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APPENDIX 1 in A new genus and tribal placement for Ceneus speculiferus Fairmaire, 1879, comments on Grundmannius Basilewsky, 1965, Mirachlaenius Facchini, 2011 and Stenoodes Basilewsky, 1953, and a key to the Oriental and Australasian genera of Chlaeniina Brullé, 1834 (Coleoptera, Carabidae)
Authors/Creators
- 1. National Museum of Natural History, Bulgarian Academy of Sciences, 1 Tsar Osvoboditel Blvd, 1000 Sofia (Bulgaria)
- 2. University of California Berkeley, Essig Museum of Entomology, 1101 VLSB # 4780, Berkeley, CA 94720 (United States)
Description
APPENDIX 1. — List of species and subspecies and number of specimens studied, with data for their provenance and repository.
| Species and subspecies | countries specimens collections | ||
|---|---|---|---|
| Callistus lunatus (Fabricius, 1775) | Bulgaria, Turkey (Koru Dağları) | 3 | NMNHS |
| Chlaenius (Achlaenius) variicornis A. Morawitz, 1863 | North Korea (Mount Myohyang) | 1 | coll. EK |
| Chlaenius (Amblygenius) anthraceus Alluaud, 1918 | Nigeria | 1 | NMNHS |
| Chlaenius (Amblygenius) episcopalis Dejean, 1831 (s.l.) | Zimbabwe (Victoria Falls) | 1 | NMNHS |
| Chlaenius (Amblygenius) glabratus Dejean, 1831 | Sudan (South Darfur) | 3 | NMNHS |
| Chlaenius (Amblygenius) lioderus Andrewes, 1923 | India (Delhi) | 1 | NMNHS |
| Chlaenius (Amblygenius) liothorax Alluaud, 1934 | Nigeria | 1 | NMNHS |
| Chlaenius (Amblygenius) macropus Chaudoir, 1876 | Nepal (Lumbini) | 1 | coll. EK |
| Chlaenius (Amblygenius) omochlorus Andrewes, 1931 | Nepal (Trisuli Bazar) | 1 | NMNHS |
| Chlaenius (Amblygenius) pretiosus Chaudoir, 1856 | India (Delhi) | 4 | NMNHS |
| Chlaenius (Amblygenius) rufomarginatus Dejean, 1831 | Zimbabwe (Victoria Falls) | 1 | NMNHS |
| Chlaenius (Amblygenius) trachys Andrewes, 1923 | Sri Lanka (Dambulla) | 1 | NMNHS |
| Chlaenius (Amblygenius) sp. | Nepal (Lumbini), Sri Lanka (Dambyla) | 2 | NMNHS |
| Chlaenius (Callistoides) amabilis Chaudoir, 1876 | Nepal (Lumbini) | 3 | NMNHS |
| Chlaenius (Callistoides) malachinus (Motschulsky, 1865) | India (Delhi) | 1 | NMNHS |
| Chlaenius (Callistoides) tripustulatus (Dejean, 1831) | Nigeria | 3 | NMNHS |
| Chlaenius (Callistoides) sp. | Sri Lanka (Dambyla) | 1 | NMNHS |
| Chlaenius (Capsochlaenius) luisae Gestro, 1895 | Somalia | 2 | coll. EK |
| Chlaenius (Chlaeniellus) atratulus Mandl, 1983 (s.l.) | India (Ladakh) | 1 | NMNHS |
| Chlaenius (Chlaeniellus) extensus Mannerheim, 1825 | Turkmenistan | 2 | NMNHS |
| Chlaenius (Chlaeniellus) flavipes Ménétriés, 1832 | Albania, Bulgaria, Turkey | 8 | NMNHS |
| Chlaenius (Chlaeniellus) impressicollis Chaudoir, 1876 | India (Elephanta Isl.) | 1 | NMNHS |
| Chlaenius (Chlaeniellus) koenigi Semenov, 1888 | Turkmenistan | 1 | coll. EK |
| Chlaenius (Chlaeniellus) nigricornis (Fabricius, 1787) | Bulgaria | 3 | NMNHS |
| Chlaenius (Chlaeniellus) semiviridis Andrewes, 1920 | Indonesia (W Timor) | 4 | NMNHS |
| Chlaenius (Chlaeniellus) stenoristus Chaudoir, 1876 | India (Delhi) | 1 | NMNHS |
| Chlaenius (Chlaeniellus) terminatus Dejean, 1826 | Bulgaria | 1 | NMNHS |
| Chlaenius (Chlaeniellus) tristis tristis (Schaller, 1783) | Bulgaria | 2 | NMNHS |
