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Published August 27, 2025 | Version v1
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Current challenges and preliminary morphological reassessment of the genus Chaerilus Simon, 1877 in China (Scorpiones: Chaerilidae)

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Tang, Victoria (2025): Current challenges and preliminary morphological reassessment of the genus Chaerilus Simon, 1877 in China (Scorpiones: Chaerilidae). Euscorpius 406: 1-89, DOI: 10.5281/zenodo.16963598

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urn:lsid:plazi.org:pub:ED232462E009FFE49A274819FF8FAA0A
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Publication: 10.1016/j.ympev.2023.107834 (DOI)
Publication: 10.1371/journal (DOI)
Publication: 10.1093/isd/ixae027 (DOI)
Publication: 10.1016/j.tree.2023.11.002 (DOI)
Publication: 10.1016/j.saa.2024.124309 (DOI)
Publication: 10.1111/icad.12684 (DOI)
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Taxonomic treatment: http://treatment.plazi.org/id/111A5C1AE005FFEB991C4E15FCEBADE9 (URL)
Taxonomic treatment: http://treatment.plazi.org/id/111A5C1AE019FFF59AE248F3FE80AEC9 (URL)
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Taxonomic treatment: http://treatment.plazi.org/id/111A5C1AE013FFFB998F49B5FE1AA8A9 (URL)
Taxonomic treatment: http://treatment.plazi.org/id/111A5C1AE016FFFB9ADE4ADFFBCBADF7 (URL)
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References

  • BASTAWADE, D. B. 1985. Scorpions (Arachnida). Records of the Zoological Survey of India, 82(1-4): 259-262.
  • BASTAWADE, D. B. 2006. Arachnida: Scorpionida, Uropygi, Schizomida and Oncopodid Opiliones (Chelicerata). Zoological Survey of India. Fauna of Arunachal Pradesh, State Fauna Series, 13: 449-465.
  • BENTON, T. G. 1991. The life history of Euscorpius flavicaudis (Scorpiones, Chactidae). The Journal of Arachnology, 19: 105-110.
  • BURBRINK. F. T., E. A. MYERS & R. A. PYRON. 2024. Understanding species limits through the formation of phylogeographic lineages. Ecology and Evolution, 14: 1-18.
  • BURRIEL-CARRANZA, B., M. ESTARELLAS, G. RIANO, A. TALAVERA, H. TEJERO-CICUENDEZ, J. ELS & S. CARRANZA. 2023. Species boundaries to the limit: integrating species delimitation methods is critical to avoid taxonomic inflation in the case of the Hajar banded ground gecko (Trachydactylus hajarensis). Molecular Phylogenetics and Evolution, 186(4): 107834. https://doi. org/10.1016/j.ympev.2023.107834
  • CAIN, S., E. GEFEN & L. PRENDINI. 2021. Systematic revision of the sand scorpions, genus Buthacus Birula, 1908 (Buthidae C.L. Koch, 1837) of the Levant, with redescription of Buthacus arenicola (Simon, 1885) from Algeria and Tunisia. Bulletin of the American Museum of Natural History, 450: 1-134.
  • COYNE, J. A. & H. A. ORR. 2004. Speciation. Sunderland, Massachusetts: Sinauer Associates, 545 pp.
  • DI, Z.-Y. 2009. [Faunistic Study of Scorpion Taxonomy from the Three Provinces of Tibet, Yunnan, and Hainan in China (Chelicerata: Arachnida)]. M.S. thesis, 211 pp. Hebei University. Hebei Province, China. [unpublished, in Chinese].
  • DI, Z.-Y., Y.-L. WU, Z.-J. CAO, L.-Q. FAN & W.-X. LI. 2009. The genus Chaerilus Simon, 1877 (Scorpiones: Chaerilidae) in China, with a description of the female C. tricostatus Pocock, 1899. Arthropoda Selecta, 18: 131-138.
