Results

We examined 21 individuals of A. nasreddini from two sites and 30 individuals of A. escherichii from one site, analysing 29 morphometric (Table 1) and eight meristic characters (Table 2). An additional 351 A. escherichii were visually examined for marked differences in body shape.All examined characters were found to be largely overlapping (Table 1 and 2). Moreover, we found no difference in the color patterns and the scale coverage of the ventral keel.

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Distribution. Alburnus escherichii is native to the Sakarya River drainage where it is widespread. It is also widespread in the Eber, Akşehir, Karamık and Çavuşcu (Ilgın) lake basins including the Selevir and Seyitler reservoirs. There are few records from the upper parts of the Kızılırmak and Yeşilırmak River drainages (Black Sea basin) and this species was introduced and has well-established populations in the Manavgat River (Mediterranean Sea basin) and in the Lake Beyşehir basin (Freyhof & Özuluğ 2009; Özuluğ et al. 2018) (Fig. 1 and 5). It cannot be fully excluded that populations in the Kızılırmak and Yeşilırmak Rivers might be introduced, as A. escherichii seems to be an invasive species in Turkey, most likely spread with fish stocked from fish farms to the wild.

Material examined and additional records without voucher materials.

Sakarya River drainage: FFR 00827, 80, 52– 101 mm SL; Ankara prov.: stream Öz at Kızılcahamam, 40.469E 32.659N. — FFR 00887, 82, 36– 102 mm SL; Ankara prov.: stream İlhanlı at Ayaş, 40.097N 32.251E. — FFR 04610, 24, 47– 118 mm SL; Ankara prov.: stream Berçin at Kızılcahamam, 40.471N 32.656E. — FFR 04616, 18, 50– 85 mm SL; K̹ tahya prov.: stream Porsuk at 10 km southwestern of K̹tahya, 39.350N 30.038E. — FFR 04617, 12, 65– 81 mm SL; Ankara prov.: stream Kirmir at 3 km north of G̹d̹l, 40.236N 32.261E. — FSJF 1921, 5, 77–85 mm SL; Bilecik prov.: a tributary of Sakarya River, 15 km northern of Boz ̹y̹k, 40.039N 30.122E. — FSJF 1964, 23, 69–102 mm SL; Sakarya prov.: Sakarya River at Doğançay, 40.627N 30.333E. — FSJF 2489, 7, 88–100 mm SL; Sakarya prov.: stream Mudurnu at Akyazı, 40.714N 30.563E. — FSJF 2627, 7, 79–95 mm SL; Ankara prov.: stream Bayındır at Çeltikçi, 40.324N 32.467E. — FSJF 3062, 11, 57–95 mm SL; Sakarya prov.: stream Doğançay at Doğançay, 40.627N 30.333E.

Additional records. Kütahya prov.: stream Pınarcık at Pınarcık village, 39.0164N 29.975E.— Eskişehir prov.: stream Allıkova at 15 km southern of İnönü, 39.682N 30.154E.— Sakarya prov.: Sakarya River at 8 km northern of Gevye, 40.596N 30.323E.— Kütahya prov.: a tributary of Sakarya River 2 km southeastern of Soğukçeşme, 39.577N 30.083E.— Eskişehir prov.: a tributary of Sakarya River at İnönü, 39.817N 30.132E.— Sakarya prov.: Sakarya River at Gevye, 40.538N 30.299E.— Eskişehir prov.: a tributary of stream Seydi at Hamidiye, 39.576N 30.907E.— Eskişehir prov.: stream Seydi at Saithalimpaşa, 39.415N 30.123E.— Afyonkarahisar prov.: a tributary of Sakarya River 15 km west of Emirdağ, 39.050N 31.323E.— Afyonkarahisar prov.: stream Pınarbaşı 27 km south of Emirdağ, 39.267N 31.148E.— Eskişehir prov.: Sakarya River at Çandır, 39.277N 31.332E.— Eskişehir prov.: a tributary of Sakarya River at Göktepe, 39.151N 31.614E.— Eskişehir prov.: Sakarya River at Ahiler, 39.195N 31.622E.— Ankara prov.: stream Koca, upper Sarıyer Dam at Davutoğlan, 40.133N 31.639E.— Konya prov.: drying canal a tributary of Sakarya River at Küçükhasan, 38.958N 31.831E.— Ankara prov.: stream Kirmir at Harmancık, 40.090N 31.889E.— Bursa prov.: stream İncirli at Köprühisar, 40.282N 30.122E.— Eskişehir prov.: a tributary of Çukurhisar Pond at Çukurhisar, 39.282N 30.122E.— Eskişehir prov.: Outlet of Porsuk Dam, 39.607N 30.265E.— Bursa prov.: stream Göksu at Köprühisar, 40.282N 29.797E.— Eskişehir prov.: a tributary of Sakarya River 2 km north of Ahiler, 39.215N 31.622E.— Ankara prov.: Aşartepe Dam at Çiğdemci, 40.144N 32.389E.

Kızılırmak River drainage: FFR 00980, 3, 55– 105 mm SL; Sivas prov.: Kızılırmak River at Zara, 39.961N 37.739E.

Yeşilırmak River drainage: FFR 00986, 26, 81– 111 mm SL; Erzincan prov.: stream Köroğlu at 3 km north of Refahiye, 39.919N 38.759E.— FFR 04692, 3, 93– 108 mm SL; Erzincan prov.: stream Köroğlu at 1 km north of Refahiye, 39.911N 38.767E.

