Omoplax desecta
Authors/Creators
- 1. Shirakami Research Center for Environmental Sciences, Faculty of Agriculture and Life Science, Hirosaki University, Aomori, Japan
Description
Omoplax desecta (Horváth, 1912)
Figs 1 B, 2 B, 3 B, 4 B, 5 B, 6 B, 7 B, 8 B, 9 B, 10 B, 11 B, 12 B, 13 B, 14 B, 15 F – H
Stephanitis (Omoplax) desecta Horváth, 1912: 337. Syntype (s): ♀; type locality: Japan • Ogasawara [= Ogasawara Islands]; Laboratory of Systematic Entomology, Faculty of Agriculture, Hokkaido University, Sapporo, Japan (Tomokuni 1994: 844; Souma 2022 a: 125).
Omoplax desecta: Takeya (1962: 74) (new combination).
References.
Drake (1948: 55) (checklist: genus); Takeya (1951: 14) (checklist: eastern Asia); Drake and Maa (1953: 100) (checklist: genus); Drake (1956: 116) (distribution); Drake and Ruhoff (1960: 83) (catalog); Drake and Ruhoff (1965: 336) (catalog); Nakane (1970: 28) (checklist: Ogasawara Islands); Miyamoto and Yasunaga (1989: 167) (checklist: Japan); Nishimura and Arai (1989: 38) (distribution); Kato (1992: 79) (checklist: Ogasawara Islands); Yasunaga et al. (1993: 179) (monograph); Tomokuni (1994: 844) (type material); Yasunaga (1995: 20) (illustration); Péricart and Golub (1996: 51) (checklist: Palearctic); Miyano (1998: 8) (distribution); Guilbert (2001: 551) (distribution); Karube and Takakuwa (2004: 83) (illustration); Ohbayashi et al. (2004: 32) (checklist: Ogasawara Islands); Government of Japan (2010: 208) (checklist: Ogasawara Islands); Yamada and Tomokuni (2012: 198) (monograph); Takahashi et al. (2014: 163) (natural enemy); Yamada and Ishikawa (2016: 432) (checklist: Japan); Souma and Kamitani (2021: 8) (distribution: part); Souma (2022 a: 125) (distribution: part); Shimamoto and Ishikawa (2023: 93) (catalog: part); Souma (2023: 9) (monograph). Several studies conducted before the 1980 s recorded this species as Stephanitis (Omoplax) desecta (Drake 1948; Takeya 1951; Drake and Maa 1953; Drake 1956; Drake and Ruhoff 1960, 1965; Nakane 1970; Nishimura and Arai 1989).
Material examined.
Non-types, Japan • 1 ♂; Ogasawara Isls, Chichijima Is.; 6 May 1974; Y. Hori leg.; NSMT • 1 ♀; same locality data as for preceding; Rhaphiolepis indica var. tashiroi; 20 Mar. 2001; G. Tokihiro; NSMT • 1 ♀; Ogasawara Isls, Hahajima Is., Higashiko; 5 Jun. 1976; T. Nakane leg.; NSMT • 1 ♀; Ogasawara Isls, Hahajima Is., Mt. Chibusa; 8 Jun. 1976; T. Nakane leg.; NSMT • 1 ♀; same locality data as for preceding; 6 Jul. 1997; T. Kishimoto leg; NSMT • 2 ♀♀; same locality data as for preceding; 7 Jul. 1997; K. Matsumoto leg; NSMT • 1 ♀; same locality data as for preceding; 27 May 2022; T. Yoshida leg.; SIHU • 2 ♂♂ 5 ♀♀; same locality data as for preceding; Rhaphiolepis indica var. tashiroi; 2 Oct. 2024; J. Souma; SIHU • 1 ♂; Ogasawara Isls, Chichijima Is., Mt. Mikazuki; 12 Jun. 1976; T. Nakane leg.; NSMT • 23 ♂♂ 18 ♀♀ 1 fifth instar nymph; same locality data as for preceding; 