Clivina wallacei subsp. philippinensis Kult 1951
Authors/Creators
- 1. Natural History Museum Bern, Department of Invertebrates, Bernastrasse 15, 3005 Bern, Switzerland. & Natural History Museum Bern, Department of Invertebrates, Bernastrasse 15, 3005 Bern, Switzerland. & Ligusterweg 9, CH- 8906 Bonstetten, Switzerland. mike. balkenohl @ bluewin. ch; https: // orcid. org / 0000 - 0001 - 9485 - 5568
Description
Clivina wallacei philippinensis Kult, 1951
(Figs 31, 66, 98, 124)
Clivina wallacei philippinensis Kult, 1951: 27; Balkenohl, 2001: 17; 2021c: 536; Lorenz, 2005: 145; 2024: 8 LTLL.
Type material. Holotype: ♀, with labels and data: white, black printed “Philippine Is. Coll. Bottcher, B.M. 1929- 201.” / “ Philippine Islands Manila 8: VII: 1914 Coll. Bottcher ” / white, handwritten in black ink and printed “Wallacei ssp. philippinensis Kult det. K. Kult 47” / red, black framed, black printed “TYPE” (NHMUK). Paratypes: 1 ♂, with labels and data: white, black printed “ Philippine Is. Coll. Bottcher, B.M. 1929-201.” / “ Philippine Islands Manila 8: X: 1914 Coll. Bottcher ” / white, handwritten in black ink and printed “Wallacei ssp. philippinensis Kult det. K. Kult 47” / “red, black framed, black printed “ PARATYPE ” / circle, yellow framed, black printed “ Paratype ” (NHMUK); 1 spec., same data as holotype but with additional label “ Clivina sp.? det. Ing. Jedlička ” [damaged, glued spec.] (NHMUK); 1 ♂, same data as holotype but “2: XI: 1914” (NHMUK); 1 spec., white, handwritten in back ink “Philipp Islands” / “Semper” / black printed “ Fry. Coll. 1905.100.” / white, circle, yellow framed “ Paratype ” / red, black printed and black framed “ PARATYPE ” / white, handwritten in black ink and black printed “ C. Wallacei s. philippinensis Kt. Det. K. Kult 1948 (NHMUK).
Additional material. Luzon: 1 ♀, 1 ♂, Los Baños P. I. Baker / 2569 / Ex Mus. Coll. Agric. Phil Is. / F. van Emden Bequest. B.M. 1960-129. / wallacei (or subsp.) / wallacei Putz. / Gesch. 8. 1938 von Andrews / Clivina vulgivaga Boh. H.E.Andrewes det.; the male with additional label: Wallacei ssp. philippinensis Kult det. K. Kult 47 (NHMUK); 1 ♀, Philippine Is. Coll. Bottcher. B.M. 1929-201. / Philippine Islands Manila 30.X.1914 Coll. Bottcher (NHMUK); 3 specs, same data but 8.VII.1914 (NHMUK); 1 spec., same data but 7.I.1916 (NHMUK); 2 specs, same data but 20.X.1914 (NHMUK); 1 spec., same data but with additional label C. Wallacei s. philippinensis Kt. det. K. Kult 1948 (NHMUK); 1 spec., Phil. Isl. / C. Wallacei s. philippinensis Kt. det. K. Kult 1948 (NHMUK); 1 spec., Manila Philipp. / H.E.Andrewes Coll. B.M. 1945-97. / Wallacei ssp. Philippinensis Kult det. K. Kult 47 (NHMUK); 1 spec., Philippine Islands Manila 6.II.1914 Coll. Bottcher / Philippine Is. Coll. Bottcher. B.M. 1929- 201. / From British Museum Collection / C. Wallacei s. philippinensis Kt. det. K. Kult 1948 / COLLECTIO KAREL KULT COLL.A.DOSTAL, 1999 (CADW); 1 ♀, PHILIP.: Luzon Mountain Prov., Sagada Echo Valley, 23./ 24.2.1999 leg. Schödl (21) (NHMW); 1 ♀, Norzagaray Bulacan, Central Luzon, January 2023, loc. Coll. leg. (CMBB); 1 ♂, Mt. Makiling Luzon Baker / Ex Mus. Coll. Agric. Phil Is. / H.E.Andrewes Coll. B.M. 1945-97. (NHMUK). Marinduque: 1 ♂, Philippines Marinduque Dec. 2013 / Isoclivina helferi-gr. (CADW). Mindanao: 1 spec., Davao Mindanao Baker / H.E.Andrewes Coll. B.M. 1945-97. (NHMUK); 1 ♀, MINDANAO, 30km E of MALYBALAY, BUSDI 5.-9. MAY 1996 BOLM lgt. 1000 m / Clivina group helferi det. Balkenohl, 1999 (SMNS). Leyte: 1 ♂, 4 ♀, PHILIPPINES: LEYTE VISCA N BayBay cultiv. land, 1991 leg. SCHAWALLER & al. / 1.3.1991 / Clivina helferi-group det. Balkenohl VI. 93 (SMNS); 1 spec., same data but sec. forest 100-200 m (SMNS); 1 ♀, 1 spec., PHILIPPINES: LEYTE SW Abuyog, 28.II.1991 river bank leg. SCHAWALLER & al. (SMNS).
