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Bringing light into deep-sea biodiversity: a systematic revision and molecular phylogeny of the genus Scaphander Montfort, 1810 (Gastropoda: Cephalaspidea), with a focus on the Indo-Pacific
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- 1. Department of Natural History, University Museum of Bergen, University of Bergen, PB7800 5020 Bergen, Norway
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Siegwald, Justine, Malaquias, Manuel António E. (2025): Bringing light into deep-sea biodiversity: a systematic revision and molecular phylogeny of the genus Scaphander Montfort, 1810 (Gastropoda: Cephalaspidea), with a focus on the Indo-Pacific. Zoological Journal of the Linnean Society 203: 1-60, DOI: 10.1093/zoolinnean/zlad201
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- https://www.checklistbank.org/dataset/308540
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- urn:lsid:zoobank.org:pub:A9AFDD7-B8BE-47EB-9676-77488EE78A24
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- urn:lsid:plazi.org:pub:FFB9FFB3FFBDFFCEFFF014351C46C22F
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- http://publication.plazi.org/id/FFB9FFB3FFBDFFCEFFF014351C46C22F
References
- Adams A. Monograph of the family Bullidae. In: Sowerby GB (ed.), Thesaurus Conchyliorum, or Monographs of Genera of Shells. London; Sowerby, 1855;2, 553-608.
- Adams A. On some new species of Cylichnidae, Bullidae, and Philinidae from the seas of China and Japan. Annales and Magazine of Natural History 1862;9:1-12.
- Akaike H. A new look at the statistical model identification. IEEE Transactions on Automatic Control 1974;19:716-23. https://doi. org/10.1109/tac.1974.1100705
- Al-Khayat JA, Vethamony P, Nanajkar M. Molluscan diversity influenced by mangrove habitat in the khors of Qatar. Wetlands 2021;41:45.
- Amorim A, Oliveira E, Malaquias MAE et al. New insights into the functional morphology of the male copulatory apparatus of bullid gastropods. Zoomorphology 2013;132:145-55. https://doi.org/10.1007/ s00435-012-0179-7
- Annandale N, Stewart FN. Illustrations of the Zoology of the Royal Indian Marine Survey Ship Investigator, under the command of Captain WG Beauchamp,RIM. Part 6. India:Superintendent Government Printing. Mollusca, 1909.
- ArnaudF.Anewspeciesof Ascorhynchus (Pycnogonida)foundparasiticon an opisthobranchiate mollusc. Zoological Journal of the Linnean Society 1978;63:99-104. https://doi.org/10.1111/j.1096-3642.1978. tb02092.x
- Beesley PL, Ross GJB, Wells A. Mollusca: the southern synthesis. Fauna of Australia. Camberra, Australia: CSIRO Publishing, 1998, 5 Part B, 565-1234.
- Bellardi L. Descrizione di un nuovo genera della famiglia delle Bullide fossile del terreno pliocenico inferior del Piemonte e della Liguria. Bullettino della Societa Malacologica Italiana 1876;2:207210.
- Bouchet P. Opisthobranches de profondeur de l'Ocean Atlantique. I - Cephaslaspidea. Cahiers de Biologie Marine 1975;16:317-65.
- Bouckaert R, Vaughan TG, Barido-Sottani J et al. BEAST 2.5: an advanced software platform for Bayesian evolutionary analysis. PLoS Computational Biology 2019;15:e1006650. https://doi.org/10.1371/ journal.pcbi.1006650
- Brouwer D, Brouwer R, van Leeuwen S et al. De schelpen van de sultan: Oman.Correspondentieblad NMV 2000;316:110-7.
- Bruguiere JG. Encyclopedie Methodique. Histoire naturelle des vers. Paris: Panckoucke, 1792.
- Bullis HR. The genus Scaphander in the Gulf of Mexico and notes on the Western Atlantic species. Bulletin of Marine Science of the Gulf and Carribean 1956;6:1-17.