18 Apr. 1997; K. Matsumoto leg.; NSMT • 1 ♀; same locality and date data as for preceding; T. Kishimoto leg.; NSMT • 1 ♂ 3 ♀♀; same locality data as for preceding; 23 Apr. 1997; K. Matsumoto leg.; NSMT • 2 ♂♂ 6 ♀♀; same locality data as for preceding; 17 May 2024; Y. Hisasue leg.; SIHU • 1 ♂ 1 ♀ 1 fifth instar nymph; same locality data as for preceding; Rhaphiolepis indica var. tashiroi; 21 Sep. 2024; J. Souma; SIHU • 2 ♀♀; same locality, host plant, and collector data as for preceding; 22 Sep. 2024; SIHU • 2 ♀♀; same locality data as for preceding; 25 Oct. 2024; Y. Hisasue; SIHU • 1 ♀; same locality and collector data as for preceding; 5 Nov. 2024; SIHU • 1 ♀; same locality and collector data as for preceding; 10 Jan. 2025; SIHU • 3 ♂♂ 4 ♀♀; same locality and collector data as for preceding; 15 May 2025; SIHU • 1 ♂ 1 ♀; Ogasawara Isls, Chichijima Is., Mt. Shigure; 8 Dec. 1977; M. Tomokuni leg.; NSMT • 1 ♀; Ogasawara Isls, Chichijima Is., Mt. Takayama; 30 Jul. 1996; K. Matsumoto leg.; NSMT • 2 ♀♀; Ogasawara Isls, Hahajima Is., Kuwanoki-zawa; 4 Aug. 1996; K. Matsumoto leg.; NSMT • 1 ♂; Ogasawara Isls, Chichijima Is., Higashidaira; 15 Apr. 1997; K. Matsumoto leg.; NSMT • 14 ♂♂ 8 ♀♀ 3 fifth instar nymphs 1 fourth instar nymph; Ogasawara Isls, Chichijima Is., Buta Beach; 16 Apr. 1997; K. Matsumoto leg.; NSMT • 7 ♂♂ 4 ♀♀; same locality and date data as for preceding; T. Kishimoto leg.; NSMT • 1 ♀; Ogasawara Isls, Chichijima Is., Nagatani – Mt. Tsutsuji; 17 Apr. 1997; K. Matsumoto leg; NSMT • 18 ♂♂ 14 ♀♀; Ogasawara Isls, Hahajima Is., Minamizaki; 19 Apr. 1997; K. Matsumoto leg; NSMT • 1 ♂; same locality data as for preceding; 4 Jul. 1997; T. Kishimoto leg; NSMT • 1 ♂; same locality data as for preceding; 15 Jul. 2024; Y. Uehara leg.; SIHU • 1 ♀; same locality data as for preceding; Rhaphiolepis indica var. tashiroi; 27 Sep. 2024; J. Souma; SIHU • 1 ♂ 3 ♀♀; Ogasawara Isls, Hahajima Is., Nakanotaira – Minamizaki; 19 Apr. 1997; K. Matsumoto leg; NSMT • 1 ♂; Ogasawara Isls, Hahajima Is., Mt. Mikazuki (non-existent place name: mislabeling?); 19 Apr. 1997; K. Matsumoto leg; NSMT • 1 ♂ 1 ♀; Ogasawara Isls, Hahajima Is., Komoridani; 20 Apr. 1997; K. Matsumoto leg; NSMT • 1 ♂ 1 ♀; Ogasawara Isls, Hahajima Is., Mt. Yakeyama; 20 Apr. 1997; K. Matsumoto leg; NSMT • 3 ♂♂; Ogasawara Isls, Hahajima Is., Oki-mura; 20 Apr. 1997; K. Matsumoto leg; NSMT • 1 ♀; same locality and date data as for preceding; T. Kishimoto leg; NSMT • 1 ♀; Ogasawara Isls, Chichijima Is., Minamifukurozawa; 23 Apr. 1997; K. Matsumoto leg; NSMT • 3 ♀♀ 1 fifth instar nymph; same locality data as for preceding; 23 Mar. 2024; Y. Hisasue leg.; SIHU • 1 ♀; Ogasawara Isls, Chichijima Is., Ichinotani; 26 Apr. 1997; K. Matsumoto leg; NSMT • 1 ♂; Ogasawara Isls, Ototojima Is., Kurohama; 27 Apr. 1997; K. Matsumoto leg; NSMT • 2 ♂♂; Ogasawara Isls, Hahajima Is., Nishiura; 4 Jul. 1997; K. Matsumoto leg; NSMT • 1 ♀; Ogasawara