  • DI, Z.-Y., X.-B. XU, Z.-J. CAO, Y.-L. WU & W.-X. LI. 2013. Notes on the scorpions (Arachnida, Scorpiones) from Xizang with the redescription of Scorpiops jendeki Kovarik, 2000 (Scorpiones, Euscorpiidae) from Yunnan (China). ZooKeys, 301: 51-99.
  • DI, Z.-Y., Z.-Z. YANG, S.-J. YIN, Z-J. CAO & W.-X. LI. 2014. History of study, updated checklist, distribution and key of scorpions (Arachnida: Scorpiones) from China. Zoological Research, 35(1): 3-19.
  • DI, Z.-Y., Z.-Z. YANG, Z.-J. CAO, Y.-L. WU & W.-X. LI. 2015. [The scorpion fauna of China (Chelicerata: Arachnida)]. Acta Arachnologica Sinica, 24: 109-115. [in Chinese].
  • DI, Z.-Y. & M.-S. ZHU. 2009. A new species of Chaerilus Simon, 1877 (Scorpiones, Chaerilidae) from China. Acta Arachnologica, 58: 97-102.
  • DU, S.-Y., Q. XUAN, R. J. HOWARD, Y.-H. WANG, M. S. ENGEL & C.- Y. CAI. 2024. A genome-scale phylogeny of scorpions: model comparison and modeling amongsite compositional heterogeneity. Palaeoentomology, 007(6): 792-801.
  • DUPRE, G. 2016. Dictionary of scientific scorpion names. Arachnides, 78: 1-68.
  • FET, V. 2000. Family Chaerilidae Pocock, 1893. Pp. 323-328 in Fet, V., W. D. Sissom, G. Lowe & M. E. Braunwalder (eds.). Catalog of the Scorpions of the World (1758-1998). New York: The New York Entomological Society, 689 pp.
  • FOERSTER, S. I. A. 2025. Body size prediction in scorpions: a phylogenetic comparative examination of linear measurements of individual body parts. PeerJ, 13(12): 1-24.
  • FORDE, A., A. JACOBSEN, M. M. DUGON & K. HEALY. 2022. Scorpion species with smaller body sizes and narrower chelae have the highest venom potency. Toxins, 14(3): 219.
  • FOX, G. A., A. M. COOPER & W. K. HAYES. 2015. The dilemma of choosing a reference character for measuring sexual size dimorphism, sexual body component dimorphism, and character scaling: cryptic dimorphism and allometry in the scorpion Hadrurus arizonensis. PLoS ONE, 10(3): e0120392. https://doi:10.1371/journal. pone.0120392
  • HENDERSON, J. R. 1913. Zoological Results of the Arbor Expedition 1911-1912. Arachnida, I. C. Scorpiones. Records of the Indian Museum, 8: 128-133.
  • HJELLE, J. T. 1990. Anatomy and morphology. Pp. 5-30 in: Polis, G. A. (ed.). The Biology of Scorpions. Stanford, CA: Stanford University Press.
  • HUBER, B. A., H. SZYMANSKI & A. BENNETT-WEST. 2024. Progress or burden? Formal description of every apparently new species available in collections is neither necessary nor useful. ZooKeys, 1214: 77-90.
  • JOSHI, M., M. ESPELAND, P. HUEMER, J. DEWAARD & M. MUTANEN. 2024. Species delimitation under allopatry: genomic insights within and across continents in Lepidoptera. Insect Systematics and Diversity, 8(5): 1-16. https://doi.org/10.1093/isd/ixae027
  • KARBSTEIN, K., L. KOSTERS, L. HODAC, M. HOFMANN, E. HORANDL, S. TOMASELLO, N. D. WAGNER, B. C. EMERSON, D. C. ALBACH, S. SCHEU, S. BRADLER, J. DE VRIES, I. IRISARRI, H. LI, P. SOLTIS, P. MADER & J. WALDCHEN. 2024. Species delimitation 4.0: integrative taxonomy meets artificial intelligence. Trends in Ecology & Evolution, 39(8): 771-784. https:// doi.org/10.1016/j.tree.2023.11.002
  • KLOOCK, C. T. 2009. Reducing scorpion fluorescence via prolonged exposure to ultraviolet light. Journal of Arachnology, 37: 368-370.