- Chaban EM, Ekimova IA, Schepetov DM et al. Meloscaphander grandis (Heterobranchia: Cephalaspidea), a deep-water species from the North Pacific: redescription and taxonomic remarks. Zootaxa 2019a;4646:zootaxa.4646.2.12. https://doi.org/10.11646/ zootaxa.4646.2.12
- Chaban EM, Ekimova IA, Schepetov DM et al. Euopisthobranch mollusks of the order Cephalaspidea (Gastropoda: Heterobranchia) of the Kuril-Kamchatka Trench and the adjacent Pacific abyssal plain with descriptions of three new species of the genus Spiraphiline (Philinidae). Progress in Oceanography 2019b;178:102185.
- Clarke AH. Annotated checklist and bibliography of the abyssal marine molluscs of the world. Bulletin (National Museum Canada) 1962;181:1-114.
- Colman JG. Studies of the life history biology of deep-sea gastropod molluscs. Doctoral dissertation, University College of Swansea, 1987.
- Cossmann M. The Gasteropods of the older tertiary of Australia. Les Opisthobranches. Transactions of the Royal Society of South Australia 1897;21:1-21.
- Dall WH. Reports on the results of dredging by the United States Coast Survey Steamer 'Blake'. XV Preliminary Report on the Mollusca. Bulletin of the Museum of Comparative Zoology 1881;9:33-144.
- Dall WH. Preliminary report on the shell-bearing mollusks and brachiopods of the southeastern coast of the United States,with illustrations of many of the species. Bulletin of the National Museum 1889a;37:1-232.
- Dall WH. Reports on the results of dredging, under the supervision of Alexander Agassiz, in the Gulf of Mexico (1877-78) and in the Caribbean Sea (1879-80), by the U. S. Coast Survey steamer 'Blake'. Bulletin of the Museum of Comparative Zoology 1889b;18:433-9.
- Dall WH. Contributions to the Tertiary fauna of Florida, with special reference to the Miocene Silex-beds of Tampa and the Pliocene beds of the Caloosahatchie River. Part I. Pulmonate, Opisthobranchiate, and orthodont gastropods. Transactions of the Wagner Free Institute of Science of Philadelphia 1890a;3:1-200.
- Dall WH. Scientific results of explorations by the U.S. Fish Commission steamer Albatross. No. VII. Preliminary report on the collection of Mollusca and Brachiopoda obtained in 1887-88. Proceedings of the United States National Museum 1890b;12:219-362. https://doi. org/10.5479/si.00963801.12-773.219
- Dall WH. Scientific results of explorations by the US Fish Commission steamer Albatross. No. XXXIV. Report on Mollusca and Brachiopoda dredged in deepwater, chiefly near the Hawaiian Islands, with illustrations of hitherto unfigured species from northwest America. Proceedings of the United States National Museum 1895;17:675-733. https://doi.org/10.5479/ si.00963801.17-1032.675
- Dall WH. Reports on the scientific results, U.S. steamer 'Albatross'. reports on the Mollusca and Brachiopoda. Bulletin of the Museum of Comparative Zoology Harvard 1908;43:205-487.
- Dall WH. Descriptions of new species of Mollusca from the North Pacific Ocean in the collection of the United States National Museum. Proceedings of the United States National Museum 1919;56:293-371. https://doi.org/10.5479/si.00963801.56-2295.293
- Dall WH.Small shells from dredgings off the southeast coast of the United States by the United States Fisheries Steamer 'Albatross' in 1885 and 1886. Proceedings of the United States National Museum 1927;70:1- 134. https://doi.org/10.5479/si.00963801.70-2667.1
- Darriba D, Taboada GL, Doallo R et al. jModelTest 2: more models, new heuristics and parallel computing. Nature Methods 2012;9:772. https://doi.org/10.1038/nmeth.2109
- Dell RK. The Archibenthal Mollusca of New Zealand. In: Dominion Museum Bulletin, Wellington, NZ: Dominion Museum, 1956, 18.