Isls, Hahajima Is., Mt. Kuwanoki; 4 Jul. 1997; T. Kishimoto leg; NSMT • 1 ♂ 1 ♀; Ogasawara Isls, Hahajima Is., Tamagawa Dam; 5 Jul. 1997; K. Matsumoto leg.; NSMT • 1 ♀; same locality data as for preceding; 14 Jul. 2024; Y. Uehara leg.; SIHU • 1 ♀; Ogasawara Isls, Chichijima Is., Mt. Tsutsuji; 10 Jul. 1997; T. Kishimoto leg.; NSMT • 1 ♂; Ogasawara Isls, Ototojima Is., Ainosawa; 1 Jul. 2021; T. Matsumoto & S. Shimamoto leg.; SIHU • 6 ♂♂ 4 ♀♀; same locality data as for preceding; 3–4 Jul. 2024; N. Tsuji leg.; SIHU • 2 ♀♀; Ogasawara Isls, Anijima Is., Central Plateau; 1 Jul. 2021; T. Matsumoto & S. Shimamoto leg.; SIHU • 1 ♂; same locality and collector data as for preceding; 2 Jul. 2021; SIHU • 1 ♀; Ogasawara Isls, Anijima Is., Mt. Togari; 29 Jul. 2021; T. Matsumoto & S. Shimamoto leg.; SIHU • 1 ♂; Ogasawara Isls, Hahajima Is., Igumadani, Sekimon; alt. 370 m; 26 May 2022; T. Yoshida leg.; SIHU • 2 ♂♂; Ogasawara Isls, Chichijima Is., Susaki; 21 May 2023; Y. Hisasue leg.; SIHU • 1 ♀; Ogasawara Isls, Chichijima Is., Mt. One; 4 Feb. 2024; Y. Hisasue leg.; SIHU • 2 ♂♂ 1 ♀; Ogasawara Isls, Meijima Is.; 14 Jun. 2024; N. Tsuji leg.; SIHU • 1 ♂; Ogasawara Isls, Hahajima Is., Nishidai; 17 Jun. 2024; N. Tsuji leg.; SIHU • 1 ♂ 5 ♀♀; Ogasawara Isls, Hahajima Is., Shizukazawa; 20 Jun. 2024; Y. Hisasue leg.; SIHU • 7 ♂♂ 6 ♀♀; Ogasawara Isls, Chichijima Is., Mt. Ogami; alt. 10–90 m; 11 Jul. 2024; Y. Uehara leg.; SIHU • 1 ♀; Ogasawara Isls, Ototojima Is., Mt. Sokuryogatake; 12 Jul. 2024; Y. Uehara leg.; SIHU • 1 ♂ 1 ♀; Ogasawara Isls., Anijima Is., Mansaku – Kanaimisaki; 12 Jul. 2024; N. Tsuji leg.; SIHU • 1 ♀; Ogasawara Isls, Hahajima Is., Nijuccho Pass; alt. 160 m; 15 Jul. 2024; Y. Uehara leg.; SIHU • 5 ♂♂ 1 ♀; Ogasawara Isls, Mukohjima Is.; alt. 0–90 m; 16 Jul. 2024; Y. Uehara leg.; SIHU • 1 ♀; Ogasawara Isls, Chichijima Is., Mt. Tsuitate; 19 Jul. 2024; Y. Hisasue leg.; SIHU • 1 ♀; Ogasawara Isls, Chichijima Is., Hatsuneura; 28 Jul. 2024; N. Tsuji leg.; SIHU • 1 ♂ 1 ♀; Ogasawara Isls, Chichijima Is., Higashi-machi; Calophyllum inophyllum; 21 Sep. 2024; J. Souma leg.; SIHU • 1 ♀; same locality, host plant, and collector data as for preceding; 8 Oct. 2024; SIHU • 1 ♀; Ogasawara Isls, Chichijima Is., Mt. Yoake; Rhaphiolepis indica var. tashiroi; 22 Sep. 2024; J. Souma leg.; SIHU • 1 ♂ 1 ♀; Ogasawara Isls., Ototojima Is., Kurohama – Ichinotani; Rhaphiolepis indica var. tashiroi; 23 Sep. 2024; J. Souma leg.; SIHU • 3 ♂♂ 1 ♀; Ogasawara Isls., Anijima Is., Tamana Beach – Mt. Mikaeri; Rhaphiolepis indica var. tashiroi; 24 Sep. 2024; J. Souma leg.; SIHU • 1 ♀; Ogasawara Isls, Chichijima Is., Ogamiyama Park; Rhaphiolepis indica var. tashiroi; 25 Sep. 2024; J. Souma leg.; SIHU • 1 ♂; Ogasawara Isls, Chichijima Is., John Beach; Calophyllum inophyllum; 7 Oct. 2024; J. Souma leg.; SIHU • 2 ♀♀; Ogasawara Isls, Nishijima Is.; 17 Nov. 2024; N. Tsuji leg.; SIHU • 1 ♂; Ogasawara Isls, Chichijima