;20:57-76.
  • Owen R. Notes on remains of fossil reptiles discovered by Prof. Henry Rodgers of Pennsylvania, U.S., in Greensand Formations of New Jersey. Quarterly Journal of the Geological Society of London 1849;5:380-3.
  • Payne ARD, Mannion PD, Lloyd GT et al. Decoupling speciation and extinction reveals both abiotic and biotic drivers shaped 250 million years of diversity in crocodile-line archosaurs.Nature Ecology & Evolution 2024;8:121-32.
  • Pol, D., and Gasparini, Z.B.. Crocodyliformes. In: ZB Gasparini, L Salgado, RA Coria (eds), Patagonian Mesozoic Reptiles (Life in the Past). Bloomington: Indiana University Press, 2007,116-142.
  • Polcyn MJ, Jacobs LL, Schulp AS et al. The North African mosasaur Globidens phosphaticus from the Maastrichtian of Angola.Historical Biology 2010;22:175-85.
  • Polcyn MJ,Jacobs LL, Araujo R et al.Physical drivers of mosasaur evolution. Palaeogeography, Palaeoclimatology, Palaeoecology 2014;400:17-27.
  • Pomel A. Decouverte de champsosauriens dans les gisements de phosphorite du suessonien de l'Algerie. Comptes Rendus de l'Academie des Sciences 1894;118:1309-10.
  • Prasad GVR, Singh V. Microvertebrates from the intertrappean beds of Rangareddi District, Andhra Pradesh and their biostratigraphic significance. Bulletin of the Indian Geological Association 1991;24:1-20.
  • R Core Team. R: A Language and Environment for Statistical Computing. Vienna: R Foundation for Statistical Computing, 2023.
  • Rambaut A. FigTree, 2018.
  • Rambaut A, Drummond AJ,Xie D et al.Posterior summarization in Bayesian phylogenetics using Tracer 1.7. Systematic Biology 2018;67:901-4.
  • Rana RS.Dyrosaurid crocodile (Mesosuchia) from the infratrappean beds of Vikarabad, Hyderabad District, Andhra Pradesh. Current Science 1987;56:532-4.
  • Rasmussen AR, Murphy JC, Ompi M et al. Marine reptiles. PLoS One 2011;6:e27373.
  • Rio JP, Mannion PD.Phylogenetic analysis of a new morphological dataset elucidates the evolutionary history of Crocodylia and resolves the long-standing gharial problem. PeerJ 2021;9:e12094.
  • Ronquist F, Teslenko M, van der Mark P et al. MrBayes 3.2:efficient Bayesian phylogenetic inference and model choice across a large model space. Systematic Biology 2012;61:539-42.
  • Saber S, Salem BS, Ouda K et al. A long-snouted dyrosaurid (Crocodyliformes, Mesoeucrocodylia) from the Campanian Quseir Formation of Egypt. Cretaceous Research 2025;165:105982.
  • Salih KAO, Evans DC, Bussert R et al. First record of Hyposaurus (Dyrosauridae, Crocodyliformes) from the Upper Cretaceous Shendi Formation of Sudan.Journal of Vertebrate Paleontology 2016;36:e1115408.
  • Salih KAO, Evans DC, Bussert R et al. Brachiosuchus kababishensis, a new long-snouted dyrosaurid (Mesoeucrocodylia) from the Late Cretaceous of North Central Sudan. Historical Biology 2022;34:821-40.
  • Schwarz D, Raddatz M, Wings O.Knoetschkesuchus langenbergensis gen.nov. sp.nov., a new atoposaurid crocodyliform from the Upper Jurassic Langenberg Quarry (Lower Saxony, northwestern Germany), and its relationships to Theriosuchus. PLoS One 2017;12:e0160617.
  • Schwarz D, Salisbury SW. A new species of Theriosuchus (Atoposauridae, Crocodylomorpha) from the Late Jurassic (Kimmeridgian) of Guimarota, Portugal. Geobios 2005;38:779-802.
  • Schwarz-Wings D, Frey E, Martin T. Reconstruction of the bracing system of the trunk and tail in hyposaurine dyrosaurids (Crocodylomorpha; Mesoeucrocodylia).Journal of Vertebrate Paleontology 2009;29:453-72.
  • Scotese CR, Verard C, Burgener L et al.The cretaceous world:plate tectonics, palaeogeography and palaeoclimate. Geological Society, London, Special Publications 2025;544:31-202.