- Edgar RC. MUSCLE: multiple sequence alignment with high accuracy and high throughput. Nucleic Acids Research 2004;32:1792-7. https://doi.org/10.1093/nar/gkh340
- Eilertsen MH, Malaquias MAE. Systematic revision of the genus Scaphander (Gastropoda, Cephalaspidea) in the Atlantic Ocean, with a molecular phylogenetic hypothesis. Zoological Journal of the Linnean Society 2013a;167:389-429.
- Eilertsen MH, Malaquias MAE. Unique digestive system, trophic specialization, and diversification in the deep-sea gastropod genus Scaphander. Biological Journal of the Linnean Society 2013b;109:512- 25. https://doi.org/10.1111/bij.12069
- Finet Y. The Marine Mollusks of the Galapagos Islands. In: James MJ (ed.), Galapagos Marine Invertebrates: Taxonomy, Biogeography, and Evolution in Darwin's Islands. New York and London: Plenum Press, 1991, 253-280.
- Finet Y, Chiriboga A, Ruiz D et al. CDF Checklist of Galapagos marine mollusks. In: Charles Darwin Foundation Galapagos species checklist. Galapagos: Charles Darwin Foundation in Puerto Ayora, 2011, 1-164.
- Gioeni G. Descrizione di una Nuova Famiglia e di un Nuova Genere di Testacei trovati nel Littorale di Catania, Con qualche Osservazione Sopra una Spezie di Ostriche, per Servire alle Conchiologia Generale. Napoli: The Author, 1783.
- Gosliner TM. The opisthobranch gastropod fauna of the Galapagos Islands.In: James MJ (ed.), Galapagos Marine Invertebrates: Taxonomy, Biogeography,and Evolution in Darwin's Islands. New York and London: Plenum Press, 1991, 281-305.
- Guangyu L. Order Cephalaspidea. Fauna Sinica (Phylum Mollusca, Class Gastropoda, Subclass Opisthobranchia). Beijing: Science Press, 1997.
- Habe T. Descriptions of new genera and species of the shell-bearing opisthobranchiate molluscs from Japan (Cephalaspidea, Tectibranchia). Venus 1952;17:69-77.
- Habe T. Report on the Mollusca chiefly collected by the SS Soyo-Maru of the imperial fisheries experimental station on the continental shelf bordering Japan during the years 1922-1930 Part 1. Cephalaspidea. Publications of the Seto Marine Biological Laboratory 1954;3:301-18. https://doi.org/10.5134/174489
- Habe T. A list of the cephalaspid Opisthobranchia of Japan. Bulletin of the Biogeographical Society of Japan 1955;16-19:54-79.
- Habe T. Shells of the Western Pacific in Color. Hoikusha, 2, 1964.
- Habe T. Two new species of Hawaiian shells. Venus 1981;40:65-7.
- Hasegawa K, Okutani T. A review of bathyal shell-bearing gastropods in Sagami Bay. Memoirs of the National Science Museum. Tokyo: 2011;47:97-144.
- Herbert DG, Jones GJ, Atkinson LJ. Phylum Mollusca. In: Atkinson LJ, Sink KJ, eds. Field Guide to the Offshore Marine Invertebrates of South Africa. Pretoria: Malachite Marketing and Media, 2018, 249-320.
- Holznagel W. Research note: a nondestructive method for cleaning gastropod radulae from frozen, alcohol-fixed, or dried material. American Malacological Bulletin 1998;14:181-3.
- Hori S. Family Cylichnidae. In: Okutani T (ed.), Marine Mollusks in Japan. Tokyo: Tokai University Press, 2000, 717-47.
- Hori S. Family Scaphandridae. In: Okutani T (ed.), Marine Mollusks in Japan. Tokyo: Tokai University Press, 2017, 1087-88.
- Huelsenbeck JP, Ronquist F. MRBAYES:Bayesian inference of phylogenetic trees. Bioinformatics 2001;17:754-5. https://doi.org/10.1093/ bioinformatics/17.8.754
- ICZN. Opinion 287. Validation, under the plenary powers, of the generic name 'Scaphander ' Montfort, 1810 (class Gastropoda, order Tectibranchiata). Opinions and Declarations rendered by the International Commission on Zoological Nomenclature 1954;8:4962.