Is., Oku-mura; 10 Dec. 2024; Y. Hisasue leg.; SIHU • 1 ♀; Ogasawara Isls, Hahajima Is., Hyogidaira; Rhaphiolepis indica var. tashiroi; 9 Mar. 2025; J. Souma leg.; SIHU • 1 ♀; same locality, host plant, and collector data as for preceding; 11 Mar. 2025; SIHU • 1 ♂; Ogasawara Isls, Chichijima Is., Kominato Beach; 25 Apr. 2025; Y. Hisasue leg.; SIHU • 1 ♀; Ogasawara Isls, Chichijima Is., Mt. Sakaiura; 27 Apr. 2025; Y. Hisasue leg.; SIHU • 1 ♂; Ogasawara Isls, Chichijima Is., Mulberry Bay; 29 Apr. 2025; Y. Hisasue leg.; SIHU • 1 ♀; Ogasawara Isls, Hahajima Is., Nishiura; 4 May 2025; Y. Hisasue leg.; SIHU • 1 ♂ 4 ♀♀; Ogasawara Isls, Ototojima Is., Shikahama – Mt. Hirone; 27 May 2025; S. Shimamoto leg.; SIHU • 3 ♂♂ 5 ♀♀; Ogasawara Isls, Ototojima Is., Kohama; 2 Jun. 2025; Y. Hisasue leg.; SIHU. The four nymphs recorded above are in poor condition and are thus not described in the present study.
Diagnosis.
Omoplax desecta is recognized among the other Omoplax species based on a combination of the following characteristics: rostrum reaching posterior margin of metasternum (Fig. 11 B); pronotal disc pale brown (Figs 3 B, 4 B, 5 B, 6 B); hood more than 0.5 times as wide as maximum width of head across compound eyes, not reaching apex of clypeus (Fig. 14 B); paranotum without areolae in middle part, with areolae in remaining parts; anterior margin of hemelytron weakly curved downward in apical half (Figs 7 B, 8 B, 9 B, 10 B); subcostal and discoidal areas of hemelytron not united; costal area narrower than combined width of subcostal and discoidal areas; Sc (subcosta) vein of hemelytron distinct in apical part of dorsal view; R + M (fused radius and media) vein of hemelytron indistinct, not carinate; and ventral surface of body in various shades of brown (Figs 12 B, 13 B).
Remarks.
In general appearance, Omoplax desecta is very similar to O. majorcarinae, whose distribution range is consistent with that of O. desecta in Chichijima Group (Guilbert 2001; Souma and Kamitani 2021; Souma 2022 a); however, the former can be distinguished from the latter based on the following five main characters (Figs 3 B, G, 4 B, G, 5 B, G, 6 B, G, 7 B, G, 8 B, G, 9 B, G, 10 B, G, 11 B, G, 14 B, G): rostrum reaching posterior margin of metasternum (reaching middle part of mesosternum in O. majorcarinae); anterior margin of hemelytron weakly curved downward in apical half (strongly curved downward in apical half in O. majorcarinae); subcostal and discoidal areas of hemelytron not united (united in O. majorcarinae); Sc (subcosta) vein of hemelytron distinct in apical part of dorsal view (indistinct in apical part of dorsal view in O. majorcarinae); and R + M (fused radius and media) vein of hemelytron distinct, carinate (indistinct, not carinate in O. majorcarinae). Morphological differences between O. desecta and the five other Omoplax species are presented in the identification key below.