  • Sertich J, Manthi FK, Sampson F et al. Rift valley dinosaurs: a new Late Cretaceous vertebrate fauna from Kenya. Journal of Vertebrate Paleontology 2006;26:124A.
  • Simoes TR, Greifer N, Barido-Sottani J et al. EvoPhylo: an r package for pre- and postprocessing of morphological data from relaxed clock Bayesian phylogenetics. Methods in Ecology and Evolution 2023;14: 1981-93.
  • Souza RG, Figueiredo RG, Azevedo SAK et al. Systematic revision of Sarcosuchus hartti (Crocodyliformes) from the Reconcavo Basin (Early Cretaceous) of Bahia, northeastern Brazil.Zoological Journal of the Linnean Society 2019;188:552-78.
  • Stadler T.Sampling-through-time in birth -death trees.Journal of Theoretical Biology 2010;267:396-404.
  • de Stefano G. Nuovi rettili degli strati a fosfato della Tunisia.Bolletino della Societa Geologia Italiana 1903;22:51-80.
  • Strganac C, Salminen J, Jacobs LL et al. Carbon isotope stratigraphy, magnetostratigraphy, and 40Ar/39Ar age of the Cretaceous South Atlantic coast, Namibe Basin, Angola. Journal of African Earth Sciences 2014;99:452-62.
  • Strganac C, Jacobs LL, Polcyn MJ et al. Stable oxygen isotope chemostratigraphy and paleotemperature regime of mosasaurs at Bentiaba, Angola. Netherlands Journal of Geosciences -Geologie en Mijnbouw 2015a;94:137-43.
  • Strganac C, Jacobs LL,Polcyn MJ et al.Geological setting and paleoecology of the Upper Cretaceous Bench 19 Marine Vertebrate Bonebed at Bentiaba,Angola.Netherlands Journal of Geosciences -Geologie en Mijnbouw 2015b;94:121-36.
  • Swinton WE. On fossil reptilia from Sokoto province. Bulletin of the Geological Survey of Nigeria 1930;13:9-56.
  • Swinton WE. On Congosaurus bequaerti Dollo. Annale du Musee du Congo Belge 1950;13:9-56.
  • Tessier F. Contributions a la stratigraphie et a la paleontologie de la partie Ouest du Senegal (Cretace et Tertiaire).Gouvernement General de l'Afrique Occidentale Francaise.Bulletin de la Direction des Mines 1952;14:7-569.
  • Thevenin A. Le Dyrosaurus des phosphates de Tunisie. Annales de Paleontologie 1911;7:95-108.
  • Troxell EL. Hyposaurus, a marine crocodilian. American Journal of Science 1925;s5-9:489-514.
  • Walker AD. A revision of the Jurassic reptile Hallopus victor (Marsh), with remarks on the classification of crocodiles.Philosophical Transactions of the Royal Society of London, Series B, Biological Sciences 1970;257: 323-72.
  • Whetstone KN, Whybrow PJ.A 'cursorial' crocodilian from the Triassic of Lesotho (Basutoland), southern Africa.Occasional Papers of the Museum of Natural History. The University of Kansas 1983;106:1-37.
  • Wilberg EW.What's in an outgroup? The impact of outgroup choice on the phylogenetic position of Thalattosuchia (Crocodylomorpha) and the origin of Crocodyliformes.Systematic Biology 2015;64:621-37.
  • Wilberg EW, Turner AH, Brochu CA. Evolutionary structure and timing of major habitat shifts in Crocodylomorpha.Scientific Reports 2019;9:514.
  • Young MT, Andrade MB. What is Geosaurus? Redescription of G. giganteus (Thalattosuchia, Metriorhynchidae) from the Upper Jurassic of Bayern, Germany.Zoological Journal of the Linnean Society 2009;157:551-85.
  • Young MT,Andrade MB, Cornee J-J et al.Re-description of a putative Early Cretaceous 'teleosaurid' from France, with implications for the survival of metriorhynchids and teleosaurids across the Jurassic-Cretaceous Boundary. Annales de Paleontologie 2014;100:165-74.
  • Young MT, Hastings AK, Allain R et al. Revision of the enigmatic crocodyliform Elosuchus felixi de Lapparent de Broin, 2002 from the Lower - Upper Cretaceous boundary of Niger: potential evidence for an early origin of the clade Dyrosauridae.Zoological Journal of the Linnean Society 2016;179:377-403.