  • KOCH, C. L. 1845. Die Arachniden. Nurnberg: C. H. Zech'sche Buchhandlung, 11, 174 pp.
  • KOVARIK, F. 1998.Stiri [Scorpiones]. Jihlava (Czech Republic): Madagaskar Publishing House, 176 pp. [in Czech].
  • KOVARIK, F. 2000. Revision of family Chaerilidae (Scorpiones), with descriptions of three new species. Serket, 7: 38-77.
  • KOVARIK, F. 2001. Catalog of the Scorpions of the World (1758-1998) by V. Fet, W. D. Sissom, G. Lowe, and M. Braunwalder (New York Entomological Society, 2000: pp. 690). Discussion and supplement for 1999 and part of 2000. Serket, 7(3): 78-93.
  • KOVARIK, F. 2005. Chaerilus pictus (Pocock, 1890). Akva tera forum, 2005: 1.
  • KOVARIK, F. 2012. Five new species of Chaerilus Simon, 1877 from China, Indonesia, Malaysia, Philippines, Thailand, and Vietnam (Scorpiones: Chaerilidae). Euscorpius, 149: 1-14.
  • KOVARIK, F. 2018. Notes on the genera Buthacus, Compsobuthus, and Lanzatus with several synonymies and corrections of published characters (Scorpiones: Buthidae). Euscorpius, 269: 1-12.
  • KOVARIK, F. 2019. Chaerilus alberti sp. n. from Malaysia (Scorpiones: Chaerilidae). Euscorpius, 274: 1-9.
  • KOVARIK, F. & G. LOWE. 2012. Review of the genus Neobuthus Hirst, 1911 with description of a new species from Ethiopia (Scorpiones: Buthidae). Euscorpius, 138: 1-25.
  • KOVARIK, F. & G. LOWE. 2022. Scorpions of the Horn of Africa (Arachnida, Scorpiones). Part XXVIII. Scorpions of Djibouti. Euscorpius, 357: 1-31.
  • KOVARIK, F., G. LOWE, D. HOFEREK, M. FORMAN & J. KRAL. 2015. Two new Chaerilus from Vietnam (Scorpiones, Chaerilidae), with observations on growth and maturation of Chaerilus granulatus sp. n. and C. hofereki Kovarik et al., 2014. Euscorpius, 213: 1-21.
  • KOVARIK, F., G. LOWE, K. B. RANAWANA, D. HOFEREK, V. A. S. JAYARATHNE, J. PLISKOVA & F. STAHLAVSKY. 2016a. Scorpions of Sri Lanka (Arachnida, Scorpiones: Buthidae, Chaerilidae, Scorpionidae) with description of four new species of the genera Charmus Karsch, 1879 and Reddyanus Vachon, 1972, stat. n. Euscorpius, 220: 1-133.
  • KOVARIK, F., G. LOWE, M. STOCKMANN & F. STAHLAVSKY. 2020. Two new Chaerilus from Thailand and Laos (Scorpiones: Chaerilidae). Euscorpius, 324: 1-20.
  • KOVARIK, F., G. LOWE & F. STAHLAVSKY. 2016b. Scorpions of the Horn of Africa (Arachnida: Scorpiones). Part IX. Lanzatus, Orthochirus, and Somalicharmus (Buthidae), with Description of Lanzatus somalilandus sp. n. and Orthochirus afar sp. n. Euscorpius, 232: 1-38.
  • KOVARIK, F., G. LOWE & F. STAHLAVSKY. 2018. Three new Chaerilus from Malaysia (Tioman Island) and Thailand (Scorpiones, Chaerilidae), with a review of C. cimrmani, C. sejnai and C. tichyi. Euscorpius, 268: 1-27.