Distribution.
Japan: Ogasawara Islands: Chichijima Group (Anijima Island, Chichijima Island, Nishijima Island, Ototojima Island), Hahajima Group (Hahajima Island, Meijima Island, Mukohjima Island), Mukojima Group (Nakodojima Island) (Fig. 18) (Government of Japan 2010; Souma and Kamitani 2021; Souma 2022 a; Shimamoto and Ishikawa 2023). Omoplax desecta is endemic to the Ogasawara Islands.
Host plant.
Rhaphiolepis indica var. tashiroi (Rosaceae) (Fig. 17 C) and Calophyllum inophyllum (Clusiaceae) (Fig. 17 D), also known as “ Shimasharinbai ” and “ Terihaboku ”, respectively, were confirmed as host plants for Omoplax desecta by the field and captive observations of adults and nymphs, suggesting the possibility of polyphagy for this lace bug species. However, no feeding behavior of O. desecta was observed on Ardisia sp. (Primulaceae), Bischofia javanica Blume (Phyllanthaceae), Cinnamomum sp. (Lauraceae), Ligustrum sp. (Oleaceae), and Terminalia sp. (Combretaceae), from which only a few adults were collected in previous studies (cf. Yasunaga et al. 1993; Guilbert 2001; Yamada and Tomokuni 2012). Therefore, these five tree species do not appear to be host plants for this lace bug species.
Bionomics.
Omoplax desecta inhabits an evergreen broad-leaved forest with a subtropical climate in the Ogasawara Islands (Souma and Kamitani 2021), and sucks sap on the abaxial side of the leaves of Rhaphiolepis indica var. tashiroi and Calophyllum inophyllum, causing irregular yellowing on the adaxial side (Fig. 17 C, D). Adults were collected in all seasons (Drake 1956; Nishimura and Arai 1989; Yasunaga et al. 1993; Tomokuni 1994; Miyano 1998; Guilbert 2001; Souma and Kamitani 2021; Souma 2022 a); nymphs were collected in March, April, and October. In Anijima Island, O. desecta has been confirmed in the stomach contents of the invasive green anole, Anolis carolinensis (Squamata, Dactyloidae) (Takahashi et al. 2014).
Notes
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Linked records
Additional details
Identifiers
Biodiversity
- Collection code
- NSMT , SIHU
- Event date
- 1974-05-06 , 1976-06-05 , 1976-06-08 , 1976-06-12 , 1977-12-08 , 1996-07-30 , 1996-08-04 , 1997-04-15 , 1997-04-16 , 1997-04-17 , 1997-04-18 , 1997-04-19 , 1997-04-20 , 1997-04-23 , 1997-04-26 , 1997-04-27 , 1997-07-04 , 1997-07-05 , 1997-07-06 , 1997-07-07 , 1997-07-10 , 2001-03-20 , 2021-07-01 , 2021-07-02 , 2021-07-29 , 2022-05-26 , 2022-05-27 , 2023-05-21 , 2024-02-04 , 2024-03-23 , 2024-05-17 , 2024-06-14 , 2024-06-17 , 2024-06-20 , 2024-07-03 , 2024-07-11 , 2024-07-12 , 2024-07-14 , 2024-07-15 , 2024-07-16 , 2024-07-19 , 2024-07-28 , 2024-09-21 , 2024-09-22 , 2024-09-23 , 2024-09-24 , 2024-09-25 , 2024-09-27 , 2024-10-02 , 2024-10-07 , 2024-10-08 , 2024-10-25 , 2024-11-05 , 2024-11-17 , 2024-12-10 , 2025-01-10 , 2025-03-09 , 2025-03-11 , 2025-04-25 , 2025-04-27 , 2025-04-29 , 2025-05-04 , 2025-05-15 , 2025-05-27 , 2025-06-02
- Verbatim event date