- ICZN. Principle of Priority. In: International Code of Zoological Nomenclature, 4th edn. London: The International Trust for Zoological Nomenclature, 1999a, 24-30.
- ICZN. Replacement of Junior Homonyms. In: International Code of Zoological Nomenclature, 4th edn. London: The International Trust for Zoological Nomenclature, 1999b, 62-63.
- Iredale T.Mollusca from the continental shelf of eastern Australia.Records of the Australian Museum 1925;14:243-70. https://doi.org/10.385 3/j.0067-1975.14.1925.845
- Johnson CW. List of the marine Mollusca of the Atlantic coast from Labrador to Texas. Proceedings of the Boston Society of Natural History 1934;40:1-204.
- Jones G. 2014. STACEY package documentation: species delimitation and species tree estimation with BEAST2. http://www.indriid.com/ software.html
- Jones G. Algorithmic improvements to species delimitation and phylogeny estimation under the multispecies coalescent. Journal of Mathematical Biology 2017;74:447-67. https://doi.org/10.1007/ s00285-016-1034-0
- Jones G, Aydin Z, Oxelman B. DISSECT: an assignment-free Bayesian discovery method for species delimitation under the multispecies coalescent. Bioinformatics 2015;31:991-8. https://doi.org/10.1093/ bioinformatics/btu770
- Kobelt W. Die Familie Bullidae. In: Kuster HC, Martini FHW, Chemnitz JH (eds.), Systematisches Conchylien-Cabinet von Martini und Chemnitz. Nurnberg: Bauer & Raspe, 1896, Vol. 1, 33-190, pls. 9-19.
- Kuroda T, Habe T. On some Japanese Mollusca described by A. Adams, whose specimens are deposited in the Redpath Museum of Canada. Venus 1954;18:1-16.
- Kuroda T, Habe T, Oyama K. The Sea Shells of Sagami Bay Collected by his Majesty the Emperor of Japan. Tokyo: Maruzen, 1971.
- Lemche H. Northern and Arctic Tectibranch Gastropods. II. A revision of the Cephalaspid species. Det Kongelige Danske Videnskabernes Selskab Biologiske Skrifter 1948;5:1-136.
- Linnaeus C. Systema Naturae per Regna tria Naturae, Secundum Classes, Ordines, Genera, Species, cum Characteribus, Differentiis, Synonymis, Locis, 10th edn. Holmiae: Laurentii Salviae, 1758.
- Locard A. Mollusces Testaces. In: Milne-Edwards A (ed.), Expedition Scientifique du Travailleur et du Talisman pendant les Annees 1880, 1881, 1882, 1883. Paris: Masson et Cie, 1897.
- Lorz AN, Berkenbusch K, Nodder S et al. A review of deep-sea benthic biodiversity associated with trench, canyon and abyssal habitats below 1500 m depth in New Zealand waters. New Zealand Aquatic Environment and Biodiversity Report 2012;92:1-133.
- Marcus EBR. On some Cephalaspidea (Gastropoda: Opisthobranchia) from the western and middle Atlantic warm waters. Bulletin of Marine Science 1974;24:300-71.
- Marcus EBR, Marcus EG. The R/V Pillsbury deep-sea biological expedition to the Gulf of Guinea, 1964-65, Opisthobranchs from tropical west Africa.Studies in Tropical Oceanography 1966;4:152-208.
- Marcus EBR, Marcus EG. Biological investigations of the Deep Sea. 33. Opisthobranchs from the Southwestern Caribbean Sea. Bulletin of Marine Science 1967;17:597-628.
- Martens E.Einige neue Arten von Meer-Conchylien aus den Sammlungen der deutschen Tiefsee-Expedition unter der Leitung von Prof. Carl Chun, 1898-99. Sitzungs-Berichte der Gesellschaft naturforschender Freunde zu Berlin 1902;1902:237-44.