  • KOVARIK, F., V. FET, B. GANTENBEIN, M. R. GRAHAM, E. A. YAGMUR, F. STAHLAVSKY, N. М. POVERENNYI & N. E. NOVRUZOV. 2022a. A revision of the genus Mesobuthus Vachon, 1950, with a description of 14 new species (Scorpiones: Buthidae). Euscorpius, 348: 1-189.
  • KOVARIK, F., G. LOWE, F. STAHLAVSKY & P. JUST. 2022b. The genus Barbaracurus in Saudi Arabia (Scorpiones: Buthidae), with description of a new species. Euscorpius, 365: 1-27.
  • KOVARIK, F. & A. A. OJANGUREN-AFFILASTRO. 2013. Illustrated catalog of scorpions. Part II. Bothriuridae; Chaerilidae; Buthidae I. Genera Compsobuthus, Hottentotta, Isometrus, Lychas, and Sassanidotus. Prague: Clairon Production, 400 pp.
  • KRAEPELIN, K. 1894. Revision der Scorpione. II. Scorpionidae und Bothriuridae. Beiheft zum Jahrbuch der Hamburgischen Wissenschaftlichen Anstalten, 11: 1-248.
  • KRAEPELIN, K. 1899. Scorpiones und Pedipalpi. In Dahl, F. (ed.). Das Tierreich. Herausgegeben von der Deutschen Zoologischen Gesellschaft. Berlin: R. Friedlander und Sohn Verlag, 8 (Arachnoidea): 1-265.
  • KRAEPELIN, K. 1913. Neue Beitrage zur Systematik der Gliederspinnen. III. A. Bemerkungen zur Skorpionenfauna Indiens. B. Die Skorpione, Pedipalpen und Solifugen Deutsch-Ost-Afrikas. Mitteilungen aus dem Naturhistorischen Museum (2. Beiheft zum Jahrbuch der Hamburgischen Wissenschaftlichen Anstalten, 1912), 30: 123-196.
  • LE, S., J. JOSSE & F. HUSSON. 2008. FactoMineR: an R package for multivariate analysis. Journal of Statistical Software, 25(1): 1-18.
  • LIU, Y.-Q., S.-R. LI, Y.-T. LI, X.-R. WANG , H.-Z. ZHANG, W.-J. WU, Z.-J. CAO, K. HUANG & Y.-L. WU. 2024. Differential fluorescence features and recovery speeds of different scorpion exoskeleton parts during the molting process. Spectrochimica Acta Part A: Molecular and Biomolecular Spectroscopy, 316, 124309. https://doi. org/10.1016/j.saa.2024.124309
  • LOPEZ-CABRERA, D., G. RAMOS-ORTIZA, E. GONZALEZ-SANTILLAN & R. ESPINOSA-LUNA. 2020. Characterization of the fluorescence intensity and color tonality in the exoskeleton of scorpions. Journal of Photochemistry & Photobiology, B: Biology, 209, 111945: 1-9.
  • LORIA, S. F., V. L. EHRENTHAL, A. D. NGUYEN & L. PRENDINI. 2022. Climate relicts: Asian scorpion family Pseudochactidae survived Miocene aridification in caves of the Annamite Mountains. Insect Systematics and Diversity, 6(6): 1-21.
  • LORIA, S. F. & L. PRENDINI. 2014. Homology of the lateral eyes of Scorpiones: a six-ocellus model. PLoSOne, 9: e112913.
  • LOURENCO, W. R. 2012. Fluorescence in scorpions under UV light; can chaerilids be a possible exception? Comptes Rendus Biologies, 335: 731-734.
  • LOURENCO, W. R. 2018. The evolution and distribution of noxious species of scorpions (Arachnida: Scorpiones). Journal of Venomous Animals and Toxins including Tropical Diseases, 24(1): 1-12.
  • LOURENCO, W. R. 2019. The genus Chaerilus Simon, 1877 (Scorpiones: Chaerilidae) in Vietnam with the description of a new species. Arachnology, 17(7): 45-53.
