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Table 1 in A revision of the genus Sclerocoelus Marshall (Diptera: Sphaeroceridae)
Authors/Creators
- 1. University of Guelph Insect Collection and Insect Systematics Laboratory, School of Environmental Sciences, University of Guelph, Guelph, Ontario, N 1 G 2 W 1, Canada.
Description
Table 1 (continued on next page). Character state matrix of Sclerocoelus Marshall, 1995, including Chespiritos Marshall, 2000, Paramosina Marshall, 2014, and Parasclerocoelus Marshall & Dong, 2008 as the outgroups. The plesiomorphic state is indicated by (0) and apomorphic state(s) by (1–5); multistate characters are treated as unordered.
| Taxon | 1 | 2 | 3 | 4 | 5 | 6 | 7 | 8 | 9 | 10 | 11 | 12 | 13 | 14 | 15 | 16 | 17 | 18 |
|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|
| Chespiritos Marshall, 2000 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 2 | 0 | 0 | 0 |
| Paramosina Marshall, 2014 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 0 | 0 | 0 | 0 | 0 | 2 | 0 | 1 | 0 |
| Parasclerocoelus Marshall & Dong, 2008 | 0 | 0 | 1 | 0 | 0 | 0 | 0 | 1 | 1 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 0 |
| Sclerocoelus aduncus sp. nov. | 0 | 1 | 0 | 0 | 0 | 0 | 0 | 1 | 0 | 0 | 0 | 1 | 0 | 0 | ? | ? | ? | ? |
| S. alpinus sp. nov. | 0 | 1 | 0 | 0 | 1 | 3 | 0 | 1 | 0 | 0 | 1 | 2 | 0 | 0 | 2 | 0 | 1 | 0 |
| S. altus sp. nov. | 0 | 1 | 1 | 0 | 1 | 2 | 0 | 0 | 1 | 2 | 0 | 1 | 0 | 0 | ? | ? | ? | ? |
| S. andensis Marshall, 1997 | 1 | 1 | 0 | 0 | 1 | 4 | 1 | 1 | 1 | 4 | 0 | 1 | 0 | 0 | 0 | 0 | 0 | 0 |
| S. argentinensis sp. nov. | 0 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 1 | 0 | 1 | 0 | 0 | ? | ? | ? | ? |
| S. azulensis sp. nov. | 0 | 1 | 0 | 0 | 2 | 4 | 2 | 1 | 1 | 0 | 0 | 1 | 0 | 0 | 2 | 0 | 1 | 0 |
| S. binus sp. nov. | 1 | 1 | 0 | 0 | 2 | 4 | 2 | 2 | 1 | 4 | 0 | 0 | 0 | 0 | 2 | 1 | 2 | 2 |
| S. brasilensis Marshall, 1997 | 1 | 1 | 0 | 0 | 2 | 4 | 2 | 1 | 1 | 4 | 0 | 2 | 0 | 0 | 0 | 0 | 0 | 0 |
| S. bucki sp. nov. | 0 | 1 | 0 | 0 | 2 | 3 | 0 | 0 | 0 | 2 | 0 | 1 | 0 | 0 | 2 | 0 | 1 | 1 |
| S. caligarius sp. nov. | 1 | 1 | 1 | 0 | 2 | 4 | 1 | 2 | 1 | 4 | 0 | 1 | 0 | 0 | 1 | 0 | 2 | 2 |
| S. caribensis Marshall, 1997 | 1 | 1 | 0 | 0 | 2 | 4 | 2 | 1 | 1 | 4 | 0 | 1 | 0 | 0 | 0 | 0 | 0 | 0 |
| S. chilensis sp. nov. | 0 | 1 | 0 | 0 | 1 | 3 | 0 | 1 | 0 | 2 | 0 | 1 | 0 | 0 | ? | ? | ? | ? |
| S. copiosus sp. nov. | 0 | 1 | 0 | 1 | 2 | 4 | 2 | 1 | 0 | 4 | 2 | 0 | 0 | 0 | 2 | 1 | 0 | 0 |
| S. costaricensis sp. nov. | 0 | 1 | 1 | 2 | 2 | 1 | 1 | 2 | 1 | 0 | 0 | 3 | 0 | 1 | 1 | 0 | 1 | 2 |
| S. cubus sp. nov. | 0 | 1 | 0 | 0 | 1 | 3 | 0 | 1 | 1 | 4 | 0 | 2 | 0 | 0 | ? | ? | ? | ? |
| S. dasysternum sp. nov. | 0 | 1 | 0 | 1 | 2 | 1 | 1 | 2 | 0 | 0 | 1 | 3 | 0 | 0 | 1 | 0 | 1 | 2 |