- 1974-05-06 , 1976-06-05 , 1976-06-08 , 1976-06-12 , 1977-12-08 , 1996-07-30 , 1996-08-04 , 1997-04-15 , 1997-04-16 , 1997-04-17 , 1997-04-18 , 1997-04-19 , 1997-04-20 , 1997-04-23 , 1997-04-26 , 1997-04-27 , 1997-07-04 , 1997-07-05 , 1997-07-06 , 1997-07-07 , 1997-07-10 , 2001-03-20 , 2021-07-01 , 2021-07-02 , 2021-07-29 , 2022-05-26 , 2022-05-27 , 2023-05-21 , 2024-02-04 , 2024-03-23 , 2024-05-17 , 2024-06-14 , 2024-06-17 , 2024-06-20 , 2024-07-03/04 , 2024-07-11 , 2024-07-12 , 2024-07-14 , 2024-07-15 , 2024-07-16 , 2024-07-19 , 2024-07-28 , 2024-09-21 , 2024-09-22 , 2024-09-23 , 2024-09-24 , 2024-09-25 , 2024-09-27 , 2024-10-02 , 2024-10-07 , 2024-10-08 , 2024-10-25 , 2024-11-05 , 2024-11-17 , 2024-12-10 , 2025-01-10 , 2025-03-09 , 2025-03-11 , 2025-04-25 , 2025-04-27 , 2025-04-29 , 2025-05-04 , 2025-05-15 , 2025-05-27 , 2025-06-02
- Scientific name authorship
- Horvath
- Kingdom
- Animalia
- Phylum
- Arthropoda
- Order
- Hemiptera
- Family
- Tingidae
- Genus
- Omoplax
- Species
- desecta
- Taxon rank
- species
- Type status
- syntype
- Taxonomic concept label
- Omoplax desecta (Horvath, 1912) sec. Souma, 2025
References
- Horvath G (1912) Species generis Tingitidarum Stephanitis. Annales Historico-Naturales Musei Nationalis Hungarici 10: 319-339.
- Tomokuni M (1994) Type materials of several Japanese and Korean Tingidae (Heteroptera), with designation of Lectotypes and notes on Matsumura's collection. Japanese Journal of Entomology 62 (4): 839–844.
- Souma J (2022 a) Two new species of the genus Omoplax (Hemiptera: Heteroptera: Tingidae) from Mukojima Island, with new records of lace bugs endemic to the Ogasawara Islands, Japan. Acta Entomologica Musei Nationalis Pragae 62 (1): 117–127. https://doi.org/10.37520/aemnp.2022.008
- Takeya C (1962) Taxonomic revision of the Tingidae of Japan, Korea, the Ryukyus and Formosa Part 1 (Hemiptera). Mushi 36 (5): 41–75.
- Drake CJ (1948) New species of Stephanitis Stål including a list of species of the World. Notes d'Entomologie Chinoise 12 (6): 45–56.
- Takeya C (1951) A tentative list of the Tingidae of Japan and her adjacent territories (Hemiptera). Kurume University Journal. Nature and Science 4 (1): 5–28.
- Drake CJ, Maa TC (1953) Chinese and other Oriental Tingoidea (Hemiptera). Quarternary Journal of Taiwan Museum 6 (2): 87–101.
- Drake CJ (1956) Hemiptera: Tingidae. Insects of Micronesia 7 (2): 101–116.
- Drake CJ, Ruhoff FA (1960) Lace-bug genera of the world (Hemiptera: Tingidae). Proceedings of the United States National Museum 112 (3431): 1–105. [pls. 1–9] https://doi.org/10.5479/si.00963801.112-3431.1
- Drake CJ, Ruhoff FA (1965) Lacebugs of the world, a catalog (Hemiptera: Tingidae). Bulletin of the United States National Museum 243: [i – viii +] 1–634. [pls. frontis piece + 1–56] https://doi.org/10.5479/si.03629236.243.1
- Nakane T (1970) The insects of the Bonin and the Volcano Islands. In: Higher Education and Science Bureau, Ministry of Education (Ed.) The nature of the Bonin and the Volcano Islands. Higher Education and Science Bureau, Ministry of Education, and Cultural Properties Protection Division, Agency for Cultural Affairs, Tokyo, Japan, 15–32. [in Japanese with English summary]
- Miyamoto S, Yasunaga T (1989) Heteroptera. In: Hirashima Y (Ed.) A Check List of Japanese Insects. I. Laboratory of Entomology, Faculty of Agriculture, Kyushu University, Fukuoka, Japan, 151–188. [in Japanese]
- Nishimura M, Arai K (1989) On insects recorded on Hahajima and Chichijima, the Bonin Islands in 1987. Kanagawa Kenritsu Hakubutsukan Kenkyu Hokoku, Shizen Kagaku 18: 35–45.