  • LOURENCO, W. R. 2020. The coevolution between telson morphology and venom glands in scorpions (Arachnida). Journal of Venomous Animals and Toxins including Tropical Diseases, 26: 1-8.
  • LOURENCO, W. R. 2023. The remarkable micro-scorpion genus Microbuthus Kraepelin, 1898 in North Africa; description of a new species with comments on its biogeography and ecology (Scorpiones: Buthidae). Serket, 20(1): 1-10.
  • LOURENCO, W. R. & B. DUHEM. 2010. The genus Chaerilus Simon, 1877 (Scorpiones, Chaerilidae) in the Himalayas and description of a new species. Zookeys, 37: 13-25.
  • LOURENCO, W. R. & A. ROSSI. 2019. The cave population of Chaerilus Simon, 1877 from Palawan, Philippines, and description of a new species (Scorpiones: Chaerilidae). Comptes Rendus Biologies, 342: 45-53.
  • LOURENCO, W. R., D. SUN & M.-S. ZHU. 2010. A new species of Chaerilus Simon, 1877 (Scorpiones, Chaerilidae) from Thailand. The Raffles Bulletin of Zoology, 58(1): 79-85.
  • LOURENCO, W. R., T.-H. TRAN & D.-S. PHAM. 2020. The genus Chaerilus Simon, 1877, in Vietnam with the description of a new species found in a volcanic cave (Scorpiones, Chaerilidae). Bulletin de la Societe entomologique de France, 125(1): 19-28.
  • LOURENCO, W. R. & E. YTHIER. 2021. A particular new species of Orthochirus Karsch, 1891 from Somalia (Scorpiones: Buthidae). Serket, 17(4): 335-349.
  • LOURENCO, W. R. & E. YTHIER. 2024. A new cave species of Chaerilus Simon, 1877 from Myanmar (Scorpiones: Chaerilidae). Revista Iberica de Aracnologia, 44: 51-56.
  • LOWE, G. 2010. New picobuthoid scorpions (Scorpiones: Buthidae) from Oman. Euscorpius, 93: 1-53.
  • LOWE, G. 2024. Star-studded stingers: fluorescent sensilla on the scorpion aculeus (Arachnida, Scorpiones). Euscorpius, 402: 1-39.
  • LOWE, G. & V. FET. 2024. A survey of proximal sensilla associated with denticle subrows on scorpion pedipalp fingers (Arachnida: Scorpiones), with observations on scorpion fluorescence. Euscorpius, 382: 1-107.
  • LOWE, G. & F. KOVARIK. 2019. Review of Grosphus Simon, 1880, with description of Teruelius gen. n., a new buthid genus from Madagascar (Scorpiones: Buthidae). Euscorpius, 281: 1-128.
  • LOWE, G. & F. KOVARIK. 2022. Reanalysis of Teruelius and Grosphus (Scorpiones: Buthidae) with descriptions of two new species. Euscorpius, 356: 1-105.
  • LOWE, G., F. KOVARIK, M. STOCKMANN, MARK & F. STAHLAVSKY. 2019. Trypanothacus gen. n., a new genus of burrowing scorpion from the Arabian Peninsula (Scorpiones: Buthidae). Euscorpius, 2019: 1-30.
  • MASON, N. A., N. K. FLETCHER, B. A. GILL, W. C. FUNK & K. R. ZAMUDIO. 2020. Coalescent-based species delimitation is sensitive to geographic sampling and isolation by distance. Systematics and Biodiversity, 0(0): 1-12.
  • MNHN-RS-RS8623. Chaerilus tessellatus Qi, Zhu & Lourenco, 2005. Paratype female. Museum National d'Histoire Naturelle, Paris, France (accessed 5. XI. 2024). https://science.mnhn.fr/institution/mnhn/collection/rs/ item/rs8623?listIndex=10&listCount=22
  • MONJARAZ-RUEDAS, R., R. W. MENDEZ & M. HEDIN. 2023. Species delimitation, biogeography, and natural history of dwarf funnel web spiders (Mygalomorphae, Hexurellidae, Hexurella) from the United States / Mexico borderlands. ZooKeys, 1167: 109-157.