| Chlaenius (Chlaeniellus) vestitus (Paykull, 1790) | Albania, Bulgaria, Greece, Turkey, Armenia | 53 | NMNHS |
| Chlaenius (Chlaeniellus) sp. | Thailand | 1 | NMNHS |
| Chlaenius (Chlaeniodromus) tetraphacus Alluaud, 1918 | Ivory Coast | 2 | coll. EK |
| Chlaenius (Chlaeniostenus) angustatus Dejean, 1831 | Nigeria | 1 | NMNHS |
| Chlaenius (Chlaeniostenus) circumdatus Brullé, 1835 (s.l.) | Nepal (Lumbini), Sri Lanka (Polonnaruwa), Indonesia (Mentawan Isl.) | 5 | NMNHS |
| Chlaenius (Chlaeniostenus) nitidicollis Dejean, 1826 | India (Delhi) | 2 | NMNHS |
| Chlaenius (Chlaeniostenus) sulcipennis sulcatulus Boheman, 1848 | Mozambique (Cabo Delgado) | 1 | NMNHS |
| Chlaenius (Chlaenites) spoliatus spoliatus (P. Rossi, 1792) | Bulgaria | 5 | NMNHS |
| Chlaenius (Chlaenius) dusaultii (L. Dufour, 1821) (s.l.) | Sudan (South Darfur & West Darfur), Nigeria | 8 | NMNHS |
| Chlaenius (Chlaenius) festivus festivus (Panzer, 1796) | Albania, North Macedonia, Bulgaria, Greece, Armenia, Iraq | 18 | NMNHS |
| Chlaenius (Chlaenius) festivus velutinus (Duftschmid, 1812) | Spain | 3 | NMNHS |
| Chlaenius (Chlaenius) meticulosus Laferté-Sénectère, 1851 | Senegal, Nigeria | 2 | coll. EK |
| Chlaenius (Chlaenius) pseudocruciatus Kirschenhofer, 2009 | Sudan (South Darfur) | 3 | NMNHS, coll. EK |
| Chlaenius (Dinodes) cruralis Fischer von Waldheim, 1829 | North Macedonia | 1 | NMNHS |
| Chlaenius (Dinodes) decipiens (L. Dufour, 1820) | North Macedonia, Bulgaria, Greece | 17 | NMNHS |
| Chlaenius (Epomis) dejeanii (Dejean, 1831) | Turkey (Adana) | 1 | NMNHS |
| Chlaenius (Epomis) duvaucelii (Dejean, 1831) | India (Delhi) | 1 | coll. EK |
| Chlaenius (Epomis) nigricans Wiedemann, 1821 | India (Dehradun, Delhi), Nepal (Lumbini), Sri Lanka (Polonnaruwa), Indonesia (Lombok) | 8 | NMNHS, coll. EK |
| Chlaenius (Haplochlaenius) baehri (Kirschenhofer, 1998) | Indonesia (Sulawesi) | 1 | SMNS |
| Chlaenius (Haplochlaenius) costiger costiger Chaudoir, 1856 | North Korea (Kumgangsan) | 1 | coll. EK |
| Chlaenius (Iberodinodes) dives Dejean, 1826 (s.l.) | Spain | 1 | NMNHS |
| Chlaenius (Lissauchenius) borgouensis Kirschenhofer, 2007 | Nigeria | 2 | NMNHS |
| Chlaenius (Lissauchenius) darfurensis Kirschenhofer, 2007 | Sudan (West Darfur) | 1 | coll. EK |
| Chlaenius (Lissauchenius) rufifemoratus lynx Chaudoir, 1856 | North Korea (Suyangsan Falls) | 1 | coll. EK |
| Chlaenius (Nectochlaenius) canariensis seminitidus Chaudoir, 1856 | Israel | 1 | coll. EK |
| Chlaenius (Nectochlaenius) persicus L. Redtenbacher, 1850 | Iran (Shiraz) | 1 | coll. EK |
| Chlaenius (Ocybatus) dyrollei (Laferté-Sénectère, 1851) (s.l.) | Nigeria | 2 | NMNHS |
| Chlaenius (Ocybatus) discicollis (Laferté-Sénectère, 1851) | Nigeria | 2 | NMNHS |
| Chlaenius (Pachydinodes) bipustulatus Boheman, 1848 | Nigeria | 1 | NMNHS |
| Chlaenius (Pachydinodes) pictus Chaudoir, 1856 | India (Delhi) | 1 | NMNHS |
| Chlaenius (Pachydinodes) posticus (Fabricius, 1798) | Sri Lanka (Polonnaruwa) | 4 | NMNHS |
| Chlaenius (Pachydinodes) virgulifer Chaudoir, 1876 | India (Delhi) | 1 | coll. EK |
| Chlaenius (Pachydinodes) sp. | Sudan (South Darfur) | 1 | NMNHS |
| Chlaenius (Pleroticus) buquetii hacquardi Chaudoir, 1883 | Zaire (Kisangani) | 1 | NMNHS |
