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TABLE 1 in Redescription of the highly specialized aquatic † Tigrivia and the classification of the Mesozoic † Coptoclavidae (Coleoptera, Adephaga, Dytiscoidea)
Authors/Creators
- 1. School of Earth Sciences and Engineering, Guangdong Provincial Key Lab of Geological Processes and Mineral Resources, Sun Yat-sen University, Guangzhou 510275, China. & School of Ecology / State Key Laboratory of Biocontrol, Shenzhen Campus of Sun Yat-sen University, Shenzhen 518107, China.
- 2. Institut für Zoologie und Evolutionsforschung, Friedrich-Schiller-Universität Jena, Jena 07743, Germany.
- 3. State Key Laboratory of Cellular Stress Biology, School of Life Sciences, Xiamen University, Xiamen 361105, China. & Yingliang Stone Natural History Museum, Nan'an 362300, China.
- 4. School of Ecology / State Key Laboratory of Biocontrol, Shenzhen Campus of Sun Yat-sen University, Shenzhen 518107, China. & Yunnan Key Laboratory for Palaeobiology, Yunnan University, Kunming 650091, China.
Description
TABLE 1. Taxa assigned to Coptoclavidae *, based on https://www.mindat.org/taxon–3263914.html#autoanchor2, modified; see also Soriano et al. 2007), taxa likely nonmonophyletic or misplaced are marked by an asterisk.
| Subfamily | Genus | Comment | Occurrence | Age | citations |
|---|---|---|---|---|---|
| † Timarchopsinae * | † Timarchopsis * | likely misplaced in Dytiscoidea | Algeria, Germany, Russia, Asia, e.g. China, Kazakhstan | 191 Ma, Lower Jurassic-133 Ma, Early/ Lower Cretaceous | e.g. Ponomarenko 1977 (as Necronectes), Wang et al. 2010 |
| † Daohugounectes * | likely misplaced | China | 166 Ma, Middle Jurassic | Wang et al. 2009 | |
| † Ditomoptera | Germany | 151 Ma, Upper Jurassic | Wang et al. 2010 | ||
| † Exedia * | likely misplaced | Kazakhstan | 130 Ma, Early/Lower Cretaceous | Ponomarenko 1977 | |
| † Ovonectes | Spain | 130 Ma, Early/Lower Cretaceous | Soriano et al. 2007 | ||
| † Protonectes (known as larva) | Germany | 232 Ma, Late/Upper Triassic | Prokin et al. 2013 | ||
| † Stygeonectes * | likely misplaced | Russia | 183 Ma, Early Jurassic-133 Ma, Early/ Lower Cretaceous | e.g. Ponomarenko 1977 | |
| † Tuhanectes | China, Xingjiang | 168 Ma, Middle Jurassic | Hong et al. 1995 | ||
| † Peudohydrophilus* | very likely misplaced in Dytiscoidea | Germany | 152 Ma, Upper Jurassic | Deichmüller 1866 | |
| †Charonoscaphinae | † Charonoscapha | Southern Kazakhstan | 166 Ma, Middle Jurassic | e.g. Ponomarenko 1977 | |
| † Charonoscaphidia | Southern Kazakhstan | 166 Ma, Middle Jurassic | Ponomarenko 1977 | ||
| †Coptoclavinae* | † Coptoclava | Russia, Asia (China, Mongolia, Korea) | 164 Ma, Middle Jurassic-113 Ma, Early/ Lower Cretaceous | e.g. Ponomarenko 1977, Zhao et al. 2018 | |
| † Tigrivia | China | 122 Ma, Early Cretaceous | Bao et al. 2023 | ||
| † Bolbonectes | Russia | 164 Ma, Middle Jurassic-129 Ma, Lower Cretaceous | Ponomarenko 1993 | ||
| † Hoyoclava | Spain | 130 Ma, Lower Cretaceous | Ponomarenko et al. 2000 | ||
| † Metacoptoclava | Spain | 130 Ma, Lower Cretaceous | Ponomarenko et al. 2000 | ||
| † Coptoclaviscinae * | † Coptoclavella | Mongolia | 147 Ma, Middle Jurassic-125 Ma, Early/ Lower Cretaceous | Soriano et al. 2007 | |
| † Coptoclavisca * | likely misplaced | Mongolia | 166 Ma, Middle Jurassic-125 Ma, Early/ Lower Cretaceous | Ponomarenko 1987 | |
| † Holcoptera | Ukraine, England, USA, Germany | 247 Ma, Early/Lower Triassic-232 Ma, Late Upper Triassic | e.g. Thomson et al. 2017, Ponomarenko et al. 2022 | ||
| † Hispanoclavinae Not assigned to subfamily | † Hispanoclavina | Spain | 130 Ma, Early/Lower Cretaceous | Soriano et al. 2007 | |