  • NAVIDPOUR, S. & G. LOWE. 2009. Revised diagnosis and redescription of Apistobuthus susanae (Scorpiones, Buthidae). The Journal of Arachnology, 37: 45-59.
  • PEREZ, S. M. 1974. Un inventario preliminar de los escorpiones de la region Paleartica y claves para la identificacion de los generos de la region Paleartica Occidental. Madrid: Universidad Complutense de Madrid, Facultad de Ciencias, Departamento de Zoologia, Catedra de Artropodos, 7: 1-45.
  • POCOCK, R. I. 1890. Description of a new genus and species of scorpion belonging to the group Jurini. Annals and Magazine of Natural History, 6 (5): 250-252.
  • POCOCK, R. I. 1894a. Scorpions from the Malay Archipelago. Pp. 84-100 in M. Weber (ed.). Zoologische Ergebnisse einer Reise in Niederlandisch Ost-lndien, 3: 84-99. Leiden: Verlag von E. J. Brill.
  • POCOCK, R. I. 1894b. A small contribution to our knowledge of the scorpions of India. Annals and Magazine of Natural History, 6(13): 72-84.
  • POCOCK, R. I. 1899. Descriptions of six new species of scorpions from India. Journal of the Bombay Natural History Society, 12: 262-268.
  • POCOCK, R. I. 1900. Arachnida. The Fauna of British India, including Ceylon and Burma. Published under the authority of the Secretary of State for India in Council. London: W. T. Blandford, xii, 279 pp.
  • PRENDINI, L., V. L. EHRENTHAL & S. F. LORIA. 2021. Systematics of the relictual Asian scorpion family Pseudochactidae Gromov, 1998, with a review of cavernicolous, troglobitic, and troglomorphic scorpions. Bulletin of the American Museum of Natural History, 453: 1-149.
  • QI, J.-X., M.-S. ZHU & W. R. LOURENCO. 2005. Eight new species of the genera Scorpiops Peters, Euscorpiops Vachon, and Chaerilus Simon (Scorpiones: Euscorpiidae, Chaerilidae) from Tibet and Yunnan, China. Euscorpius, 32: 1-40.
  • QUEIROZ de, K.. 2007. Species concepts and species delimitation. Systematic Biology, 56(6): 879-886.
  • RANASINGHE, U. G. S. L., J. THORMANN, S. P. BENJAMIN, A. BEZDEK, J. EBERLE & D. AHRENS. 2023. Contrasting results of multiple species delimitation approaches cause uncertainty in synecological studies: A case study on Sri Lankan chafers. Insect Conservation and Diversity, 16(6): 870-885. https://doi.org/10.1111/ icad.12684
  • SANCHEZ-PINERO, F., F. URBANO-TENORIO & L. PUERTA-RODRIGUEZ. 2024. Foraging strategies, prey selection and size- and microhabitat-related diet variation in Buthus.
  • SANTIAGO-BLAY, J. A., V. FET, M. E. SOLEGLAD & S. R. ANDERSON. 2004. A new genus and subfamily of scorpions from Lower Cretaceous Burmese amber (Scorpiones: Chaerilidae). Revista Iberica de Aracnologia, 9: 3-14.
  • SHARMA, P. P., C. M. BAKER, J. G. COSGROVE, J. E. JOHNSON, J. T. OBERSKI, R. J. RAVEN, M. S. HARVEY, S. L. BOYER & G. GIRIBET. 2018. A revised dated phylogeny of scorpions: phylogenomic support for ancient divergence of the temperate Gondwanan family Bothriuridae. Molecular Phylogenetics and Evolution, 122: 37-45.