| S. dominicensis sp. nov. | 0 | 1 | 0 | 3 | 1 | 1 | 1 | 1 | 1 | 3 | 0 | 2 | 1 | 2 | 1 | 0 | 0 | 2 |
| S. dryadalis sp. nov. | 0 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 3 | 0 | 2 | 0 | 0 | 2 | 0 | 1 | 1 |
| S. elephas sp. nov. | 1 | 1 | 0 | 2 | 2 | 4 | 2 | 1 | 0 | 4 | 2 | 0 | 0 | 0 | 2 | 1 | 2 | 2 |
| S. espeletia sp. nov. | 0 | 1 | 0 | 0 | 2 | 3 | 1 | 1 | 1 | 2 | 1 | 1 | 0 | 0 | ? | ? | ? | ? |
| S. flavus sp. nov. | 0 | 1 | 0 | 0 | 1 | 0 | 0 | 0 | 0 | 3 | 0 | 2 | 0 | 0 | 2 | 0 | 1 | 1 |
| S. frigidifrons sp. nov. | 0 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 2 | 0 | 0 | 2 | 0 | 1 | 0 |
| S. galapagensis Marshall, 1997 | 1 | 1 | 0 | 0 | 2 | 4 | 1 | 1 | 1 | 4 | 1 | 3 | 0 | 0 | 0 | 0 | 0 | 0 |
| S. grandicercus sp. nov. | 0 | 1 | 1 | 1 | 2 | 3 | 1 | 2 | 1 | 0 | 1 | 3 | 0 | 0 | 1 | 0 | 1 | 0 |
| S. hemorrhoidalis Marshall, 1997 | 1 | 1 | 0 | 0 | 2 | 4 | 2 | 1 | 1 | 4 | 0 | 1 | 0 | 0 | 0 | 0 | 0 | 0 |
| S. inornatus sp. nov. | 0 | 0 | 0 | 0 | 1 | 3 | 0 | 0 | 0 | 0 | 1 | 0 | 0 | 0 | ? | ? | ? | ? |
| S. irregularis sp. nov. | 0 | 1 | 0 | 3 | 1 | 1 | 1 | 1 | 1 | 3 | 0 | 2 | 1 | 2 | 1 | 0 | 0 | 2 |
| S. latibarbus sp. nov. | 0 | 1 | 1 | 2 | 2 | 4 | 2 | 1 | 1 | 2 | 0 | 3 | 0 | 1 | 1 | 0 | 1 | 2 |
| S. lazulita sp. nov. | 0 | 1 | 0 | 0 | 2 | 2 | 0 | 0 | 0 | 2 | 0 | 2 | 0 | 0 | 2 | 0 | 1 | 0 |
| S. limbus sp. nov. | 0 | 1 | 2 | 0 | 0 | 0 | 0 | 0 | 1 | 0 | 0 | 1 | 0 | 0 | 2 | 0 | 1 | 0 |
| S. longibarbus sp. nov. | 0 | 1 | 1 | 1 | 2 | 4 | 2 | 1 | 1 | 0 | 0 | 2 | 1 | 0 | 2 | 0 | 0 | 0 |
| S. lutosus sp. nov. | 0 | 0 | 0 | 0 | 1 | 3 | 0 | 0 | 1 | 1 | 0 | 0 | 0 | 0 | 1 | 0 | 1 | 1 |
| S. mandibulum sp. nov. | 0 | 1 | 0 | 0 | 1 | 2 | 0 | 1 | 0 | 5 | 1 | 2 | 0 | 0 | 0 | 1 | 0 | 2 |
| S. masneri sp. nov. | 0 | 1 | 1 | 1 | 2 | 3 | 2 | 1 | 1 | 4 | 0 | 4 | 0 | 0 | 1 | 0 | 2 | 2 |
| S. meridensis sp. nov. | 0 | 1 | 0 | 0 | 1 | 0 | 0 | 0 | 0 | 3 | 0 | 0 | 0 | 0 | 2 | 0 | 1 | 1 |
| S. nebulosus sp. nov. | 0 | 1 | 0 | 0 | 2 | 2 | 0 | 1 | 0 | 0 | 0 | 2 | 0 | 0 | 2 | 0 | 1 | 1 |
| S. nitidistylus sp. nov. | 0 | 1 | 0 | 1 | 2 | 3 | 1 | 2 | 1 | 2 | 1 | 3 | 0 | 1 | 0 | 0 | 1 | 2 |
| S. ocellatus sp. nov. | 0 | 1 | 0 | 1 | 2 | 3 | 1 | 2 | 1 | 3 | 1 | 4 | 0 | 0 | 2 | 0 | 1 | 2 |
| S. paranebulosus sp. nov. | 0 | 1 | 0 | 0 | 2 | 3 | 0 | 1 | 0 | 0 | 0 | 1 | 0 | 0 | ? | ? | ? | ? |
| S. pararegularis sp. nov. | 0 | 1 | 0 | 3 | 1 | 1 | 1 | 1 | 1 | 3 | 0 | 2 | 1 | 2 | ? | ? | ? | ? |
| S. parasordipes sp. nov. | 0 | 1 | 0 | 1 | 2 | 3 | 1 | 2 | 0 | 5 | 0 | 1 | 0 | 0 | 2 | 0 | 2 | 1 |
| S. penai sp. nov. | 0 | 1 | 2 | 0 | 2 | 2 | 0 | 1 | 0 | 2 | 0 | 1 | 0 | 0 | ? | ? | ? | ? |
| S. plumiseta (Duda, 1925) | 0 | 1 | 0 | 0 | 1 | 1 | 0 | 1 | 1 | 1 | 0 | 3 | 1 | 0 | 2 | 0 | 0 | 2 |
| S. punensis sp. nov. | 0 | 1 | 0 | 0 | 1 | 3 | 0 | 0 | 1 | 2 | 1 | 2 | 0 | 0 | ? | ? | ? | ? |