- Kato M (1992) List of insects in the Ogasawara Islands. WWFJ Science Report 1 (1): 73–105. [in Japanese with English summary]
- Yasunaga T, Takai M, Yamashita I, Kawamura M, Kawasawa T (1993) A field guide to Japanese bugs. Terrestrial heteropterans. Zenkoku Noson Kyoiku Kyokai, Publishing Co., Ltd., Tokyo, Japan, 380 pp. [in Japanese]
- Yasunaga T (1995) The heteropteran fauna and zoogeography of Bonin Islands. The Nature & Insects 30 (1): 18–21.
- Péricart J, Golub VB (1996) Superfamily Tingoidea Laporte, 1832. In: Aukema B, Rieger C (Eds) Catalogue of the Heteroptera of the Palaearctic Region. Vol. 2, Cimicomorpha I. Netherlands Entomological Society, Amsterdam, Netherlands, 3–78.
- Miyano S (1998) Insects of Chichijima and Hahajima in Bonin Islands. Ogasawara Research 24: 1–15. [in Japanese with English summary]
- Guilbert E (2001) Western Pacific Tingidae (Heteroptera): New species and new records. Memoirs of the Queensland Museum 46 (2): 543–554.
- Karube H, Takakuwa M [Eds] (2004) Oriental Galapagos, the Ogasawara Islands: Charm of endemic organisms and the threats they face. Kanagawa Prefectural Museum of Natural History, Kanagawa, Japan, 169 pp. [in Japanese]
- Ohbayashi T, Inaba M, Suzuki H, Kato M (2004) List of insects in Ogasawara (Bonin) Islands, Japan (2002). Ogasawara Research 29: 17–74. [in Japanese]
- Government of Japan (2010) 3-2. Species List. In: Government of Japan (Eds) Letter of recommendation for inscription on the UNESCO World Natural Heritage sites: Ogasawara Islands: Supplementary information. Government of Japan, Tokyo, Japan 147–241. [in Japanese]
- Yamada K, Tomokuni M (2012) Family Tingidae Laporte, 1832. In: Ishikawa T, Takai M, Yasunaga T (Eds) A field guide to Japanese bugs. Terrestrial heteropterans III. Zenkoku Noson Kyoiku Kyokai, Publishing Co., Ltd., Tokyo, Japan, 180–213. [pl. 2–13] [in Japanese]
- Takahashi H, Yamaki A, Akita K, Kishimoto T, Toda M (2014) Feeding habits of the green anole in the Ogasawara Islands and its impact on the native insect community. Bulletin of the Herpetological Society of Japan 2014 (2): 158–167. [in Japanese]
- Yamada K, Ishikawa T (2016) Family Tingidae. In: Hayashi M, Tomokuni M, Yoshizawa K, Ishikawa T (Eds) Catalogue of the insects of Japan. Volume 4. Paraneoptera. Touka-shobo, Fukuoka, Japan, 429–435. [in Japanese]
- Souma J, Kamitani S (2021) Taxonomic review of the lace bug genus Omoplax (Hemiptera: Heteroptera: Tingidae) endemic to " Oriental Galapagos " (the Ogasawara Islands, Japan) with the description of its new allied genus and species. Entomological Science 24 (1): 3–11. https://doi.org/10.1111/ens.12445
- Shimamoto S, Ishikawa T (2023) Catalog of Heteroptera (Insecta, Hemiptera) of the Ogasawara Islands. Japanese Journal of Entomology 26 (2): 84–106. [New Series] https://doi.org/10.20848/kontyu.26.2_84
- Souma J (2023) New Japanese names or recently revised scientific names for 31 species in 15 genera of lace bugs (Heteroptera, Tingidae). Rostria 68: 1–19. [in Japanese]