  • SHARMA, P. P., R. FERNANDEZ, L. A. ESPOSITO, E. GONZALEZ-SANTILLAN & L. MONOD. 2015. Phylogenomic resolution of scorpions reveals multilevel discordance with morphological phylogenetic signal. Proceedings of the Royal Society B, Biological Sciences, 282: 20142953.
  • SIMON, E. 1877. Etudes arachnologiques. 6e Memoire. X. Arachnides nouveaux ou peu connus. Annales de la Societe Entomologique de France, Series 1-6, 7: 225-242.
  • SISSOM, W. D., G. A. POLIS & D. D. WATT. 1990. Field and laboratory methods. Pp. 215-221 in: Polis, G. A. (ed.). The Biology of Scorpions. Stanford, CA: Stanford University Press.
  • SOLEGLAD, M. E. & V. FET. 2003a. The scorpion sternum: structure and phylogeny (Scorpiones: Orthosterni). Euscorpius, 5: 1-34.
  • SOLEGLAD, M. E. & V. FET. 2003b. High-level systematics and phylogeny of the extant scorpions (Scorpiones: Orthosterni). Euscorpius, 11: 1-175.
  • SOLEGLAD, M. E. & W. D. SISSOM, 2001. Phylogeny of the family Euscorpiidae Laurie, 1896: a major revision. Pp. 25-111 in Fet, V. & P.A. Selden (eds.). Scorpions 2001. In memoriam Gary A. Polis. Burnham Beeches, Bucks: British Arachnological Society.
  • SOUSA, P., M. A. ARNEDO & D. HARRIS, DAVID. 2017. Updated catalogue and taxonomic notes on the Old- World scorpion genus Buthus Leach, 1815 (Scorpiones, Buthidae). ZooKeys, 686: 15-84.
  • STAHNKE, H. L. 1970. Scorpion nomenclature and mensuration. Entomological News, 81(12): 297-316.
  • SUN, D. 2010. [Taxonomy and Status of Resources of the Scorpiones from China (Chelicerata: Arachnida)]. M.S. thesis, 274 pp. Hebei University. Hebei Province, China. [unpublished, in Chinese].
  • SUN, D. & Z.-N. SUN. 2011. Notes on the genus Mesobuthus (Scorpiones: Buthidae) in China, with description of a new species. The Journal of Arachnology, 39: 59-75.
  • TAKASHIMA, H. 1945. [Scorpions of Eastern Asia]. Acta Arachnologica, 9(3-4): 68-106. [in Japanese].
  • TANG, V. 2022a. A standardized list of scorpion names in Chinese, with an etymological approach. Euscorpius, 350: 1-91. Supplementary file (updated checklist) available at: https://www.researchgate. net/publication/369762475_A_standardized_list_of_ scorpion_names_in_Chinese_with_an_etymological_ approach
  • TANG, V. 2022b. Scorpions of China: an updated catalogue with comments on some taxonomic issues (Arachnida: Scorpiones). Euscorpius, 355: 1-18.
  • TANG, V. 2022c. A new species of Scorpiops Peters, 1861 from Yunnan Province, China, with a preliminary review of its congeners in Yunnan (Scorpiones: Scorpiopidae). Euscorpius, 360: 1-45.
  • TANG, V. 2023. Description of the adult male Scorpiops tongtongi Tang, 2022, with further comments on the genus Scorpiops Peters, 1861 in China (Scorpiones: Scorpiopidae). Euscorpius, 377: 1-52.
  • TANG, V. 2024a. Scorpion external morphology glossary. Zenodo. 32 pp. https://zenodo.org/records/13310174
  • TANG, V. 2024b. Confirmation of basitarsal compound slit sensilla (BCSS) in scorpion genera Chaerilus and Scorpiops (Arachnida: Scorpiones). Zenodo. 29 pp. https://zenodo.org/records/13310127
  • TANG, V. 2025. A review of scorpiofauna of China: nomenclatural notes and updated faunistic catalog (Arachnida: Scorpiones). Euscorpius, 404: 1-24.