Lake Beyşehir basin: FFR 00801, 28, 57– 131 mm SL; Konya prov.: stream Kuğuluçay at Seydişehir, 37.379E 31.878N.— FFR 04628, 28, 57– 131 mm SL; Konya prov.: irrigation canal at 5 km southwestern of Seydişehir, 37.379N 31.878E.— FFR uncat., 2, 60– 70 mm SL; Konya prov.: stream Sarıöz at Beyşehir, 37.749N 31.780E.— FSJF 2618, 27, 81–124 mm SL; Konya prov.: Outlet of Lake Beyşehir below dam in the Beyşehir at Beyşehir, 37.680N 31.723E.

Additional records. Konya prov.: stream B̹y̹kköpr̹ at Üst̹nler, 37.621N 31.590E.— Konya prov.: stream Büyükçay at Adaköy, 37.512N 31.590E.— Konya prov.: Karstik spring of Beyşehir 10 km north of Sarayönü, 38.353N 32.367E.— Stream Eylikler at Beyşehir, 710N 31.736E.

Lake Akşehir basin: FSJF 1911, 11, 68–98 mm SL; Konya prov.: stream Ortaköy at Ortaköy, 38.447N 31.518E.

Additional records. Afyonkarahisar prov.: Inlet water of Lake Akşehir at Üçkuyu, 38.670N 31.530E.— Konya prov.: Lake Akşehir at Akşehir, 38.456N 31.416E.— Konya prov.: a tributary of Lake Akşehir at Karabulut, 38.440N 31.457E.— Konya prov.: Lake Akşehir 10 km west of Pazarkaya, 38.530N 31.465E.

Lake Karamik basin: Additional records. Afyonkarahisar prov.: Lake Karamik Bataklığı at Çayıryazı, 38.384N 30.751E.— Afyonkarahisar prov.: Lake Karamik Bataklığı at Armutlu, 38.418N 30.820E

Lake Eber basin: FFR 04629, 10, 63– 106 mm SL; Afyonkarahisar prov.: creek Kali at Inli, 38.517N 30.773E.— FFR08648, 10, 72– 90 mm SL; Afyonkarahisar prov.: creek Kali, a tributary to Lake Eber, 38.448N 30.643E.

Additional records. Afyonkarahisar prov.: creek Kali 5 km east of İnli, 38.541N 30.845E.— Afyonkarahisar prov.: creek Kali at Maltepe, 38.602N 30.894E.— Afyonkarahisar prov.: a tributary of Selevir Reservoir at Şuhut, 38.475N 30.682E.— Afyonkarahisar prov.: creek Kali 5 km west of Pazarağaç, 38.576N 30.872E.— Afyonkarahisar prov.: Lake Eber at Derekarabağ, 38.689N 31.162E.— Afyonkarahisar prov.: Lake Eber at Eber, 38.626N 31.151E.

Lake Çavuşcu (Ilgın) basin: FFR 04641, 11, 59– 104 mm SL; Konya prov.: stream Deliköyboğazı at 10 km south of Lake Ilgın, 38.237N 31.883E.— FSJF 2577, 13, 61–93 mm SL; Konya prov.: stream Battal 5 km south of Ilgın at road to Beyşehir, 38.236N 31.880E.— FSJF 3092, 1, 67 mm SL; Konya prov.: Ilgın Lake, 38.321N 31.890E.

Additional records. Konya prov.: stream Deliköyboğazı at 6 km north of Ilgın, 38.236N 31.880E.— Konya prov.: Lake Ilgın at Gedikören, 38.321N 31.890E.— Konya prov.: a tributary of stream Deliköyboğazı at Ilgın, 38.240N 31.880E.— Konya prov.: a tributary of Lake Ilgın at Ilgın, 38.270N 31.856E.— Konya prov.: stream Deliköyboğazı at 6 km north of Ilgın, 38.236N 31.880E.— Konya prov.: stream Deliköyboğazı at 5 km north of Ilgın, 38.237N 31.883E.— Konya prov.: stream Deliköyboğazı at 6 km north of Ilgın, 38.236N 31.880E.

Akarçay Basin: Additional records. Afyonkarahisar prov.: a tributary of Selevir Reservoir at Şuhut, 38.511N 30.705E.— Afyonkarahisar prov.: Selevir Reservoir at Şuhut, 38.475N 30.682E.— Afyonkarahisar prov.: Seyitler Reservoir at Gebeceler, 38.794N 30.789E.

Karman closed basin: Additional records. Karaman prov.: stream Buğdaylı at Ayrancı, 37.312N 33.753E.— Not preserved, Karaman prov.: stream Uludere at Ayrancı. 37.289N 33.791E.

Mediterranean Sea Basin: FFR 04680, 59, 49– 119 mm SL; Antalya prov.: stream Ilıca at Ilıca, 36.818N 31.353E.— FSJF 3118, 4, 78–82 mm SL; Antalya prov.: stream Manavgat below Oymapınar Dam, 36.899N 31.527E.

Additional records. Antalya prov.: Manavgat Dam at Manavgat, 36.878N 31.522E.— Antalya prov.: stream Karpuz at Hacıobası, 36.729N 31.594E.