| S. puyensis sp. nov. | 0 | 1 | 1 | 0 | 1 | 3 | 0 | 0 | 0 | 0 | 2 | 2 | 0 | 0 | 2 | 0 | 1 | 2 |
| S. rectangularis (Malloch, 1914) | 0 | 1 | 0 | 0 | 2 | 4 | 2 | 2 | 0 | 1 | 0 | 3 | 1 | 0 | 2 | 0 | 2 | 2 |
| S. recurvatus sp. nov. | 0 | 1 | 0 | 1 | 2 | 2 | 2 | 2 | 1 | 0 | 1 | 3 | 0 | 0 | 0 | 0 | 3 | 0 |
| S. regularis (Malloch, 1914) | 0 | 1 | 0 | 3 | 1 | 1 | 1 | 1 | 1 | 3 | 0 | 2 | 1 | 2 | 1 | 0 | 2 | 2 |
| S. riparius sp. nov. | 0 | 1 | 0 | 0 | 1 | 3 | 1 | 1 | 1 | 1 | 0 | 0 | 0 | 0 | 1 | 0 | 1 | 2 |
| S. rostrum sp. nov. | 0 | 1 | 1 | 1 | 2 | 3 | 2 | 1 | 1 | 2 | 0 | 3 | 0 | 1 | ? | ? | ? | ? |
| S. sordipes (Adams, 1904) | 0 | 1 | 0 | 1 | 2 | 3 | 1 | 2 | 0 | 5 | 0 | 1 | 0 | 0 | 2 | 0 | 0 | 0 |
| S. subbrevipennis (Frey, 1954) | 1 | 1 | 0 | 0 | 2 | 4 | 1 | 1 | 1 | 4 | 0 | 2 | 0 | 0 | 0 | 0 | 0 | 0 |
| S. synorios sp. nov. | 0 | 1 | 1 | 1 | 2 | 4 | 2 | 1 | 1 | 1 | 1 | 3 | 0 | 1 | 2 | 0 | 0 | 2 |
| S. tantus sp. nov. | 1 | 1 | 0 | 2 | 2 | 4 | 2 | 2 | 0 | 2 | 2 | 0 | 0 | 0 | 2 | 1 | 0 | 0 |
| S. tridens sp. nov. | 0 | 1 | 0 | 0 | 2 | 3 | 0 | 1 | 1 | 1 | 0 | 1 | 0 | 0 | 2 | 0 | 1 | 1 |
| S. turpis sp. nov. | 0 | 1 | 0 | 2 | 1 | 1 | 1 | 1 | 1 | 3 | 1 | 2 | 1 | 0 | ? | ? | ? | ? |
| S. vulgatus sp. nov. | 0 | 1 | 0 | 1 | 2 | 3 | 2 | 2 | 2 | 2 | 1 | 2 | 0 | 1 | 1 | 0 | 0 | 2 |
| S. xynos sp. nov. | 0 | 1 | 2 | 0 | 2 | 2 | 0 | 1 | 0 | 2 | 0 | 1 | 0 | 0 | 1 | 0 | 1 | 0 |
Notes
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Identifiers
References
- Marshall S. A. 1995. Sclerocoelus and Druciatus, new genera of New World Sphaeroceridae (Diptera; Sphaeroceridae; Limosininae). Insecta Mundi 9 (3 - 4): 283-289.
- Marshall S. A. 2000. Chespiritos, a new genus of Limosininae (Diptera: Sphaeroceridae) from Costa Rica. Proceedings of the Entomological Society of Washington 102 (3): 609-612.
- Marshall S. A. & Dong H. 2008. Parasclerocoelus, a new south temperate genus of Limosininae (Diptera: Sphaeroceridae). Studia Dipterologica 15: 223-230.
- Marshall S. A. 1997. A revision of the Sclerocoelus galapagensis group (Diptera: Sphaeroceridae: Limosininae). Insecta Mundi 11 (2): 97-115.
- Duda O. 1925. Die aussereuropaischen Arten der Gattung Leptocera Olivier - Limosina Macquart (Dipteren) mit Berucksichtigung der europaischen Arten. Archiv fur Naturgeschichte Abteilung A, Berlin 90 (11): 5-215.
- Malloch J. R. 1914. Costa Rican Diptera collected by Philip P. Calvert, Ph. D., 1909 - 1910. Transactions of the American Entomological Society, Philadelphia 40: 1-36.
- Adams C. F. 1904. Notes and descriptions of North American Diptera. The Kansas University Science Bulletin 2 (14): 433-455.
- Frey R. 1954. Diptera Brachycera and Sciaridae von Tristan da Cunha (with a contribution by J. Bequaert). Results of the Norwegian Scientific Expedition to Tristan da Cunha 1937 - 1938, Oslo 4 (26): 1-55.