Tornus subcarinatus
Authors/Creators
- 1. Université de Rennes, Campus de Beaulieu, 263 avenue Général Leclerc 35042 Rennes (France) bouchardblanche @ gmail. com (corresponding author)
- 2. Naturalis Biodiversity Center, P. O. Box 9517, 2300 RA Leiden (The Netherlands) and Department of Earth Sciences, Utrecht University, Princetonlaan 8 a, 3584 CB Utrecht (The Netherlands)
- 3. Naturalis Biodiversity Center, P. O. Box 9517, 2300 RA Leiden (The Netherlands)
Description
Tornus subcarinatus (Montagu, 1803) (Fig. 9F 1-F 3)
Helix subcarinata Montagu, 1803: 438, pl. 7, fig. 9.
Tornus subcarinatus – Van Dingenen et al. 2016: 149, pl. 11, fig. 9, pl. 12, fig. 3. — Landau et al. 2018: 303, pl. 131, fig. 1.
For more, see synonymy list in Van Dingenen et al. (2016) and Landau et al. (2018).
MATERIAL AND DIMENSIONS. — Maximum height 0.8 mm, width 1.2 mm. — RGM.1364994 (19), leg. ACJ; RGM.1365009 (93), leg. AWJ; RGM.1365299 (1), leg. AWJ; RGM.1405531 (1), leg. ACJ.
SPECIES CHARACTERISATION. — Small, depressed, subcircular shell. Protoconch of 2.5 smooth whorls; three teleoconch whorls with carinate shoulder, mid-whorl and peribasal cords, sinuous ribs on dorsum and venter; base flattened, bearing two further elevated cords, the medial periumbilical cord delimiting wide, deep umbilicus with ribs extending within; aperture wide, rounded, strongly oblique in profile.
DISTRIBUTION. — Middle Miocene: Atlantic, Loire Basin, France (Glibert 1949). — Upper Miocene: Atlantic, NW France (Landau et al. 2018). — Lower Pliocene: NSB, Coralline Crag, England (Wood 1848; Harmer 1923; Atlantic, NW France (Van Dingenen et al. 2016); central Mediterranean, Italy (Chirli 2006). — Upper Pliocene: NSB, Red Crag, England (Wood 1848; Harmer 1923). — Upper Pliocene-Pleistocene: Atlantic, NW France (Cossmann 1918; Brébion 1964). — Lower Pleistocene: Atlantic, Selsoif, NW France (this paper). — Upper Pleistocene: NSB, The Netherlands (Van Regteren Altena et al. 1954); Atlantic, British Isles (Harmer 1923). Today this species is present in the Atlantic coasts of Europe from British Isles to Tarifa (Fretter & Graham 1978).
REMARK
Van Dingenen et al. (2016:149) highlighted the problem with the Tornus subcarinatus (Montagu, 1803) species concept as accepted at present, in which specimens with paucispiral and multispiral protoconchs are considered conspecific. They concluded that there were two species: an Atlantic species T. subcarinatus, with a multispiral protoconch of about 2.1- 2.25 whorls, with a small nucleus (counting including first half whorl: Rolán & Rubio 2002: figs 5, 6; Van Aartsen et al. 1998: fig. 7), and a Mediterranean species that is at present unnamed with a paucispiral protoconch of 1.25-1.4 whorls, with a larger nucleus (Rolán & Rubio 2002: figs 13, 14; Van Aartsen et al. 1998: fig. 7). With a protoconch of 2.5 whorls the specimens from Selsoif are typical of the Atlantic species T. subcarinatus.
Notes
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Linked records
Additional details
Identifiers
Biodiversity
- Collection code
- RGM
- Material sample ID
- RGM.1364994 , RGM.1365009 , RGM.1365299 , RGM.1405531
- Scientific name authorship
- Montagu
- Kingdom
- Animalia
- Phylum
- Mollusca
- Order
- Littorinimorpha
- Family
- Tornidae
- Genus
- Tornus
- Species
- subcarinatus
- Taxon rank
- species
- Taxonomic concept label
- Tornus subcarinatus (Montagu, 1803) sec. Bouchard, Wesselingh, Pouwer & Landau, 2025
References
- MONTAGU G. 1803. - Testacea Britannica or Natural History of British Shells, Marine, Land, and Fresh-Water, Including the most Minute: Systematically Arranged and Embellished with Figures. J. White, London, Vol. 1, xxxvii + 291 p.; Vol. 2, p. 293-606, pl. 1 - 16.
- VAN DINGENEN F., CEULEMANS L. & M. LANDAU B. 2016. - The lower Pliocene gastropods of Le Pigeon Blanc (Loire-Atlantique, north west France), 2. Caenogastropoda. Cainozoic Research 16 (2): 109-219.
- LANDAU B., CEULEMANS L. & VAN DINGENEN F. 2018 - The upper Miocene gastropods of northwestern France, 2. Caenogastropoda. Cainozoic Research 18 (2): 177-368.
- GLIBERT M. 1949. - Gastropodes du Miocene moyen du Bassin de la Loire. Premiere partie. Institut royal des Sciences naturelles de Belgique, Bruxelles, 289 p.
- WOOD S. V. 1848. - A Monograph of the Crag Mollusca: or, Descriptions of Shells from the Middle and Upper Tertiaries of the East of England: Volume 1: Univalves. Cambridge University Press, Cambridge. https://doi.org/10.1017/CBO9781139680745
- CHIRLI C. 2006. - Malacofauna Pliocenica Toscana, 5. Caenogastropoda. Firenze, 144 p.
- BREBION P. 1964. - Les gasteropodes du Redonien et leur signification. 1 - 2. These de doctorat des Sciences. Faculte des Sciences de l'Universite de Paris, Paris, 775 p.
- FRETTER V. & GRAHAM A. 1978. - The Prosobranch Molluscs of Britain and Denmark. Part 4. Marine Rissoacea. Journal of Molluscan Studies, suppl. 6: 153-241.
- ROLAN E. & RUBIO F. 2002. - The Family Tornidae (Gastropoda, Rissooidea) in the East Atlantic 13 (supplement). Sociedad Espanola de Malacologia, special number of II International Congress of the European Malacological Societies, 98 p.
- VAN AARTSEN J. J., CARROZZA F. & MENKHORST H. 1998. - Tornus mienisi, a new species of Tornus from the Eastern Mediterranean (Mollusca: Prosobranchia). Bollettino Malacologico 33: 135-138.