- Martens E. A. Systematisch-geographischer Teil, mit Tafel I -V und I Addildung im Text. In: Martens E, Thiele J. 1904. Die beschalten Gastropoden der Deutschen Tiefsee-Expedition 1898-1899. Germany: G. Fischer, 1903, 7, 1-179.
- Martens E, Thiele J. Die beschalten Gasteropoden der deutschen Tiefsee-Expedition, 1898-1899.In: Chun C (ed.), Wissenschaftliche Ergebnisse der deutschen Tiefsee-Expedition auf dem Dampfer 'Valdivia', 1898- 1899. Jena: Gustav Fischer, 1903, 3-179.
- Martini FHW. 1769. Neues systematisches Conchylien-Cabinet. Berlin, Germany: G. N. Raspe.
- Maury CJ. Recent species from the Gulf of Mexico and Pleistocene and Pliocene species from the Gulf states. Bulletins of American Paleontology 1922;9:34-172.
- Melvill JC. Descriptions of thirty-one Gastropoda and one scaphopod from the Persian Gulf and Gulf of Oman, dredged by Mr. F.W. Townsend, 1902-1904. Proceedings of the Malacological Society of London 1906;7:69-80.
- Melvill JC, Standen R. The Mollusca of the Persian Gulf, Gulf of Oman and Arabian Seas as evidenced mainly through the collections of Mr. F. W. Townsend, 1893-1900, with descriptions of new species. Part 1, Cephalopoda, Gastropoda, Scaphopoda. Proceedings of the Zoological Society of London 1901;71:327-460. https://doi. org/10.1111/j.1469-7998.1901.tb08181.x
- Menke CT. Zur Familie Bullacea und deren Gattungen und Arten. Malakozoologische Blatter 1954;1:33-48.
- Mighels JW, Adams CB. Descriptions of twenty-four species of the shells of New England. Boston Journal of Natural History 1842;4:37-54.
- Minichev YS. Studies on the morphology of the lower Opisthobranchia (on the evolutionary significance of the detorsion-process). Proceedings of the Zoological Institute 1967;44:109-81.
- Minichev YS. Opisthobranch mollusks (Gastropoda, Opistobranchia). In: Zenkevich LA, Kort VG (eds.), Biology of the Pacific Ocean. 2. Deep-sea benthos. Moscow: Nauka, 1969, 42-44.
- MolluscaBase eds. 2022a. MolluscaBase. Scaphander Montfort, 1810. http://www.molluscabase.org/aphia.php?p=taxdetails&id=137871 (13 December 2022, date last accessed).
- MolluscaBase eds. 2022b. MolluscaBase. Scaphander bushirensis (Melvill & Standen, 1901). https://www.molluscabase.org/aphia. php?p=taxdetails&id=1417603 (13 December 2022, date last accessed).
- Montfort PD. Conchyliologie Systematique, et Classification Methodique des Coquilles. Paris: Schoell, Frederic, 1810.
- Morariu VI, Srinivasan BV, Raykar VC et al. Automatic online tuning for fast Gaussian summation. In: Koller D, Schuurmans D, Bengio Y et al. (eds.), Advances in Neural Information Processing Systems 2009;21:1113-20.
- Morley MS, Hayward BW. Marine Mollusca of Great Barrier Island, New Zealand. Records of the Auckland Museum 2009;46:15-51.
- Muller OF.Zoologia danica seu animalium Daniae et Norvegiae rariorum ac minus notorum historia descriptiones et historia. Weygand 1784;2:1-124.
- Nordsieck F. Die Europaischen Meeresschnecken (Opisthobranchia mit Pyramidellidae; Rissoacea). Vom Eismeer bis Kapverden, Mittelmeer und Schwarzes Meer. Stuttgart: Gustav Fischer, 1972.
- Okutani T. Archibenthal and abyssal Mollusca collected by the RV Soyo-Maru from Japanese waters during 1964. Bulletin of the Tokai Regional Fisheries Research Laboratory 1966;46:1-32.