  • TANG, V., Q.-Q. JIA & L. LIU. 2023. A new monotypic genus and species from China, Langxie feti gen. et sp. n. (Scorpiones: Buthidae). Euscorpius, 370: 1-101.
  • TANG, V. & Z.-B. LIU. 2024. Albinism in Olivierus martensii (Karsch, 1879) (Scorpiones: Buthidae). Euscorpius, 396: 1-33.
  • TANG, V., Z.-B. LIU, M. R. GRAHAM, V. FET, F. KOVARIK& F. STAHLAVSKY. 2024a. Revision of the genus Olivierus in Xinjiang, China, with comments on Mesobuthus thersites (Scorpiones: Buthidae). Euscorpius, 383: 1-58.
  • TANG, V., K.-C. OUYANG, Z.-B. LIU & F. STAHLAVSKY. 2024b. Three new species of genus Scorpiops Peters, 1861 from Tibet, China (Scorpiones: Scorpiopidae), with implications for the diagnostic values of qualitative characters. Euscorpius, 394: 1-40.
  • TARKHNISHVILI, D. 2024. Molecular markers and taxonomic explosion in herpetology: more or less robust taxonomy? True and false advantages of DNA markers. International Journal of Zoology and Animal Biology, 7(2): 1-14.
  • ERUEL, R. & A. K. TIETZ. 2008. The true identity of Rhopalurus pintoi Mello-Leitao, 1932, with notes on the status and distribution of Rhopalurus crassicauda Caporiacco, 1947 (Scorpiones: Buthidae). Euscorpius, 70: 1-14.
  • TIKADER, B. K. & D. B. BASTAWADE. 1983. Scorpions (Scorpionida: Arachnida). In The Fauna of India, Vol. 3. (Edited by the Director). Calcutta: Zoological Survey of India, 671 pp.
  • WU, W., W.-L. NG, J.-X. YANG, W.-M. LI & X.-J. GE. 2018. High cryptic species diversity is revealed by genomewide polymorphisms in a wild relative of banana, Musa itinerans, and implications for its conservation in subtropical China. BMC Plant Biology, 18: 194.
  • VENCES, M., A. MIRALLES & C. DUFRESNES. 2024. Next-generation species delimitation and taxonomy: implications for biogeography. Journal of Biogeography, 00: 1-14.
  • WIKIPEDIA. Tibet Autonomous Region - Administrative divisions of Tibet Autonomous Region. (accessed 14. XI. 2024). https://en.wikipedia.org/wiki/Tibet_Autonomous_ Region
  • XUAN, Q., L. PRENDINI, M. S. ENGEL, C.-Y. CAI & D.-Y. HUANG. 2025. Extinct scorpion family Chaerilobuthidae from Mid-Cretaceous Burmese amber reinterpreted as subfamily of extant family Pseudochactidae (Chelicerata: Scorpiones). Zoological Journal of the Linnean Society, 203: 1-60.
  • YANG, X.-F. 2008. [Scorpion Fauna and Taxonomy in China (Chelicerata: Arachnida)]. M.S. thesis, 108 pp. Hebei University. Hebei Province, China. [unpublished, in Chinese].
  • YIN, S.-J., Y.-.N. QIU, Z.-H. PAN, S.-B. L. & Z.-Y. DI. 2015. Chaerilus pseudoconchiformus sp. n. and an updated key of the chaerilid scorpions from China (Scorpiones, Chaerilidae). ZooKeys, 495: 41-51.
  • ZHU, M.-S., G.-X. HAN & W. R. LOURENCO. 2008. The chaerilid scorpions of China (Scorpiones: Chaerilidae). Zootaxa 1943: 37-52.
  • ZHU, M.-S., QI J.-X. & SONG D.-X. 2004. [A Checklist of Scorpions from China (Arachnida: Scorpiones)]. Acta Arachnologica Sinica, 13(2): 111-118. [in Chinese]