Genus Teulisna Walker, 1862

Teulisna Walker, 1862, Journal of the proceedings of the Linnean Society (Zoology), 6: 109 (Type species: Teulisna plagiata Walker, 1862, by original designation).

Diagnosis. Species of the genus (Figs 11–91) are diverse externally, usually have distally angular forewing and display a moderate or limited sexual dimorphism. In most groups, males have a forewing with an elongate cluster of androconial scales below the vein Cu (similar to the Thysanoptyx generic complex). The male genitalia of the genus (Figs 98–144) are characterised by the combination of the following features. (1) The scaphium is thin, gelatinous or weakly sclerotised. (2) The vinculum is trapezoidal with thin but heavily sclerotised arms. The intravincular area has a medial saccate corema and a complex of sclerotisations: the processus intrinsicus, medius, introrsum, and the paired plostri bearing gelatinous and easily detachable bodies termed the caselli (Birket-Smith 1965; Volynkin 2024). Some of the aforementioned intravincular structures are absent in certain groups. (3) The juxta is frame-like with a membranous ventro-medial section and is extended into caulis dorsally. (4) The basis valvae is well-developed. (5) The costa is very short and occupies only the basal section of the dorsal margin of the valva, in most groups it lays only on the outer wall of the valva. (6) The editum is short, thin and gelatinous or weakly sclerotised. The tendon is present. (7) The sacculus is proximally broad, forming a fold, the dorsal margin of which is connected with the apex of the juxta by the membranous commissure. (8) The distal saccular process has a dorsal plate-like branch lying in the plane of the outer wall of the valva. (9) The lamella centralis is present, distally articulated with the dorsal branch of the distal saccular process, and in a number of groups bears a process. In the recent paper devoted to the genitalia morphology of Lithosiini (Volynkin 2024), it was suggested to use the descriptive term the crest of lamella centralis to refer to the protrusions of this sclerite. However, as in most groups of Teulisna it is not crest-shaped but thorn-shaped or clavate, the new term pseudoampulla is introduced herein to refer to the process of the lamella centralis. The phallus and vesica structures are diverse within the genus. The female genitalia of Teulisna (Figs 150–181) are characterised by the combination of the following features. (1) The ductus bursae is relatively short, dorso-ventrally flattened, gelatinous or with sclerotised plates. (2) The corpus bursae is saccate and has a gelatinous posterior section in many species bearing irregular areas of sclerotisation, wrinkles or clusters of spinulose scobination. In certain groups the corpus bursae also has diverticula and/or protrusions. (3) The appendix bursae is short conical, can be situated in various parts of the corpus bursae depending on the group; reduced in certain species.

Notes. (1) The genus is subdivided into four subgenera in the present paper. The names Teulisna, Tegulata Walker 1862 and Corcura Walker 1862 were introduced in the same publication (Walker 1862), and Birket-Smith (1965), acting as the First Reviser (see Article 24.2 of ICZN (1999)), determined the priority of Teulisna over Tegulata and Corcura. (2) The following species are herein excluded from Teulisna and transferred to the genus Mithuna: Mithuna tongdzuythanhi (Dubatolov & Bucsek, 2016), comb. n., Mithuna flava (Dubatolov & Bucsek, 2016), comb. n., Mithuna varia (Bucsek, 2020), comb. n., Mithuna mithunoides (Černý, 2009), comb. n., Mithuna tenebrosus (N. Singh & Kirti, 2016), comb. n.