| Chlaenius (Prochlaeniellus) senegalensis Dejean, 1831 | Sudan (South Darfur), Nigeria | 2 | NMNHS |
| Chlaenius (Pseudochlaeniellus) chitwanensis Kirschenhofer, 2005 Nepal (Lumbini) | 1 | NMNHS | |
| Chlaenius (Rhopalopalous) janthinus L.Redtenbacher, 1844 | India (Jammu & Kashmir) | 5 | NHMUK |
| Chlaenius (Rhysotrachelus) nepalensis Hope, 1831 | Nepal (Betrawati) | 1 | NMNHS |
| Chlaenius (Sphodromimus) holzschuhi Casale, 1984 | Nepal | 1 | SMNS |
| Chlaenius (Stenochlaenius) coeruleus (Steven, 1809) | Turkey (Artvin) | 4 | NMNHS |
| Species and subspecies | countries | specimens collections | |
| Chlaenius (Tomochilus) alternans (Imhoff, 1843) (s.l.) | Nigeria | 2 | NMNHS |
| Chlaenius (Tomochilus) carbonatus Chaudoir, 1876 | Senegal, Burkina Faso, Nigeria, Ethiopia | 5 | NMNHS, coll. EK |
| Chlaenius (Trichochlaenius) aeneocephalus aeneocephalus Dejean, 1826 | Bulgaria | 10 | NMNHS |
| Chlaenius (unplaced) rufithorax Wiedemann, 1821 | India | 2 | NHMUK |
| Grundmannius dispar (Péringuey, 1896) | Mozambique, South Africa | 4 | CAS, NMNHS |
| Harpaglossus laevigatus (Dejean, 1828) | “W. Africa”, Senegal | 3 | NHMUK, SMNS |
| Harpaglossus opacus (Chaudoir, 1857) | Eritrea, Saudi Arabia, India | 7 | IZAS, NHMUK |
| Hololeius ceylanicus (Nietner, 1856) | Sri Lanka, Myanmar, Indonesia (Bali), Australia | 19 | NMNHS, NHMUK, SMNS |
| Holosoma hedini (Andrewes, 1935) | China (Gansu) | 2 | NMNHS |
| Holosoma weigoldi (Heller, 1923) | China (Sichuan) | 3 | NMNHS |
| Lapitachlaenius speculiferus (Fairmaire, 1879) n. comb. | Fiji | 17 | ANIC, BPBM, NZAC, SAMA |
| Lapita chlaenius specularis (Emden, 1937) n. comb. | Vanuatu | 3 | NHMUK |
| Mirachlaenius barbarae Facchini, 2011 | India (Nilgiri Hills) | 1 | SMNS |
| Parachlaenius (Parachlaenius) eminus Kolbe, 1894 | Uganda, Tanzania (Bukoba) | 3 | NHMUK, SMNS |
| Parachlaenius (Parachlaenius) violaceus Péringuey, 1899 | “E. Africa”, Zimbabwe | 5 | NHMUK, SMNS |
| Procletodema parallelum Péringuey, 1899 | Zimbabwe | 3 | NHMUK, SMNS |
| Procletus minor Basilewsky, 1968 | Nigeria (Samaru) | 1 | NHMUK |
| Procletus singularis Péringuey, 1896 | Zimbabwe | 3 | NHMUK, SMNS |
| Rhopalomelus angusticollis Boheman, 1848 | South Africa | 6 | NHMUK, SMNS |
| Stenoodes jeanneli Basilewsky, 1953 | Madagascar | 1 | MNHN |
| Stuhlmannium mirabile Kolbe, 1894 | “E. Africa” | 1 | NHMUK |
Notes
Published as part of Guéorguiev, Borislav & Will, Kipling, 2025, A new genus and tribal placement for Ceneus speculiferus Fairmaire, 1879, comments on Grundmannius Basilewsky, 1965, Mirachlaenius Facchini, 2011 and Stenoodes Basilewsky, 1953, and a key to the Oriental and Australasian genera of Chlaeniina Brullé, 1834 (Coleoptera, Carabidae), pp. 258-259 in Zoosystema 47 (15) on page 258, DOI: 10.5252/zoosystema2025v47a15, http://zenodo.org/record/15502564
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Related works
- Is part of
- Journal article: 10.5252/zoosystema2025v47a15 (DOI)
- Journal article: urn:lsid:plazi.org:pub:FFD25810C85FFFDDFFD6FFDFFF868266 (LSID)
- Journal article: http://publication.plazi.org/id/FFD25810C85FFFDDFFD6FFDFFF868266 (URL)
- Journal article: http://zenodo.org/record/15502564 (URL)
- Journal article: http://zoobank.org/11740D9A-34E8-48B0-A6C9-15AB438BA100 (URL)