The subspecies is described by Kult (1951).
Additional description. Measurements. Body length 5.29–6.05 mm (ẍ = 5.73 mm *), width 1.49–1.82 mm (ẍ = 1.66 mm *), ratio L/W of pronotum 0.89–0.95 (ẍ = 0.93*), ratio L/W of elytra 1.84–1.95 (ẍ = 1.89*); (*n = 10).
The subspecies seems to be slightly larger than the nominotypical subspecies. It differs mainly in the following characters: colour piceous.
Head with neck constriction more distinctly closed by band of punctures, with slight step.
Pronotum (Fig. 66) with lateral margin slightly contracted anteriorly, channel wider.
Elytron (Fig. 31) with intervals more convex, interval seven at apex almost convex, with very short rim at apex.
Male genitalia (Fig. 98). Median lobe of moderate length, arcuate basally, somewhat bulgy in apical half, apex like elongated slight parable. Parameres relatively stable, moderately slender, distorted in apical quarter, with medium long seta each.
Female gonocoxites and epipleurite (Fig. 124). Basal gonocoxite with apex regularly acuminated, with two long setae, apical gonocoxite elongated, slightly curved, cylindrical in cross section in apical two thirds, with two medium sized and one small setae basally, with two long setae apically. Epipleurite trapezoid, with two setae.
Variation. The lateral margin of the pronotum is more or less convex. In some of the specimens including paratypes, one side is straighter whereas the other side is more convex.
Distribution. Known from the following Philippine Islands: Luzon, Marinduque, Mindanao, and Leyte.
Notes
Files
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Linked records
Additional details
Identifiers
Biodiversity
- Collection code
- NHMUK
- Scientific name authorship
- Kult
- Kingdom
- Animalia
- Phylum
- Arthropoda
- Order
- Coleoptera
- Family
- Carabidae
- Genus
- Clivina
- Species
- philippinensis
- Taxon rank
- subSpecies
- Type status
- holotype , paratype
- Taxonomic concept label
- Clivina wallacei subsp. philippinensis Kult, 1951 sec. Balkenohl, 2025
References
- Kult, K. (1951) Revision of the genus Clivina, Latr., from Oriental region. (Col. Carabidae). Casopis Ceskoslovenske Spolecnosti Entomologicke, 48, 16-32.
- Balkenohl, M. (2001) Key and Catalogue of the tribe Clivinini from the Oriental realm, with revisions of the genera Thliboclivina KULT, and Trilophidius JEANNEL (Insecta, Coleoptera, Carabidae, Scarititae, Clivinini). Pensoft Series Faunistica 21. Pensoft, Sofia-Moscow, 83 pp.
- Balkenohl, M. (2021 c) The Clivina komareki - species group from Asia with the description of a new species from the Philippines (Coleoptera: Carabidae: Scaritinae). Linzer biologische Beitrage, 53, 535-543.
- Lorenz, W. (2005) Systematic list of extant Ground Beetles of the World (Insecta Coleoptera " Geadephaga ": Trachypachidae and Carabidae incl. Paussinae, Cicindelinae, Rhysodidae). 2 nd Edition. Printed by the author, Tutzing, 530 pp.
- Lorenz, W. (2024) CarabCat: Global database of ground beetles. Version 15 December 2023. In: Roskov, Y., Ower, G., Orrell, T., Nicolson, D., Bailly, N., Kirk, P. M., Bourgoin, T., DeWalt, R. E., Decock, W., Nieukerken, E. van, Zarucchi, J. & Penev, L. (Eds.), Species 2000 & IT IS catalogue of life. 2021 Annual Checklist. Species 2000. Naturalis, Leiden. Available from: https://www.catalogueoflife.org/annual-checklist/2022 (accessed 31 January 2024)
- Balkenohl, M. (1999) New Clivinini from the Oriental region. 2. Clivina rugosofemoralis nov. spec. and Rugiluclivina leonina nov. spec. from Laos (Coleoptera, Carabidae, Scaritinae). Linzer biologische Beitrage, 31, 337-344.