- Okutani T. Description of a new gigantic scaphanderid gastropod, Bucconia centa, from off Koshiki Islet, west of Kyushu. Venus 1987;45:219-21.
- Pallary P. Exploration scientifique du Maroc. Deuxieme fascicule. Malacologie, Archives Scientifiques du Protectorat Francais, 1912, 1-107.
- Pequegnat WE.The Ecological Communities of the Continental Slope and Ajacent Regimes of the Northern Gulf of Mexico. College Station, Texas: United States Department of the Interior, Minerals Management Services, 1983.
- Philipsson LM. Dissertatio Historico Naturalis Sistens Testaceorum Genera. Lundae, Berlingianis, 1788.
- PilsbryHA.ManualofConchology,StructuralandSystematic,withIllustrations of the Species, Vol. 15, Polyplacophora, Acanthochitidae, Cryptoplacidae and Appendix. Tectibranchiata. Philadelphia: Conchological Section of the Academy of Natural Sciences of Philadelphia, 1893.
- Pilsbry HA. Manual of Conchology, Structural and Systematic, with Illustrations of the Species, Vol. 16, Philinidae, Gastropteridae, Aglajidae, Aplysiidae, Oxynoeidae, Runcinidae, Umbraculidae, Pleurobranchidae. Philadelphia: Conchological Section of the Academy of Natural Sciences of Philadelphia, 1895.
- Poppe GT. Philippine Marine Mollusks, Vol. 3, Gastropoda. Hackenheim: ConchBooks, 2010.
- Powell AWB. New Zealand Mollusca: Marine, Land and Freshwater Shells. Auckland: Collins, 1979.
- Puillandre N, Brouillet S, Achaz G. ASAP: assemble species by automatic partitioning. Molecular Ecology Resources 2021;21:609-20. https:// doi.org/10.1111/1755-0998.13281
- Qi Z. Seashells of China. Beijing: China Ocean Press, 2004.
- R Core Team. R: a Language and Environment for Statistical Computing. Vienna: R Foundation for Statistical Computing, 2021. https:// www.R-project.org/
- Rambaut A, Drummond AJ, Xie D et al. Posterior summarization in Bayesian phylogenetics using Tracer 1.7. Systematic Biology 2018;67:901-4.
- Rowden AA, Leduc D, Clark MR et al. Habitat differences in deep-sea megafaunal communities off New Zealand: implications for vulnerability to anthropogenic disturbance and management. Frontiers in Marine Science 2016;3:241.
- Rudman WB. A new species and genus of the Aglajidae and the evolution of the philinacean opisthobranch molluscs. Zoological Journal of the Linnean Society 1978;62:89-107. https://doi. org/10.1111/j.1096-3642.1978.tb00524.x
- Sars GO. Bidrag til Kundskaben om Norges Arktiske Fauna: I. Mollusca Regionis Arcticae Norvegiae. Norway: Christiania, 1878.
- Schepman MM. The Prosobranchia, Pulmonata and Opisthobranchia Tectibranchiata, Tribe Bullomorpha of the Siboga Expedition, Vol. 32, No. 1-6, Leiden: Brill Publishers and Printers, 1913, 6, 59-494.
- Schumacher CF. Essai d'un Nouveau Systeme des Habitations des vers Testaces. Copenhagen: Schultz, 1817.
- Siegwald J, Malaquias MAE. Revisiting the deep-sea Atlantic gastropod species Scaphandergracilis Watson,1883(Gastropoda:Cephalaspidea: Scaphandridae): first data on its anatomy, systematics, and ecology. Journal of Natural History 2022;55:3053-66. https://doi.org/10.108 0/00222933.2022.2036378
- Siegwald J, Pastorino G, Oskars T et al. A new species of the deep-sea genus Scaphander (Gastropoda, Cephalaspidea) from the Mar del Plata submarine canyon off Argentina. Bulletin of Marine Science 2020;96:111-26. https://doi.org/10.5343/bms.2019.0069
- Siegwald J, Oskars TR, Kano Y et al. A global phylogeny of the deep-sea gastropod family Scaphandridae (Heterobranchia: Cephalaspidea): redefinition and generic classification. Molecular Phylogenetics and Evolution 2022;169:107415.