| † Agrascapha | China | 237 Ma, Middle Triassic | Zhang et al. 2022 | ||
| † Amblycephalonius * | family placement doubtful | Germany | 183 Ma, Early Jurassic | Bode 1953 | |
| † Euroscapha | China | 237 Ma, Middle Triassic | Zhang et al. 2022 |
Notes
Published as part of Bao, Liang, Beutel, Rolf Georg, Niu, Kecheng & Bao, Tong, 2025, Redescription of the highly specialized aquatic † Tigrivia and the classification of the Mesozoic † Coptoclavidae (Coleoptera, Adephaga, Dytiscoidea), pp. 411 in Zootaxa 5588 (3) on page 411, DOI: 10.11646/zootaxa.5588.3.1, http://zenodo.org/record/14896574
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- Journal article: 10.11646/zootaxa.5588.3.1 (DOI)
- Journal article: urn:lsid:plazi.org:pub:FFE5FFEEFF8E66679532FFDA0639FF90 (LSID)
- Journal article: http://publication.plazi.org/id/FFE5FFEEFF8E66679532FFDA0639FF90 (URL)
- Journal article: http://zenodo.org/record/14896574 (URL)
- Journal article: http://zoobank.org/E3697AAB-FB6D-440B-89C7-3AD757157481 (URL)
References
- Soriano, C., Ponomarenko, A. G. & Delclos, X. (2007) Coptoclavid beetles (Coleoptera: Adephaga) from the Lower Cretaceous of Spain: a new feeding strategy in beetles. Palaeontology, 50, 525-536. https://doi.org/10.1111/j.1475-4983.2007.00642.x
- Ponomarenko, A. G. (1977) Suborder Adephaga, etc. In: Arnoldi, L. V., Zherikin, V. V., Nikritin, L. M. & Ponomarenko, A. G. (Eds.), Mesozoic Coleoptera. Trudy Paleonthologicheskogo Instituta Akademiya Nauk SSSR, 161, pp. 1-204. [in Russian]
- Wang, B., Ponomarenko, A. G. & Zhang, H. (2010) Middle Jurassic Coptoclavidae (Insecta: Coleoptera: Dytiscoidea) from China: a good example of mosaic evolution. Acta Geologica Sinica-English Edition, 84 (4), 680-687. https://doi.org/10.1111/j.1755-6724.2010.00272.x
- Wang, B., Ponomarenko, A. G. & Zhang, H. C. (2009) A new coptoclavid larva (Coleoptera: Adephaga: Dytiscoidea) from the Middle Jurassic of China, and its phylogenetic implication. Paleontological Journal, 43, 652-659. https://doi.org/10.1134/S0031030109060082
- Prokin, A. A., Makarov, K. V., Ponomarenko, A. G. & Bashkuev, A. S. (2013) New beetle larvae (Coleoptera: Coptoclavidae, Caraboidea, Polyphaga) from the Upper Triassic of Germany. Russian Entomologcal Journal, 22, 259-274.
- Hong, Y. C., Liang, S. & Hu, T. (1995) Study on geology and paleontological assemblage from Tuha Basin of Xinjiang, China. Geoscience, 9 (4), 435.
- Bao, L., Li, L., Niu, K., Wang, N., Kroeck, D. M. & Bao, T. (2023) Retracted: A new aquatic beetle (Adephaga: Coptoclavidae) from the Middle Jurassic Daohugou Biota. The Anatomical Record, 306 (8), E 1-E6. https://doi.org/10.1002/ar.25221
- Ponomarenko, A. G. (1987) New Mesozoic aquatic beetles. Paleontologicheskii Zhurnal, 2, 83-97.
- Thomson, U., Ross, A. J. & Davidson, P. (2017) Mesozoic Holcoptera (Coleoptera: Coptoclavidae) from England and the United States. Proceedings of the Geologists' Association, 128, 659-674. https://doi.org/10.1016/j.pgeola.2017.05.009
- Ponomarenko, A. G., Fedorenko, D. N. & Bashkuev, A. S. (2022) A new species of the beetle genus Holcoptera (Coleoptera: Coptoclavidae) from the Upper Triassic of east Ukraine. Russian Entomological Journal, 31 (2), 128-131. https://doi.org/10.15298/rusentj.31.2.06
- Zhang, Q., Zheng, D., Wang, B. & Zhang, H. (2022) A review of Triassic insects in China. Geological Society, London, Special Publications 521. Geological Society, London, pp. 16 pp. [pp. 45-60] https://doi.org/10.1144/SP521-2021-121
- Bode, A. (1953) Die Insektenfauna des ostniedersachsischen oberen Lias. Palaeontographica, 103 (1), 1-375.