- Smith EA.Natural history notes from H.M. Indian Marine Survey Steamer 'Investigator', Commander C.F. Oldham, R.N.-Series II, No. 10. Report upon some Mollusca dredged in the Bay of Bengal and the Arabian Sea. Annals and Magazine of Natural History, ser.6 1894;14:157-74.
- Smith EA. Natural history notes from H.M. Indian Marine Survey Steamer 'Investigator', Commander T.H. Heming, R.N. On Mollusca from the Bay of Bengal and the Arabian Sea. Annals and Magazine of Natural History, ser. 7 1904;14:1-14.
- Smith EA. On Mollusca from the Bay of Bengal and the Arabian sea. Annals and Magazine of Natural History 1906;18:245-64. https://doi. org/10.1080/00222930608562609
- Steyn DG, Lussi M. Offshore Shells of Southern Africa: a Pictorial Guide to more than 750 Gastropods. South Africa: The authors, 2005.
- Talavera G, Castresana J. Improvement of phylogenies after removing divergent and ambiguously aligned blocks from protein sequence alignments. Systematic Biology 2007;56:564-77. https://doi. org/10.1080/10635150701472164
- Thiele J. Gastropoda der Deutschen Tiefsee-Expedition.II Teil. In: Chun C (ed.), Wissenschaftliche ergebnisse der Deutschen Tiefsee- expedition auf dem Dampfer 'Valdivia' 1898-1899. Jena:Gustav Fischer, 1925, 17, 35-382.
- Valdes A. Deep-sea 'cephalaspidean' heterobranchs (Gastropoda) from the tropical southwest Pacific. In: Heros V, Cowie RH, Bouchet P (eds.), Tropical Deep-Sea Benthos, Vol. 25. Paris: Memoires du Museum national d'Histoire naturelle, 2008,. Vol. 196, 1-806.
- Valdes A. Northeast Pacific benthic shelled sea slugs. Zoosymposia 2019;13:242-304. https://doi.org/10.11646/zoosymposia.13.1.21
- Valdes A, McLean JH. On two abyssal species of Scaphandridae G.O. Sars, 1878 (Gastropoda: Cephalaspidea) from the eastern Pacific.The Nautilus 2015;129:118-25.
- Verrill AE. Second catalogue of Mollusca recently added to the fauna of the New England coast and the adjacent parts of the Atlantic, consisting mostly of deep sea species, with notes on others previously recorded. Transactions of the Connecticut Academy of Arts and Sciences 1884;6:139-600. https://doi.org/10.5962/bhl.part.7412
- Vogler RE. The radula of the extinct freshwater snail Aylacostoma stigmaticum (Caenogastropoda: Thiaridae) from Argentina and Paraguay. Malacologia 2013;56:329-32. https://doi.org/10.4002/ 040.056.0221
- Watson RB. Mollusca of the 'Challenger' Expedition. Part XX.The Journal of the Linnean Society 1883;17:341-6.
- Watson RB. Report on the Scaphopoda and Gasteropoda collected by H.M.S. Challenger during the years 1873-76. Report of the scientific results of the Voyage of H.M.S. Challenger during the years 1873-76. Report of the scientific results of the Voyage of H.M.S. Challenger during the years 1873-76. Zoology 1886;15:1-756.
- Weinkauff HC. Catalogue des coquilles marines recueillies sur les cotes de l'Algerie.Journal de Conchyliologie, Paris 1862;10:301-71.
- Winckworth R. The British marine Mollusca. Journal of Conchology 1932;19:211-52.
- Zaharias P, Kantor YI, Fedosov AE et al. Just the once will not hurt: DNA suggests species lumping over two oceans in deep-sea snails (Cryptogemma). Zoological Journal of the Linnean Society 2020;190:532-57. https://doi.org/10.1093/zoolinnean/zlaa010